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76 FieldBryology No119 | May18

The BBS and Data Protection RegulationsThanks to all members who have returned the “contact preferences” form (enclosed with the previous Field Bryology, or available on the website). Please would the rest do so. Permission to contact members directly allows the BBS to send information on all matters which may be of interest.

Data protection rules do not restrict any normal administrative functions and the BBS will continue to carry these out as normal. It is also a legitimate interest of the Society to use your personal information to enable supply of the Society Bulletin, Field Bryology, and the Journal of Bryology, both of which are entitlements of membership. The BBS compliance with the General Data Protection Regulations is detailed in the BBS Privacy Policy, posted on the website (or available as a print-out on request).

The BBS now has “data processing agreements” in place with both Latimer Trend (which prints and mails Field Bryology) and Taylor & Francis (the journal publisher) by which they undertake to keep BBS members’ data securely (both receive lists of your names and addresses). There is now an option for members to opt out of receiving the printed ‘hard copy’ of the Journal of Bryology; if you want the journal on-line only, you need to indicate your preference on the form.

All members have on-line access to the journal, both current issues and all back numbers; if you haven’t got this set up and would like to do so please contact the membership secretary: Mr P. Martin, 60 West Street, Tetbury, Gloucestershire, GL8 8DR (e [email protected]).

Rachel Carter, General Secretary, September 2018

From the new editorsAs Silvia Pressel and Jo Wilbraham said in the last issue, we have taken over from them as editors and must start by thanking them for the work they have done over the last few years. We also thank Silvia for helping us put together this issue, and especially for stepping in at short notice to complete it when Mari had an unexpected opportunity to spend some months in the Antarctic with BAS. Having previously been readers who have admired from afar the polished product which Jo and Silvia have produced, it is rather intimidating to take on the responsibility ourselves.

Howard Matcham has stood down as a proof reader of Field Bryology, and we thank him warmly for his service to the Society. We also thank Mark Lawley, who will continue as a proof reader, and Agneta Burton and Sue Grahame for kindly offering to join the team.

Contributions for the next issue of Field Bryology should be sent to [email protected] by 1 March 2019. Chris Preston & Mari Whitelaw

News and Society Business

76 FieldBryology No120 | Nov18

FieldBryology No119 | May18 77FieldBryology No120 | Nov18

General Referee: R.J. Fisk

HornwortsAnthoceros D.G. LongPhaeoceros D.G. Long

LiverwortsAcrobolbus G.P. RotheroAdelanthus S.D.S. BosanquetAnastrepta S.D.S. Bosanquet

Anastrophyllum D.G. LongAneura D.G. LongAnthelia G.P. RotheroAphanolejeunea N.G. HodgettsAthalamia D.G. LongBarbilophozia T.H. BlackstockBazzania T.H. BlackstockBlasia D.G. LongBlepharostoma T.H. BlackstockCalypogeia S.D.S. Bosanquet

Referee service – a helpline for identificationSome members may not be aware of the refereeing service that the BBS provides. It is intended to be a helpline to members who have difficulty in naming their British and Irish bryophytes. The referees will be delighted to help. It is better to send specimens rather than photographs, but please avoid sending large numbers of specimens at any one time. You should include collection details for the specimen(s), ideally including habitat, locality, altitude, and a grid reference (as precise as possible).

It is worth taking care in packing your specimen(s). Let them dry at air temperature before sending them. However oil-bodies are important for some leafy liverworts (e.g. Scapania, Lejeunea), and in such cases it is a good idea to separate out a few moist stems into a small plastic bag or tube (assuming of course that your specimen is still fresh). Many specimens will travel happily in a normal envelope, but many others will need protection from crushing in the post. This is particularly important for bryophytes growing on soil, such as Riccia species and other colonists of bare earth. If sent without protection, the soil and specimen will arrive as a pile of dust! Use small rigid boxes or tins, and it is a good idea to pad the specimen with tissue paper. Mini-packets within the main packet are useful for separating off plants of the problem species, for example stems bearing tubers or bulbils.

