in copyright - non-commercial use permitted rights ... · 1!/*+,#-%#.#! +!/*+,#-%#.#!/*+,#-%#.#!
TRANSCRIPT
Research Collection
Doctoral Thesis
Altitudinal adaptation in Arabis thaliana
Author(s): Quèbre, Nicolas
Publication Date: 2012
Permanent Link: https://doi.org/10.3929/ethz-a-007632306
Rights / License: In Copyright - Non-Commercial Use Permitted
This page was generated automatically upon download from the ETH Zurich Research Collection. For moreinformation please consult the Terms of use.
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Brassicaceae Familly - Worldwide distributed, from costal lands to up to 3500 m a.s.l. Annual and rapid cycling – Self fertlizer - Opportunistic weed growing in disturbed areas
Arabidopsis thaliana (L.) Heynh. - The Mouse-ear cress
Kingdom Subkingdom Superdivision Division Class Subclass Order Family Genus Species
Plantae – PlantsTracheobionta – Vascular plantsSpermatophyta – Seed plantsMagnoliophyta – Flowering plantsMagnoliopsida – DicotyledonsDilleniidaeCapparalesBrassicaceae – Mustard familyArabidopsis Heynh. – rockcressArabidopsis thaliana (L.) Heynh.
Classification
Inflorescence (Corymb)Fruit (Silique)
Main stemStem leaf
Secondary stemLeaves Rosette form
Morphology
© Biopix via eol.org, CC by-nc
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Pop ID Location name Families Altitude
(m a.s.l.) Altitude
grouping International Coordinates
BRI Bristen 7 800 Low alt. origin 46°46'15.92"N / 8°42'6.37"E NAT Naters 7 850 Low alt. origin 46°20'0.43"N / 7°59'15.72"E EGG Eggerberg 7 900 Low alt. origin 46°18'51.53"N / 7°52'42.38"E AUS Ausserberg 6 1000 Low alt. origin 46°18'54.88"N / 7°52'4.55"E SAE Saas Fee 7 1792 High alt. origin 46°6'37.65"N / 7°55'49.88"E GRA Grächen 7 1850 High alt. origin 46°9'52.12"N / 7°50'15.87"E SAB Saas Fee 7 1949 High alt. origin 46°7'30.68"N / 7°55'40.53"E SAO Saas Fee 7 2012 High alt. origin 46°6'24.65"N / 7°54'38.60"E BRA Graubünden 3 2040 High alt. origin 46°39'25.23"N / 10°1'12.41"E ZIN Zinal 7 2700 High alt. origin 46°8'52.51"N / 7°38'48.49"E
!
24!
Experimental sites
Region Tamins Eichwald (Pradamal) 626 Low altitude 46°50'19.12"N / 9°25'55.85"E Tamins Lusbühel (Girsch) 796 Low altitude 46°50'36.97"N / 9°24'13.24"E Haldenstein Arella 1004 Low altitude 46°52'30.37"N / 9°30'36.34"E Haldenstein Nesselboden 1403 Low altitude 46°52'14.02"N / 9°29'29.38"E Haldenstein Calandahütte 1999 High altitude 46°53'20.75"N / 9°29'23.62"E Laax Nagens 2177 High altitude 46°51'56.79"N / 9°14'13.21"E Laax Mutta Rodunda 2401 High altitude 46°52'12.67"N / 9°13'33.73"E !
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Full
MA
M m
odel
slm
ergl
mer
glm
erlm
erlm
erlm
erlm
erlm
erlm
erEffect
pMCMC-value
Pr(>|z|)
Pr(>|z|)
pMCMC-value
pMCMC-value
pMCMC-value
pMCMC-value
pMCMC-value
pMCMC-value
alt.origin
0.37
400.
0048
*0.
2231
0.39
220.
3246
0.28
920.
7498
0.80
700.
3958
alt.planted
0.32
680.
3254
0.07
130.
0034
*0.
0314
*0.
0044
*0.
4372
0.97
480.
0001
*cohort
0.00
01*
0.02
35*
0.00
00*
0.00
01*
0.00
01*
0.03
98*
0.00
01*
0.00
01*
0.01
72*
alt.origin*alt.planted
0.01
34*
0.00
01*
----
----
----
----
----
----
0.59
620.
