bollino_2010_leodonta tagaste

8/3/2019 Bollino_2010_Leodonta tagaste

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Genus Vol. 21(1): 89-100 Wrcaw, 30 III 2010

Tanmical nte nL (Felder& Felder, 1859)(Lepiptera: Pieriae)

Maurizio BollinoVia Raplla 40, I-73100 Lecce, Ital. E-mail: [email protected]

aBstract.Brief nte cncerning te tanm an itribtin fL (Felder& Felder, 1859) are prvie, wit ecriptin f tree new bpecie.

Ke wr: entmlg, tanm, nmenclatre, Blivia, Clmbia, itribtin, Ecar,L, Netrpical Pierinae, Per.

INTRoduCTIoN

Te Pieri genL wa ecribe in 1870 b Butler. It i clel relate(VenaBles 1993; LaMas & Bollino 2004; BraBy l. 2007) t Pru HerricH-scHFFer, 1867 an CicButler, 1870 an range frm Cta Rica t Blivia,ccrring nl in mntane Anean abitat. Te tematic an tanm fLare prl knwn, relting in tatement c a te fllwing: Te nmber f pe-cie in ti gen i a matter f cnjectre me atr claim n mre tan tw r

tree, ter a a mc a ten! (dABrera 1984: 66) r It i till nknwn wman pecie tere are inL (deVries 1987: 90). Te rtL lit wa

pblie b talBot in 1932, recgniing 3 pecie: L. yi (douBleday, 1847), L. ll (Hewitson, 1860), an L. (Felder & Felder, 1859), wile temt recent ne (LaMas 2004) recgnie 5 pecie: L. yi,L. ,L. ll,L. zbi (Felder& Felder, 1865) anL. zbi (HoppFer, 1869).

Te aim f ti paper i t tart relving at leat me f tee prblematic i-e. It i evte tL (Felder& Felder, 1859), te nl pecie inte gen caracterie b bt male an female aving te frewing ical cellmtl ccpie b wite.


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MATERIAL, METhods, ACRoNyMs ANd ABBREVIATIoNs

In preparatin fr a reviin f te gen, a lt f ata n L were acc-

mlate ring reearc viit b te atr t vari intittin, an al tank tte cperatin f clleage. Nearl 400 pecimen f te pecie treate ere wereeamine in te fllwing cllectin: Natral hitr Mem, Lnn, unite King-m; Mm Natinal hitire Natrelle, Pari, France; Mem fr Tierkne,dreen, German; Me e hitria Natral, univeria Nacinal Mar e sanMarc, Lima, Per; Natinal Mem f Natral hitr, smitnian Intittin,Waingtn, usA; zlgice Mem er hmblt univeritt, Berlin, German;zlgice staatammlng, Mncen, German; staatlice Mem fr Natrkne,stttgart, German; zlgical Mem, Jagellnian univerit, Krakw, Plan;Pierre Boyer, Le P ste. Rparae, France; stepane Attal, Pari, France; Gabriel

Rodriguez, Enviga, Clmbia; Jean-Frani Le CroM, Bgt, Clmbia, pl teprivate cllectin f te atr.

Acrnm fr intittin an private cllectin ling pecimen eamine inti reviin mtl fllw EVenHuis (2008).

BMNh: Te Natral hitr Mem, Lnn, Englan;CRMC: cllectin f Gabriel rodriguez, Meellin, Clmbia;ICN: Intitt e Ciencia Natrale, Bgt, Clmbia;JLBC: cllectin f Jean-Frani Le CroM, Bgt, Clmbia;

MBLI: cllectin f Marii Bollino, Lecce, Ital;MusM: Me e hitria Natral, univeria Nacinal Mar e san Marc,

Lima, Per;MzuJ: zlgical Mem, Jagellnian univerit, Krakw, Plan;PBPF: cllectin f Pierre Boyer, Le P ste. Rparae, France;uNCC: Me e hitria Natral, univeria Nacinal e Cala, Maniale,

Clmbia;zsM: zlgice staatammlng, Mncen, German.

Abbreviatin

FWd: frewing, ral;FWV: frewing, ventral;hWd: inwing, ral;hWV: inwing, ventral;hT: hltpe;PT: Paratpe;LT: Lecttpe.