When posting packets, remember that even small envelopes will be classed as “large letters” if they exceed 5 mm thickness, and as “packets” if they exceed 25 mm. Enclose a stamped, addressed envelope (or label) with your specimen(s) if you would like them to be returned. However it is not practical or necessary to enclose a pre-paid envelope if the referee is based in a different country from yourself.

The General Referee will help beginners who are having difficulty in placing their material in a genus. If you are reasonably confident about the genus but not the species, please send your gathering to the appropriate referee, as listed below. If you encounter any other problems, refer to the respective recorder, Tom Blockeel for mosses, and Nick Hodgetts for hornworts and liverworts.

The names of genera in the following list follow the current Census Catalogue (Hill et al., 2008), with the addition of Crossidium, Dicranoloma, Ephemerella and Oxystegus.

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Cephalozia T.H. BlackstockCephaloziella (see note on p. 81 below)Chiloscyphus S.D.S. BosanquetCladopodiella T.H. BlackstockCololejeunea N.G. HodgettsColura N.G. HodgettsConocephalum D.G. LongDiplophyllum D.G. LongDouinia D.G. LongDrepanolejeunea N.G. HodgettsDumortiera D.G. LongEremonotus S.D.S. BosanquetFossombronia D.G. LongFrullania N.G. HodgettsGeocalyx G.P. RotheroGongylanthus S.D.S. BosanquetGymnocolea T.H. BlackstockGymnomitrion G.P. RotheroHaplomitrium D.G. LongHarpalejeunea N.G. HodgettsHarpanthus G.P. RotheroHerbertus D. BellHeteroscyphus S.D.S. BosanquetHygrobiella T.H. BlackstockJamesoniella S.D.S. BosanquetJubula N.G. HodgettsJungermannia S.D.S. BosanquetKurzia T.H. BlackstockLeiocolea S.D.S. BosanquetLejeunea N.G. HodgettsLepidozia T.H. BlackstockLeptoscyphus S.D.S. BosanquetLiochlaena S.D.S. BosanquetLophocolea S.D.S. BosanquetLophozia S.D.S. BosanquetLunularia D.G. LongMarchantia D.G. LongMarchesinia N.G. HodgettsMarsupella G.P. RotheroMastigophora T.H. BlackstockMetzgeria D.G. LongMicrolejeunea N.G. HodgettsMoerckia D.G. LongMylia G.P. RotheroNardia T.H. Blackstock

Nowellia T.H. BlackstockOdontoschisma T.H. BlackstockPallavicinia D.G. LongPedinophyllum S.D.S. BosanquetPellia D.G. LongPetalophyllum D.G. LongPlagiochila S.D.S. BosanquetPleurocladula T.H. BlackstockPleurozia D.G. LongPorella N.G. HodgettsPreissia D.G. LongPtilidium T.H. BlackstockRadula N.G. HodgettsReboulia D.G. LongRiccardia D.G. LongRiccia D.G. LongRicciocarpos D.G. LongSaccogyna G.P. RotheroScapania D.G. LongSolenostoma S.D.S. BosanquetSouthbya S.D.S. BosanquetSphaerocarpos D.G. LongSphenolobopsis S.D.S. BosanquetTargionia D.G. LongTelaranea T.H. BlackstockTetralophozia S.D.S. BosanquetTrichocolea T.H. BlackstockTritomaria S.D.S. Bosanquet

MossesAbietinella T.L. BlockeelAcaulon D.F. ChamberlainAchrophyllum T.L. BlockeelAloina D.F. ChamberlainAmblyodon N.G. HodgettsAmblystegium T.L. BlockeelAmphidium G.P. RotheroAndreaea M.O. HillAnoectangium D.F. ChamberlainAnomobryum D.T. HolyoakAnomodon R.J. FiskAntitrichia R.J. FiskAongstroemia G.P. RotheroAphanorrhegma N.G. HodgettsAplodon N.G. Hodgetts