0001
*--
----
----
----
--alt.origin*cohort
0.14
740.
0046
*--
----
----
----
----
----
--0.
4718
----
----
0.00
01*
0.00
82*
alt.planted*cohort
0.00
04*
0.00
00*
0.00
77*
0.00
01*
0.00
01*
0.00
10*
0.00
01*
0.00
01*
0.00
01*
alt.origin*alt.planted*cohort
0.01
64*
0.00
30*
----
----
----
----
----
----
0.02
04*
----
----
----
----
----
----
Su
bm
odel
s
alt.origin
0.53
580.
0052
*0.
3338
0.46
340.
4632
0.18
720.
6652
0.50
900.
3602
alt.planted
0.39
880.
3253
0.06
040.
0036
*0.
0242
*0.
0086
*0.
5956
0.79
140.
0182
*alt.origin*alt.planted
0.01
66*
0.00
01*
----
----
----
----
----
----
----
----
0.02
20*
----
----
----
----
alt.origin
0.87
660.
6230
0.34
110.
9670
0.82
420.
0970
0.79
860.
3520
0.96
58alt.planted
0.94
360.
0000
*0.
0002
*0.
4318
0.79
920.
0028
*0.
0286
*0.
0034
*0.
0010
*alt.origin*alt.planted
----
----
----
----
----
----
----
----
----
----
0.00
24*
0.00
01*
----
----
----
----
Trai
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ter
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al
coho
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Sum
mer
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co
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Ros
ette
siz
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Low
alt.
plan
ted
alt.origin
0.33
960.
1070
0.17
240.
2718
0.70
16H
igh
alt.
plan
ted
alt.origin
0.47
900.
0050
*0.
3792
0.35
720.
1658
Low
alt.
orig
inalt.planted
0.66
220.
0000
*0.
7211
0.00
06*
0.01
72*
Hig
h al
t.or
igin
alt.planted
0.37
500.
3240
0.42
950.
0080
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0048
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#71#$I3#:#7542LG! $83K#K21! 924423! 54! ;#B;! 5L4#4812! _:632! 4;57! Q,v! 6C! $83K#K5La! 4;57! 54!
L6]!5L4#4812!_L2$$!4;57!\Yv!$83K#K5L!`!A#B832!P9a%!?7!4;2!$5:2!]5GJ!4;2!5L4#4812!NL57421!
5L$6! #7CL827I21! 4;2! CL6]23#7B! #7I#127I2! _'59L2! +a%! U2$$! 4;57! /,v! 6C! 4;2! NL574$! 4;54!
$83K#K21!54! L6]!5L4#4812!$#42$!CL6]2321J!];#L2!Q,v!6C!NL574$!54!;#B;!5L4#4812!4357$NL574!
$#42$! CL6]2321! _A#B832! Pa%! 0LL! CL6]23#7B! NL574$! N3618I21! L6]! 984! $#:#L53! 5:6874$! 6C!
$#L#d82$! _924]227!+,! 571!P+,a! _'59L2!+J! A#B832!Pa%! ?7! 4;#$! I6;634J! 4;2! 6K235LL! K5L82! 6C!
NL574!C#472$$!#7!964;!5L4#4812!NL57421!571!5L4#4812!6C!63#B#7!]5$!32L54#K2LG!L6]!I6:N5321!
46!4;2!]#7423!57785L!I6;634!571!1#1!764!$;6]!46!92! #:N5I421!9G!964;!2CC2I4$!_'59L2!+J!
A#B832!P9a%!
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5:67B!5L4#4812$!6C!63#B#7!63!5L4#4812$!NL57421! #7! 4;#$!I6;634J!];#L2!36$2442!1#5:2423!54!
CL6]23#7B!4#:2!]5$!B27235LLG!;#B;23!C63!NL574$!4357$NL57421!46!;#B;!5L4#4812!4;57!4;6$2!
4357$NL57421!46!L6]!5L4#4812!_'59L2!+a%!AL6]23#7B!4#:2!5C423!$76]:2L4!]5$!$#B7#C#I574LG!