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91TAxoNoMICAL NoTEs oNLeodontatagaste

REsuLTs ANd dIsCussIoN

Leodonta tagastetagaste (Felder& Felder, 1859)

(g. 1-4)

eurp tNb. Felder& Felder, 1859b: 396, Pl. 9: Fig. 1.

typelocality[Nrt-wetern] Per.

typeMaterialLecttpe male (BMNh), eignate b ABadjieV (2006: 118), illtrate b

ABadjieV (2006: Fig 89, 90).

L wa riginall ecribe frm Per an, ntil recentl, wa cn-iere t be a mntpic pecie ranging frm Clmbia t Blivia. onl a few earag constantino an salazar (in le croMet al. 2004) ecribe a new bpeciefrm Clmbia. Apparentl, n previ atr ave ever eamine te tpe, r if teave, never realie tat it i ver ifferent frm all knwn eatern Anean pecimenf te pecie. Recentl ample pecimen frm t-wetern Ecar an nrt-wetern Per perfectl matc te lecttpe (g. 1), cnrming tat it wa prbablcllecte mewere in nrt-wetern Per, terefre te nmintpical pplatinare gegrapicall eparate an penticall ifferentiate frm an eatern Anean

repreentative f te pecie.specimen fL (g. 2-4) are ver rare in bt pblican private cllectin, an I wa able t eamine nl 8 male an 3 female, plte tpe. specimen are al rarel met wit in te el: ti partiall eplain tetanmic miinterpretatin b pat atr. A cnrmatin f it carcit, I fnnl ne bibligrapical qtatin (zelada, 2004).

exaMinedMaterial (9 male, 3 female)LT male, Per / Feler / Cll N, in BMNh; 1 female, Per Cajamarca / El

Clqe / Via Celenin-Bala / ~ 652s 7805W/ m. 1800-2000 VI-VII.2006

/ Lg. lcal cllectr; 2 male, 1 female , Ecar Lja/ km. 19 Lja-La Tma / ~359s 7818W / m. 2300 xI.1999 / lg. Imael Ala, all in MBLI; 1 male, RteLja-Catama, 2250 m, km. 19, 7.xI.1996, Ecar; 5 male, 1 female, n ata, allin cll. attal (Pari, France).

diagnosticcHaractershWV ellw ical ban large, cmpe f pale ellw merge pt, wit el-

lw cale al cvering te vein. Te ban i brken nl at vein M2. hWV ctalblack pt rt an qare ff, clearl cring vein sC+R

1an eparate frm baal

ark area in cell sC+R1.

Te caracter lite abve permit an immeiate ifferentiatin between te nmi-ntpical bpecie an all eatern Anean pplatin, wic are te fllwing


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caracter: hWV ellw ical ban cmpe f ellw pt, eparate frm eacter b ark-cale vein. hWV ctal black pt tick an lng, an cntaine inievein sC+R

1

an cnnecte t baal ark area in cell sC+R1

b a ark cale vein.

distriButionAccring t crrent knwlege, nmintpical pplatin range frm t-

wetern Ecar (Lja) twar t nrt-wetern Per (Cajamarca).Eatern Anean pecimen, are, n te cntrar, mc cmmner in te el an

well-repreente in mt cllectin. Teir pattern i apparentl qite cnitent, bteaminatin f lng erie allwe me t acertain tat te cl be eparate intfr bpecie, incling te nrternmt p. clmbi, an ne new Pervianbpecie ptlate n b LaMas (2004). separatin f te vari eatern Anean

pplatin i mainl bae n te wit f te hWd marginal black ban in btmale an female, an ifference fn in te hWV ical ban.

1-4.L : 1 LT ver; 2 male ral: Ecar Lja/ km. 19 Lja-La Tma/ ~ 359s7818W/ m. 2300 xI.1999/ lg. Imael Ala, cll. MBLI; 3 ame, ventral; 4 ame, female ral:Per Cajamarca/ El Clqe/ Via Celenin-Bala/ ~ 652s 7805W/ m. 1800-2000 VI-VII.2006/

Lg. lcal cllectr, cll. MBLI


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5-7.L clmbi: 5 male ral: Clmbia Cninamarca / Cia La Flreta / ~452N 7404W / m. 2600-2700 26.VII.2007 / Lg. Tma Prc, cll. MBLI; 6 ame, ventral; 7 ame,female ral: Ecar Lja/ near Lja/ m. 2500 x.2000, cll. MBLI. 8-10.L clri:8hT ral: Per Pac / Cr. yanacaga / El Pajnal / ~ 1038s 7518W/ m. 2900 xII.2004 /Lg. J Bttger, preentl in cll. MBLI; 9 ame, ventral; 10 ame, female PT ral: Per Pac /