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Archidium G.P. RotheroArctoa G.P. RotheroAtrichum M.O. HillAulacomnium N.G. HodgettsBarbula D.F. ChamberlainBartramia N.G. HodgettsBlindia N.G. HodgettsBrachydontium N.G. HodgettsBrachytheciastrum T.L. BlockeelBrachythecium T.L. BlockeelBreutelia R.J. FiskBruchia G.P. RotheroBryoerythrophyllum D.F. ChamberlainBryum D.T. HolyoakBuxbaumia M.O. HillCalliergon T.L. BlockeelCalliergonella T.L. BlockeelCalomnion T.L. BlockeelCalyptrochaeta T.L. BlockeelCampyliadelphus T.L. BlockeelCampylium T.L. BlockeelCampylophyllum T.L. BlockeelCampylopus G.P. RotheroCampylostelium N.G. HodgettsCatoscopium N.G. HodgettsCeratodon G.P. RotheroCheilothela G.P. RotheroCinclidium T.W. OttleyCinclidotus T.L. BlockeelCirriphyllum T.L. BlockeelClimacium R.J. FiskConardia T.L. BlockeelConostomum N.G. HodgettsCoscinodon T.L. BlockeelCratoneuron T.L. BlockeelCrossidium D.F. ChamberlainCryphaea R.J. FiskCtenidium G.P. RotheroCyclodictyon T.L. BlockeelCynodontium G.P. RotheroDaltonia T.L. BlockeelDendrocryphaea R.J. FiskDialytrichia D.F. ChamberlainDichodontium G.P. RotheroDicranella G.P. Rothero

Dicranodontium G.P. RotheroDicranoloma T.L. BlockeelDicranoweisia G.P. RotheroDicranum G.P. RotheroDidymodon T.L. BlockeelDiphyscium M.O. HillDiscelium N.G. HodgettsDistichium G.P. RotheroDitrichum G.P. RotheroDrepanocladus M.O. HillEncalypta N.G. HodgettsEntodon R.J. FiskEntosthodon N.G. HodgettsEphemerella N.G. HodgettsEphemerum D.T. HolyoakEpipterygium N.G. HodgettsEucladium D.F. ChamberlainEurhynchiastrum T.L. BlockeelEurhynchium T.L. BlockeelFissidens T.W. OttleyFontinalis T.L. BlockeelFunaria N.G. HodgettsGlyphomitrium N.G. HodgettsGrimmia R.D. PorleyGymnostomum D.F. ChamberlainGyroweisia D.F. ChamberlainHabrodon S.D.S. BosanquetHageniella T.L. BlockeelHamatocaulis M.O. HillHedwigia T.L. BlockeelHelodium T.L. BlockeelHennediella D.F. ChamberlainHerzogiella N.G. HodgettsHeterocladium S.D.S. BosanquetHomalia R.J. FiskHomalothecium T.L. BlockeelHomomallium T.L. BlockeelHookeria R.J. FiskHygroamblystegium T.L. BlockeelHygrohypnum T.L. BlockeelHylocomiastrum S.D.S. BosanquetHylocomium R.J. FiskHymenostylium D.F. ChamberlainHyocomium G.P. RotheroHypnum G.P. Rothero