12L5G21!#7!;#B;!5L4#4812J!$#:#L53LG!C63!964;!B368N$!6C!N6N8L54#67$!_'59L2!PJ!A#B832!+Ca%!b2!
69$23K21!5!$#B7#C#I574!#74235I4#67*924]227!4;2!5L4#4812!6C!63#B#7!571!4;2!5L4#4812!NL57421*
C63! 4;2! 36$2442! $#R2! 5C423! 4;322! ]22^$! 571! C63! EU0! _'59L2! +a%! @674353G! 46! 4;2! ]#7423!
57785L!I6;634J!5!;#B;23!K5L82!6C!EU0!]5$!;232!5$$6I#5421!]#4;!57!#7I325$2!#7!C#472$$!67LG!
54!;#B;!5L4#4812!4357$NL574!$#42$!571!1#1!764!I6743#9842!46!C#472$$!54! L6]!5L4#4812!_'59L2!
/a%! EU0! ]5$! #:N5I421! 9G! 4;2! 2CC2I4! 6C! 5L4#4812! NL57421! #7! $89:612L$! 575LG$#7B!
p S p S p S p S
Rosette size 3 weeks 0.0001 * 1.81 0.0001 * 2.37 0.0166 * 0.80 0.0001 * 0.42Rosette size flowering 0.0001 * 1.75 0.0001 * 2.32 0.0238 * 0.54 0.0001 * 0.83SLA 0.4886 0.83 0.0020 * -1.26 0.4562 1.14 0.0290 * 0.57Flowering time 0.0001 * -0.54 0.0605 -0.59 0.0001 * -3.02 0.0010 * -0.50
Traits
Low altitude High altitude Low altitude High altitude
Winter annual cohort Summer annual cohort
!
32!
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$8BB2$4#67! 6C! n5]2I^#! 571! &9234! _+,,/a!]2! 8$21! $89:612L$! 46! C834;23! 42$4! 2CC2I4$! 6C!
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]#7423! 57785L! I6;634J! 571! ];27! 4357$NL57421! 54! ;#B;! 5L4#4812J! 67LG! N6N8L54#67$! 6C!
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N6N8L54#67$! C36:! L6]! 5L4#4812! 6C! 63#B#7! #7! 423:$! 6C! $83K#K5L%! )6! C834;23! B2764GN2!
1#CC23274#54#67!#7!4;2!64;23!I671#4#67$!6C!683!2SN23#:274!]5$!C6871!984!4;2!N6N8L54#67$!
63#B#754#7B! C36:! L6]! 5L4#4812$! 1#$NL5G21! ;#B;23! $83K#K5L! ];27! NL57421! 7253! 4;2#3!
5L4#4812! 6C! 63#B#7! 4;57! NL57421! 54! ;#B;! 5L4#4812$%! '6B24;23J! 4;#$! $8BB2$4$! 515N4#K2!
1#CC23274#54#67! C63! 4;2! ]#7423! 57785L! I6;634%! ?7! 4;2! $8::23! I6;634J! #4! $22:$! 4;54!
27K#367:2745L!I671#4#67$!]232!:632!L#^2LG!4;2!12423:#7574$!6C!4;2!C#472$$!C63!5LL!NL574$!
5$! 5LL! NL574! 325I421! $#:#L53LG! 46! 4357$NL574! L6I54#67$%! b2! 1#1! 764! #7IL812! 2CC2I4$! 6C!
B23:#754#67J! 127$#4G! 12N27127I2! 571! 64;23! 9#64#I! #74235I4#67$! L#^2! B35R#7BJ! ^76]7! 46!
5CC2I4!12K2L6N:274!$45B2$!#7!!/*+,#-%#.#!_"676;82!24!5L%!+,,Y9J!"676;82!+,,QJ!0^#G5:5!
571!sB327!+,P+a%!b#4;!5!32I#N36I5L!4357$NL574!67!5!$:5LL!$I5L2!924]227!$571!1872!571!
#7L571!;59#454$! #7!)24;23L571$J! #4!]5$!5L$6!69$23K21! 4;54!$6:2!!/*+,#-%#.#!N6N8L54#67$!
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964;! N6N8L54#67$! ]232! 515N421! 46! 4;2#3! 754#K2! 27K#367:274$! 984! C6871! 4;54! $83K#K5L!