Meapata / ~ 1046s 7522W / m. 1200-1400, Ix.2003, lg. lcal cllectr, cll. MBLI


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LeodontatagastecolombianaConstantino & salazar 2004(g. 5-7)

Lclmbi p. nv. In le croM et al., 2004: 86, Pl. Pieriae 37 Tip.

typelocality[Clmbia], Cala, salamina, san Fli.

typeMaterialhltpe male in ICN, riginall eignate.

exaMinedMaterial(14 male, 6 female)hT male, Clmbia Cala / salamina san Feli / 3100 m feb. 1995 / C. sar-

mient leg. / in ICN; AT female, Qin / Reerva Acame, salent / 2700 m., 11.I.89/ J. Vle lg., in uNCC; 1 male, Clmbia Antiqia / Amaga / m. 2500; 1 female,Clmbia Antiqia / san Feli / 2700 m. 10/08/2000, all in CRMC; 1 male, LaBnita/ scmbi/ Ecar/ VI-1996; 1 male, san Anre, 1800 m/ Rgin e zmba/13-VIII-1998/ zamra-Cincipe/ Ecar/ Cae s. Et M. Attal; 1 male, Rgin eVallali/ 2600 m., 8-Ix-2001/zamra Cincipe/ Ecar; 1 female, Ancienne rteLja-zamra/ 2500 m/ 8-V-2000/ zamra Cincipe/ Ecar; 1 male, Ancienne rteLja-zamra/ 2500 m/ 31-V-2000/ zamra Cincipe/ Ecar, all in cll.attal (Pari,France); 1 female, Clmbia Antiqia / El Retir / ~ 604N 7530W / m. 2800

10.VIII.2007 / Lg. Tma Prc; 3 male, Clmbia Antiqia / Alt el Pare

Amaa / ~ 617N 7542W / m. 2800 12.VIII.2007 / Lg. Tma Prc; 1 male,Ecar Lja/ yangana/ ~ 422s 7911W/ m. 2500 V.2000/ lg. Imael Ala; 1male, Ecar Lja/ l ra Lja-zamra/ m. 2600 Nvember 1999/ lg. I. Ala;1 male, Ecar Aa Prv./ Mala/ ~ 258s 7842W/ m. 3200 14-I-2004/ lg.Bllin & Vitale; 1 male, Ecar Aa Prv./ Mala/ ~ 258s 7842W/ m. 3200

xII-2003/ lg. lcal cllectr; 1 male, Ecar Aa Prv./ Maila (16 km EsEGalace)/ ~ 258s 7842W/ m. 3200-3350 I.2003, lg. Eclie Ala; 1 female,Ecar Lja/ Cenen Alt (Lja)/ m. 2800 Nvember 1999/ lg. I. Ala; 1 female,Ecar Lja/ near Lja/ m. 2500 x.2000, all in MBLI

diagnosisTi bpecie i caracterie b bt male an female aving a black marginal

hWd ban ccping apprimatel ne-tir f te pace frm te rigin f vein M3

t te ege f te wing. hWd marginal black ban i itinctl eparate frm icalwite area, bt in te anal prtin it i ffe wit wite cale in cell 2A-1A-C

2

an ccainall in cell C1, t frming a tick grei ban apprimatel 1 mm

wie. hWV ical ban wit ellw pt in cell M2-M

3-C

1f nearl te ame ie,

pentagnal, tcing eac ter alng a plane f everal mm., tat in cell M2ccain-

all being ligtl maller tat te in cell M3-C

1.


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noteconstantino an salazar(in Le CroMet al. 2004), wilt ecribing teir new

bpecie, tate [th w ubpci] iffr frm miypicl ubpci frmecur Pru fr wi hp p whi clr f mi b, which i

lih crm yllw i L. . . [tranlate frm spani tet]. Frm te riginalecriptin, it i nt at all clear weter te cnier te wetern r eatern Eca-rian an Pervian pplatin t be te nmintpical bpecie. Anwa, te ligtcream ellw clr a n iagntic vale, a it ccainall ccr in all eaternAnean pplatin.

distriButionTi bpecie i preent frm Clmbia (Central Crillera an central-tern

prtin f oriental Crillera) twar t Lja Prvince (Ecar).