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Isopterygiopsis N.G. HodgettsIsothecium T.L. BlockeelKiaeria G.P. RotheroKindbergia T.L. BlockeelLeptobarbula D.F. ChamberlainLeptobryum N.G. HodgettsLeptodictyum T.L. BlockeelLeptodon R.J. FiskLeptodontium D.F. ChamberlainLeptophascum D.F. ChamberlainLeptotheca T.L. BlockeelLescuraea T.L. BlockeelLeskea R.J. FiskLeucobryum G.P. RotheroLeucodon R.J. FiskLoeskeobryum S.D.S. BosanquetMeesia N.G. HodgettsMicrobryum D.F. ChamberlainMicromitrium D.T. HolyoakMielichhoferia N.G. HodgettsMnium T.W. OttleyMolendoa D.F. ChamberlainMyrinia T.L. BlockeelMyurella N.G. HodgettsMyurium T.L. BlockeelNeckera R.J. FiskOedipodium N.G. HodgettsOligotrichum M.O. HillOncophorus G.P. RotheroOrthodontium T.L. BlockeelOrthothecium N.G. HodgettsOrthotrichum T.L. BlockeelOxyrrhynchium T.L. BlockeelOxystegus T.L. BlockeelPaludella N.G. HodgettsPalustriella T.L. BlockeelParaleptodontium D.F. ChamberlainParaleucobryum G.P. RotheroPhascum D.F. ChamberlainPhilonotis N.G. HodgettsPhyscomitrium N.G. HodgettsPlagiobryum D.T. HolyoakPlagiomnium T.W. OttleyPlagiopus N.G. HodgettsPlagiothecium N.G. Hodgetts

Plasteurhynchium T.L. BlockeelPlatydictya N.G. HodgettsPlatygyrium T.L. BlockeelPlatyhypnidium T.L. BlockeelPleuridium G.P. RotheroPleurochaete D.F. ChamberlainPleurozium R.J. FiskPogonatum M.O. HillPohlia N.G. HodgettsPolytrichastrum M.O. HillPolytrichum M.O. HillPottiopsis D.F. ChamberlainPseudephemerum G.P. RotheroPseudobryum T.W. OttleyPseudocalliergon T.L. BlockeelPseudocrossidium D.F. ChamberlainPseudoleskea T.L. BlockeelPseudoleskeella T.L. BlockeelPseudoscleropodium T.L. BlockeelPseudotaxiphyllum N.G. HodgettsPterigynandrum S.D.S. BosanquetPterogonium R.J. FiskPterygoneurum T.L. BlockeelPtilium G.P. RotheroPtychodium T.L. BlockeelPtychomitrium N.G. HodgettsPylaisia T.L. BlockeelRacomitrium T.L. BlockeelRhabdoweisia G.P. RotheroRhizomnium T.W. OttleyRhodobryum D.T. HolyoakRhynchostegiella T.L. BlockeelRhynchostegium T.L. BlockeelRhytidiadelphus S.D.S. BosanquetRhytidium G.P. RotheroSaelania G.P. RotheroSanionia M.O. HillSarmentypnum T.L. BlockeelSchistidium S.D.S. BosanquetSchistostega N.G. HodgettsSciuro-hypnum T.L. BlockeelScleropodium T.L. BlockeelScopelophila D.F. ChamberlainScorpidium T.L. BlockeelScorpiurium T.L. Blockeel



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Seligeria D.T. HolyoakSematophyllum T.L. BlockeelSphagnum M.O. HillSplachnum N.G. HodgettsStegonia D.F. ChamberlainStraminergon T.L. BlockeelSyntrichia D.F. ChamberlainTaxiphyllum N.G. HodgettsTayloria N.G. HodgettsTetraphis M.O. HillTetraplodon N.G. HodgettsTetrodontium M.O. HillThamnobryum R.J. FiskThuidium T.L. BlockeelTimmia N.G. HodgettsTomentypnum M.O. HillTortella D.F. ChamberlainTortula D.F. ChamberlainTrematodon G.P. RotheroTrichodon G.P. RotheroTrichostomum T.L. BlockeelUlota T.L. BlockeelWarnstorfia M.O. HillWeissia D.F. ChamberlainZygodon T.L. Blockeel

Note: DNA work at the Royal Botanic Garden, Edinburgh (Bell et al., 2013) had shown that our current understanding of the genus Cephaloziella is inadequate and a complete revision is needed before most of the species can be identified reliably.