N544237$! 924]227! $#42$! 571! N6N8L54#67$! 1#CC2321! #7! :5B7#4812! 5I36$$! G253$%! ';8$J!
!
33!
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4357$NL574! $481#2$%! D83! $481G! 32K25L$! 4;54! #4! :5G! 5L$6! 12N271! 67! 4;2! L#C2! ;#$463G!
2SN32$$21!];24;23!515N4#K2!1#CC23274#54#67!#7!!/*+,#-%#.#!N6N8L54#67$!I57!92!#7C23321%!!
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1G75:#I$! 6C! 754835L! N6N8L54#67$!:5G! 4;232C632! L#:#4! 32I274! #7C2327I2$! 59684! IL#:54#I!
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^76]!G24!];#I;!L#C2!;#$463G!683!N6N8L54#67$!2SN32$$!#7!4;2#3!754835L!;59#454%!b2!5$$8:2J!
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4;2! N32K#68$T:274#6721! $481#2$J! ]2! 8$21! 571! B368N21! :632! 4;57! 4]6! $#42$! 571!
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1#CC2327I2$!924]227!435#4$!2SN32$$21!;#B;!571!L6]!$#42$J!$8BB2$4#7B!5!;#B;!N36N27$#4G!C63!
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34!
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12:67$435421! IL6$2! L#7^$! 924]227! $#R2! 435#4$! 571! C#472$$! _U2#:8! 571! A#$I;23! +,,-J!
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!
!
Pop ID Location Altitude (a.s.l.) Nbr of Genotypes
BRI Bristen 800 46°46'15.92"N / 8°42'6.37"E 4NAT Naters 850 46°20'0.43"N / 7°59'15.72"E 4AUS Ausserberg 1000 46°18'54.88"N / 7°52'4.55"E 4SAB Saas Fee 1949 46°7'30.68"N / 7°55'40.53"E 4SAO Saas Fee 2012 46°6'24.65"N / 7°54'38.60"E 4ZIN Zinal 2700 46°8'52.51"N / 7°38'48.49"E 4
International Coordinates
Region Location Altitude (a.s.l.) Sowing autumn Snow fall 2008 Snowmelt 2009 Sowing Summer Snow fall 2009 Snowmelt 2010
Tamins Eichwald (Pradamal) 626 46°50'19.12"N / 9°25'55.85"E 15.nov.08 20-30.nov 17-19.march 30.may 15-20.nov 20-22.marchTamins Lusbühel (Girsch) 796 46°50'36.97"N / 9°24'13.24"E 07.nov.08 20-30.nov 04-06.april 15.june 26.oct 01-06.aprilHaldenstein Arella 1004 46°52'30.37"N / 9°30'36.34"E 31.oct.08 10.nov 10-12.april 15.june 20-25.oct 01-06.aprilHaldenstein Nesselboden 1403 46°52'14.02"N / 9°29'29.38"E 24.oct.08 20-30.oct 01-04.may 03.july 8-10.oct 01-06.aprilHaldenstein Calandahütte 1999 46°53'20.75"N / 9°29'23.62"E 17.oct.08 20-30.oct 07-08.may 08.aug 8-10.oct 18-22.mayLaax Nagens 2177 46°51'56.79"N / 9°14'13.21"E 13.oct.08 16.oct 20-25.june ----- 30.sept 10-15.juneLaax Mutta Rodunda 2401 46°52'12.67"N / 9°13'33.73"E 13.oct.08 16.oct 20-25.june ----- 30.sept 10-15.june
International Coordinates
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Site 0.0000 * 0.0000 * 0.0000 * 0.0720Population 0.1052 0.1468 0.4295 0.0516Site x Population -------- -------- -------- --------
Site 0.0000 * 0.0000 * 0.0001 *Population 0.0276 * 0.4301 0.2745Site x Population -------- -------- 0.0002 *
Site 0.0000 * 0.4738 0.0544Population 0.0211 * 0.0685 0.6866Site x Population -------- -------- --------
(a)
(b)
(c)
Source of variation Germinant number
Reproductive plant number
Survivors Autumn germinants
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51
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=4338"*)'@%.>*
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