Tree mre far necribe bpecie can be peneticall ifferentiate. Iterefre brie ecribe tem belw.

Leodonta tagaste calderonin. ssp.(g. 8-10)

typeserieshltpe: male, Per Pac/ Cr. yanacaga/ El Pajnal/ ~ 1038s 7518W/

m. 2900 xII.2004/ Lg. J Bttger, preentl in MBLI, will be epite in MusM

; Paratpe (186 male, 10 female): 1 male, Per Amana / Ri Vere / ~ 628s7735W / m. 1200 xII.1997 / Lg. Benign Calern; 4 male, Per san Martin /Pente El Aente / ~ 542s 7735W / m. 1600-1800 xII-2003 / lg. lcal cllectr;1 male, Per Amana / Pente Nieva / ~ 541s 7747W / m. 2300 III.2005 / Lg.Benign Calern; 1 male, Per Amana / san J e Mlinpampa / ~ 613s7738W / m. 2300 I-II.2006 / Lg. Benign Calern; 2 male, Per Amana/ Per Ri Gall / ~ 5 51s 7758W / m. 1650-1850 III.2005 / Lg. BenignCalern; 22 male, Per - dept. Amana / Rrige e Mena / Qebraa ePirrr / ~ 623s 7726W / m. 1700 -1800 / Lg. Benign Calern; 27 male, Per

Amana / Abve Per Ri Gall / ~ 553s 7813W / m. 1550-1700 Agt

2002 / lg. Benign Calern; 32 male, Per Amana / Per Ri Gall / ~ 553s7813W / m. 1600-1700 VIII 2001 / lg. Benign Calern; 2 male, Per Ama -na / near Rrige e Mena / ~ 620s 7724W / m. 1700-2000 / lg. BenignCalern; 6 male, Per Amana / Abve Per Ri Gall / ~ 553s 7813W/ m. 1800 8-15.xII.2001 / lg. Benign Calern; 1 male, Per Amana / Track

between / Mlinpampa an Granaa / m. 3150 -3250 / octber 2000 / Lg. Calern;2 male, Per hanc / Carpi Pa / ~ 942s 7605W / m. 2800 III-V-2001/lg. lcal cllectr; 2 , Per hanc / abve Ting Maria / m. 1800 x.1998; 5male, Per hanc / hanacare / ~ 946s 7554W / m. 2700-3000 xII-2006/ Lg. J Bettger; 1 male, Per Pac/ Palcama/ hancabamba ditr./ ~ 1025s

7534W/ m. 2200 VIII.2003/ Lg. lcal cllectr; 4 male, Per Pac/ Palcama/


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11-13.L pru: 11 hT ral: Per Cc / Ra Acjanac-Pillcpata / ~ 1308s7125W / m. 2100-2500 xI-xII.2004/ lg. lcal cllectr, preentl in cll. MBLI; 12 ame, ventral;13 ame, female ral: Per Cc / Ri Cipata / m. 1500-2000, I.1999, lg. lcal cllectr, cll.MBLI. 14-16.L xrm : 14 hT ral: Blivia, ynga e Crani, 2500 m.,2.x.1953,leg.

W.Frter,in zsMC; 15 ame, ventral; 16 ame, female PT ral: Per Pn dept./ Aqele (sania)/ ~ 1404s 6942W / m. 2100-2200 xI.2004 / Lg. J Bettger, cll. MBLI


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hancabamba ditr./ ~ 1025s 7534W/ m. 2200 x.2003/ Lg. lcal cllectr; 4male, Per Pac/ Ceva Blanca/ hancabamba ditr./ ~ 1031s 7534W/ m. 2600

x-xI.2003/ Lg. J Bettger; 4 male, Per Pac/ Ceva Blanca/ hancabambaditr./ ~ 1031s 7534W/ m. 2600 xI.2003/ Lg. J Bettger; 1 male, 1 female,Per Pac/ Meapata/ ~ 1046s 7522W/ m. 1200-1400 Ix.2003/ Lg. lcal cl-lectr; 1 male, Per Pac / san Antni e Pc / ~ 1043s 7532W / m. 2000