Contact details

Mr D. Bell, Royal Botanic Garden, Inverleith Row, Edinburgh EH3 5LR. e: [email protected] T.H. Blackstock, Eirianallt, Glanrafon, Llangoed, Anglesey, LL58 8PG. e: [email protected] T.L. Blockeel, 9 Ashfurlong Close, Dore, Sheffield, S17 3NN. e: [email protected].

Mr S.D.S. Bosanquet, Dingestow Court, Monmouth, Monmouthshire, NP25 4DY. e: [email protected]. Twitter: @sambbryo.Dr D.F. Chamberlain, Royal Botanic Garden, Inverleith Row, Edinburgh, EH3 5LR. e: [email protected] R.J. Fisk, 35 Fair Close, Beccles, Suffolk, NR34 9QR. e: [email protected] M.O. Hill, 11 Chaucer Road, Cambridge, CB2 7EB. e: [email protected] N.G. Hodgetts, 15 Earlish, Portree, Isle of Skye, IV51 9XL. e: [email protected] D.T. Holyoak, Quinta da Cachopa, Rua da Barcoila no. 274, 6100-014 Cabeçudo, Portugal. e: [email protected] D.G. Long, Royal Botanic Garden, Inverleith Row, Edinburgh, EH3 5LR. e: [email protected] T.W. Ottley, Valley Forge, Goginan, Aberystwyth, SY23 3PD. e: [email protected] R.D. Porley, Cerca dos Pomares, CxP 409M, 8670-052 Aljezur, Portugal. e: [email protected] G.P. Rothero, 8 Deer Park, Glenmassan, Dunoon, Argyll, PA23 8RA. e: [email protected].

ReferencesBell, D., Long, D.G. & Hollingsworth, P.M. (2013). The use of DNA barcoding to address major taxonomic problems for rare British bryophytes. Royal Botanic Garden, Edinburgh.Hill M.O., Blackstock T.H., Long D.G. & Rothero, G.P. (2008). A checklist and census catalogue of British and Irish bryophyte, updated 2008. British Bryological Society, Middlewich.

Tom Blockeel & Nick Hodgetts


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364 Atlas of British & Irish Bryophytes (1)

Sphagnum palustre

Britain 1990–2013 1358 1950–1989 337 pre-1950 42

Ireland 1990–2013 332 1950–1989 122 pre-1950 2

oose cushions and patches of Sphagnum palustre are found in a wide variety of habitats, including wet woods,

boggy grassland, ditches, flushed peaty banks, marshes and streamsides. Unlike S. papillosum, it is tolerant of shade and is sometimes abundant in damp conifer plantations and swampy carr. It is one of the less acid-demanding sphagna, growing with S. fimbriatum, S. squarrosum and S. subnitens. It is also common on oceanic blanket bogs, where it occupies small declivities receiving surface flow in wet weather. Altitudinal range: 0–1100 m.

Dioicous; capsules are occasional, August.

Plants in our area are mostly var. palustre. Var. centrale is often treated as a separate species, S. centrale, but has proved very difficult to distinguish because some collections

have leaf cross-sections of intermediate and unstable shape, differing between leaves on the same plant. Var. centrale has been found in four places in England, two in Wales, two in the Isle of Man, and as two separate gatherings from the Morrone Birkwood near Braemar in eastern Scotland. The Morrone locality is at about 500 m altitude and is the only one that fits with its distribution in Eurasia and North America, where it has a continental distribution, penetrating continental interiors from which var. palustre is absent. Genetic analysis in North America (Karlin et al., 2010) indicates that there is a clear distinction between the two taxa. Photos in Flatberg (2013) and Hölzer (2010) show S. centrale as remaining greenish or yellowish in autumn, and not turning pinkish as is normal for var. palustre.

Circumpolar Boreo-temperate. Widespread in Europe except the south and far north. Northern Asia and N America, mainly as var. palustre in coastal areas, mainly as var. centrale in continental interiors.