I.2005 / Lg. J Bttger; 1 male, Per Pac/ Trrebamba/ hancabamba ditr./ ~1031s 7538W/ m. 2500 Ix.2003/ Lg. lcal cllectr; 1 male, Per Pac/ sanAlbert/ oapampa ditr./ ~ 1036s 7520W/ m. 2000 Ix.2003/ Lg. lcal cllectr;2 male, 1 female, Per Pac/ Cr. yanacaga/ El Pajnal/ ~ 1038s 7518W/ m.2800 xI.2004/ Lg. J Bttger; 8 male, 2 female, Per Pac/ Cr. yanacaga/El Pajnal/ ~ 1038s 7518W/ m. 2900 xII.2004/ Lg. J Bttger; 6 male, Per

Pac/ Cr. yanacaga/ El Pajnal/ ~ 1038s 7518W/ m. 2800 xI.2005/ Lg.

J Bttger; 9 male, 2 female, Per Pac/ Cr. yanacaga/ El Pajnal/ ~ 1038s7518W/ m. 2800 Ix.2006/ Lg. J Bttger, all in MBLI; 13 male, 1 female, [Per Amana], Qa. El Ci, Pmacca / 15.09.1999 / B.Calern leg. / 2500 m;3 male, [Per Amana], Cer, Mena / 02.2002 / B.Calern leg. / 2000 m;2 male, 1 female, ame ata, bt 10.2002; 3 male, [Per Amana], Alt Ri

Nieva / 06.2002 / 2200-2500 m / B. Calern leg.; 1 male, [Per Amana], Qa.hingilpata / Mena / 25.08.1998 / J.Wjtiak leg.; 1 male, [Per Amana],Qa. Pirrr / Mena / 1700 m / 28.08.1998 / T.Prc leg.; 1 female, [Per Ama-na], Pea Blanca / Pmacca / 02.07.1999 / 2800 m / B. Calern leg.; 1 male,[Per Amana], Qa. Llanaic / 14.10.1998 / J.Wjtiak leg.; 1 male, [Per

Pac], Villa Rica / oapampa / 2000-2100 m / 06.10.2002 / T.Prc leg.; 1 male,Per/ dep. Jnn / Cancama / Caciac Caca / Qa. Malamb / , 2600-2700m. / leg. T. Prc., all in MzuJ; 1 male, Rrige e Mena (Amana) / 1999 ?r 2000 ? / Benign Calern leg.; 1 male, 1 female, La sia / 2400/2700m (Pac)/ 31/10/2004 / P.Ber leg.; 1 male, La sia / 2200/2400m / oapampa (Pac) /28/10/2006 / P.Ber leg.; 5 male, Rte Pacacpan ver hanacare km39 / Et eAcma / 2600m (hanc) / 09 45 412 s, 075 54 844 W / 23/10/2006 / P.Ber leg.;3 male, Ceva Blanca / 2300m / hancabamba, (Pac) / 6/12/2003 / P.Ber leg.; 1male, sa (Icc) / a e e Milp 3100/3300m / 10 22 968 s, 075 38 021W ( (Pac) 27/5/2005 / P.Ber leg.; 1 male, Ceva Blanca / 2500m / hancabamba

(Pac) / 11/2004 / J.Bttger leg., all in PBPF.

diagnosisTi bpecie i caracterie b bt male an female aving a black marginal

hWd ban ccping apprimatel ne-tir f te pace frm te rigin f vein M3

t te ege f te wing. hWd marginal black ban i eparate frm ical wite areab a ark ban ffe wit wite cale; ti grei ban i apprimatel 3 mm wiein cell 2A an becme prgreivel narrwer in cell 1A-C

2-C

1-M

3. hWV ical

ban wit ellw pt in cell M2-M

3-C

1prgreivel maller, pentagnal, tcing

eac ter alng a plane f everal mm., tat in cell M2

alwa being te larget.


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distriButionTi new bpecie i itribte, an lcall qite cmmn, frm nrtern Per

(Amana) twar t central Jnn. It tern limit i till nknwn.

deriVationoMinisTi new bpecie i name after Benign Calderon NoVoa (Rrige e

Mena, Amana, Per) w a ent me a lt f intereting material. Man f iL were critical fr cmpletin f te crrent reearc.

Leodonta tagaste peruana n. ssp.(g. 11-13)

typeserieshltpe: male, Per Cc/ Ra Acjanac-Pillcpata/ ~ 1308s 7125W/

m. 2100-2500 xI-xII.2004/ lg. lcal cllectr, preentl in MBLI, will be e -pite in MusM; Paratpe (65 male, 6 female): 2 male, Per dept. Cc/Qincemil/ ~ 1313s 7045W/ m. 1000 II.1998/ lg. lcal cllectr; 6 male, 1female, Per Cc/ Ri Cipata/ m. 1500-2000, I.1999, lg. lcal cllectr; 7male, Per Cc/ Ra Acjanac-Pillcpata/ ~ 1308s 7125W/ m. 2100-2500

xI-xII.2004/ lg. lcal cllectr; 15 male, Per Cc/ Inkatamb/ ~ 1303s7225W/ m. 2150-2200 III.2005/ Lg. J Bettger; 14 male, 4 female, Per

Cc/ Carrial (hapata)/ ~ 1305s 7223W/ m. 3200-3250 III.2005/ Lg.

J Bettger; 1 male, Per Cc/ Carrial (hapata)/ ~ 1305s 7223W/m. 3000 III.2006/ Lg. J Bettger; 10 male, Per Cc/ Amparae/ ~ 1317s7156W/ m. 2800 III.2005/ Lg. J Bettger, all in MBLI; 1 female, Abra Malagaver Qillabamba, 3200/3400m (Cc) 18/5/2003; 1 male, Qebraa san Li, via aQillabamba, 3000/3200m (Cc)14/5/2003; 2 male, Abra Malaga ver Qillabambakm 24 26 3000/3100m (Cc)26/2/2005; 2 male, Abra Malaga ver Qillabambakm27 2900m (Cc) 26/2/2005; 3 male, 6km N e Amparae, Calca ver Qebraahna km62,5, 3100m (Cc)17/5/2005; 2 male, 6km N e Amparae, via a Qebraahna, 3200m (Cc)15/5/2003, all in PBPF

diagnosisTi new bpecie i caracterie b male aving a wie black marginal hWd

ban ccping abt alf f te pace frm te rigin f vein M3

t te ege f tewing. hWd marginal black ban i eparate frm ical wite area b a ark banffe wit wite cale; ti grei ban i apprimatel 5 mm wie in cell 2Aan becme prgreivel narrwer in cell 1A-C

2-C

1-M

3- M

2-M

1. hWV ical

ban wit ellw pt in cell M2-M

3-C

1trianglar, nl tcing eac ter at a

pint, tat in cell M2

being at leat twice te ie f te in cell M3-C

1. Female

ave a wier black ban tan male.


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distriButionPreentl, te new bpecie i nl knwn t ccr in Cc; n pecimen

frm eiter Aacc r tern Jnn are knwn t me wic cl etabli itnrtern limit.

Leodonta tagaste extrema n.ssp.(g. 14-16)

typeserieshltpe: male, Blivia, ynga e Crani, 2500 m., 2.x.1953, leg. W. Frter,

in zsM; Paratpe (80 male, 9 female): 1 male, PerPn dept./Near Para/ ~1409s 6939W/ m. 3400-3600 xI.2004/Lg. J Bettger; 8 male, Per Pn

dept./ Carcel Pnc (sania)/ ~ 1403s 6939W/ m. 2700 V.2005/ Lg. JBettger; 21 male, 3 female, Per Pn dept./ Carcel Pnc (sania)/ ~ 1403s6939W/ m. 2500-2600 III.2006/ Lg. J Bettger; 3 male, Per Pn dept./Paccani (sania)/ ~ 1404s 6941W/ m. 2200 x.2004/ Lg. J Bettger; 36 male,2 female, Per Pn dept./ Aqele (sania)/ ~ 1404s 6942W/ m. 2100-2200

xI.2004/ Lg. J Bettger; 6 male, 1 female, Blivia Ccabamba, near CritalMa, ~ 1711s 6536W, m. 1000-1500 VII-VIII.2006, all in MBLI; 1 male,Blivia, ynga e Arepc, sienca, 2200-2500 m., 20.Ix.1953, leg. W. Frter;1 female, Blivia, ynga e Palmar, 2000 m., 3.5.50, leg. zicka; 1 female, Blivia,ynga e Palmar, 2000 m., leg. R. scnfeler, all in zsM; 1 female, Limbani ver