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SPECIAL OFFER –Atlas of British and Irish BryophytesMembers may be interested to know that the 2-volume Atlas of British and Irish Bryophytes, published in 2014, is now available at a reduced price of £50 plus p&p (full price £75 plus p&p) during the pre-Christmas period. This is an opportunity not to be missed if you do not

already have a copy. All 1069 species of hornworts, mosses and liverworts known from Britain and Ireland at the time of publication are mapped in colour. The Atlas incorporates the results of 20 years’ fieldwork by members of the British Bryological Society. The hectad (10 × 10 km square) maps reveal dramatic changes in the distribution of some species since 1990. Each map is accompanied by a detailed text covering habitat, reproductive biology, significant range changes, taxonomic or recording problems and world distribution. The altitudinal distribution of most species in Britain is illustrated by a novel diagram, and the species accounts are illustrated by numerous habitat photographs. The introductory chapters describe the history of the BBS recording scheme, highlighting the effect of technological changes since 1990, and map environmental factors relevant to the interpretation of the maps. An important chapter presents a new analysis of the records to identify changes in frequency in recent decades in response to drivers such as air pollution, land-use and climate change.

To order at the reduced price visit Nature Bureau’s website: https://www.naturebureau.co.uk/bookshop/special-offers

An overview of the BBS meeting programme for 2019 is given below. Many areas also have BBS local group meetings and details of these can also be found on the BBS website.

BBS Spring Meeting North Devon, 10–17 April 2019North Devon (vc 4) is a large and varied area, with habitats ranging from the heights of Exmoor and Dartmoor to the nationally famous sand dunes of Braunton Burrows and the impressive cliffs of the north and west coasts. There is also a lot of potentially interesting country inland, much of which has still to be properly checked for its bryophytes. The headquarters for the week is Hallsannery, approximately a mile south of Bideford. It consists of a large country house in its own grounds, and has been booked for the exclusive use of the BBS for the week. A full description is given on the Hallsannery website (www.hallsannery.co.uk); anyone interested is advised to look at the floor plans (follow “About” and then “Links”), as the one disadvantage is that there are relatively few single rooms. As the premises have already been reserved for us, individual reservations should be made through the Local Secretary, Mark Pool, who will also be able to provide further information if needed. The cost per person has not yet been finalised, but is likely to be around £35 per night. For anyone not wishing to stay at the headquarters, there is a variety of suitable accommodation in and around Bideford.

Local Secretary: Mark Pool (91 Warbro Road, Torquay, TQ1 3PS; email [email protected]; phone 01803-316154).

BBS Summer meeting, June–July 2019 Week 1: County Wicklow, 29 June to 6 July 2019. Week 2: Iveragh Peninsula, County Kerry, 7–14 July 2019.

AGM, Autumn 2019University of Leicester Botanic Garden, 4–6 October 2019.Further information on the summer meetings and the AGM will be published on the website as soon as it is available.

Would you like to host a meeting in your county or area? Please get in touch with Lucia Ruffino ([email protected]).

BBS workshop 2019

Acrocarp Identification Workshop, 8–10 February 2019Preston Montford Field Studies Centre, Shrewsbury, 8–11 February 2019.Tutor: Sharon Pilkington.A repeat of a very popular workshop held in 2013 and 2016, this course will enable beginners and improvers to learn how to identify many widespread acrocarpous mosses. An intensive mixture of lectures, microscope exercises and a short excursion will provide opportunities to study acrocarps in depth. Workshop participants will also hone field skills and master some of the trickier microscope techniques required to identify species with confidence. Numbers will be limited to 12. The cost will be £175 for a shared room or £195 single occupancy. This includes accommodation and all meals from Friday evening until Sunday afternoon.Please e-mail Sharon Pilkington ([email protected]) to book a place on this course.

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Future Meetings and Courses

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