Carcel Pnc km4, 3000m (Pn) 16/10/2004; 1 female, Carcel Pnc, 2600m (Pn)3/2006; 1 male, 1 female, Ccabamba ver Villa Tnari km77, 2900m (Ccabamba)Blivia 18/11/2001, all in PBPF; 1 male, Lctal, via Ccabamba-VillaTnari, Blivia,30.08.2000, T.Prc leg., in MzuJ.

diagnosisTi new bpecie i caracterie b male aving a black marginal hWd

ban ccping le tan ne-tir f te pace frm te rigin f vein M3

t te egef te wing. hWd marginal black ban i eparate frm ical wite area b a arkban ffe wit wite cale; ti grei ban i apprimatel 4-5 mm wie in

cell 2A an becme mc tinner in cell 1A-C2-C1-M3-M2-M1. hWV ical banwit ellw pt in cell M

2-M

3-C

1trianglar, nl tcing eac ter at a pint,

tat in cell M2

being at leat twice te ie f te in cell M3-C

1. Female ave a

wier black ban tan male.

distriButionTe new bpecie i fn frm Pn (Per) twar t central Blivia.

acknowledgMentsI wi epre m incere tank t Pierre Boyer(Le P ste. Rparae, France),

Tma Pyrcz (Waraw, Plan), Jan Wojtusiak(Mem zlgicne uniwer-


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tet Jagiellkieg, Krakw, Plan), Pilip J. Ackery an Kim Goodger(depart-ment f Entmlg, Te Natral hitr Mem, Lnn, Englan), Ael hausMann(zlgice sammlng e Baericen staate, Mnic, German), Wlfram Mey(zlgice Mem, hmblt univeritt,Berlin, German); Critp huser(staatlice Mem fr Natrkne, stttgart, German), Gabriel Rodrguez (Envi-ga, Clmbia), Jean Frani le CroM (Bgt, Clmbia), stpane attal (Pari,France). Finall I wi t epre m gratite t Bb wortHy (srre, Englan) fr ikin aitance wit te Engli langage, an t stanilavaBadjieV (sa, Blgaria):we tarte wrking tgeter n L, bt e wa frce t abann te prjecte t circmtance ben i cntrl.

REFERENCEs

aBadjieV, s. P., 2006. Tpe f Netrpical Pierinae in te cllectin f department f Entmlg, Natralhitr Mem, Lnn (Lepiptera: Pieriae). ztaa,1143: 1218.

BraBy, M. F., pierce, N. E., Vila, R., 2007. Plgen an itrical bigegrap f te btribe Apriina(Lepiptera: Pieriae): implicatin fr te rigin f Atralian btterie. Bil. Jrn. Linn. sc.,90: 413440.

daBrera, B., 1984. Btterie f st America. hill he, Fern Creek, 256 pp.deVries, P. J., 1987. Te Btterie f Cta Rica an teir natral itr. Papiliniae, Pieriae,

Nmpaliae. Princetn univerit Pre, Princetn, ii + 328 pp., 50 cl. pl.eVenHuis, N. L., 2008. Inect an spier Cllectin f te Wrl. Available frm: ttp://b.bipmem.

rg/cen/cen-r-.tml; available nw.laMas, G., 2004. Pieriae. In: laMas, G. (E.), Cecklit: Part 4A.heperiieaPapiliniea. In: Heppner,

J.B. (E.), Atla f Netrpical Lepiptera. Vlme 5A. Aciatin fr Trpical Lepiptera/scientic Pblier, Gaineville, 99117.laMas, G., Bollino, M., 2004. Reviinal nte n te mri grp f Cic Butler, 1870

(Lepiptera: Pieriae), wit ecriptin f new pecie an bpecie. ztaa, 605:1-19lecroM, J. F., llorente, J. E., constantino, L. M., salazar. J. A., 2004. Maripa e Clmbia. Tm

2: Pierie. Bgt.talBot, G., 1932. Pieriae. Lepipterrm Catalg, 53: 1-320.VenaBles, B. a. B., 1993. Plgen f te wite an lpr btterie (Pieriae). unpblie mancript.

75 pp.zelada, W. h., 2004. La maripa irna (Lepiptera: heperiiea Papiliniea) el Bqe e

Ca, Aabaca, Pira, Per. Revita perana.Entml.,44: 37-41, 1 g., 1 tab.

bollino_2010_leodonta tagaste

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