Acarina 27 (2): 117–144 © Acarina 2019

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IN MEMORIAM OF ANDREI VLADIMIROVICH BOCHKOV: SCIENTIFIC BIOGRAPHY

Sergey V. Mironov1, Dmitry A. Apanaskevich1,2 and Sergey G. Medvedev1

1Zoological Institute of the Russian Academy of Sciences, Saint Petersburg, Russia2United States National Tick Collection, James H. Oliver, Jr. Institute for Coastal Plain Science, Georgia Southern University, Statesboro, USA

Slightly more than a year ago, the world aca-rological and parasitological community suffered from the loss of Andrei (André) Vladimirovich Bochkov (27 August 1968—9 May 2018)—an eminent acarologist, one of the greatest experts on systematics of parasitic acariform mites—as he died on the 50th year of his life, in the prime of his scientific career. Andrei’s death is an irretrievable loss for the national and world science. His col-leagues and friends dedicate this special issue of Acarina to the memory of this outstanding scientist and extraordinary man.

The entire 32-year scientific career of Andrei Bochkov was associated with the Zoological Insti-tute of the Russian Academy of Sciences (Saint Petersburg). At this institute, he undertook his long and challenging academic path from Lab Assistant to Head Researcher, becoming one of the greatest experts in acarology.

From Laboratory Assistant to PhD (1986–1999). Andrei Bochkov was born on 28 August 1968 in Novosibirsk (Russia). In the mid-1970s, his family moved to a suburb of Leningrad (now Saint Petersburg). Since his early years, Andrei expressed great interest in biology, which was in-spired by his school teacher E.V. Kudryashov. In 1986, after graduating from high school in a small town of Taitsy (Gatchina District, Leningrad Re-gion, Russia), he came to the Zoological Institute with a naive desire “to study tigers”. However, at that time, the Institute could only offer young An-drei a more prosaic position of a lab assistant at the Department of Parasitology. Nevertheless, he was delighted by the opportunity to work at the Institute with parasites of vertebrates. Although the labora-tory assistant duties in the animal facility were not particularly exciting, later in his mature years, Andrei spoke with considerable pride of the fact that he had passed all of the steps of the scientific hierarchy, starting from the very bottom.

From the first days of his work at the Depart-ment, Andrei was not willing to limit himself with simple tasks, like technical work in the mite col-

lection and looking after animals in the vivarium. His interest in acarology prompted Andrei to ask his supervisor, Dr. H.V. Dubinina, an expert on mammal-associated acariforms and house dust mites, for an opportunity to work on a research project in this field. This request resulted in his first scientific works dealing with the systematics of myobiid mites (Myobiidae) parasitizing on murid rodents.

Parallel with his work at the Institute, he was a part-time graduate student at the Department of Entomology of the Biological Faculty in St. Peters-burg State University, from which he graduated in 1996. Later, Andrei declared that the Department is where “the real systematists are formed”. By the time of his graduation from the university, his sci-entific interest was clearly focused on the system-atics of a particular category of acariform mites—permanent parasites of mammals. He remained faithful to this group of arthropods until the end of his life. Upon graduation, while still working as a lab assistant, he independently and enthusiasti-cally began conducting research on his favorite group of mites—myobiids (Myobiidae).

Gradually, his research began to expand to other groups of parasitic mites associated with small mammals, both Astigmata and Prostigmata. In particular, under the supervision of H.V. Du-

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binina, Andrei began studying fur mites Chirodis-cidae, Listrophoridae and Myocoptidae, mange psoric mites Psoroptidae, house dust mites Pyro-glyphidae (Astigmata) and predatory mites Chey-letidae (Prostigmata). The development of Andrei’s

scientific mindset was significantly broadened by contacts and collaborations with senior researchers in the Department of Parasitology. Under the influ-ence of Dr. S.V. Mironov—a prominent expert on feather mites, who subsequently became his PhD

Fig. 1. Andrei Vladimirovich Bochkov and some parasitic mite specimens. A—searching through feathers of a live eared owl Asio otus infected by quill mites (Syringophilidae) (“Rybachy” bird banding station, Kaliningrad Region, September 2006). B—European robin Erythacus rubecula with web huts of Bakericheyla chanayi (Chetyletidae). C—working tent near Kyzylkol Lake (expedition to Kazakhstan, October 2007). D—Ornithocheyletia sp. (Cheyletidae). E—Soricilichus scutisorex (Chirodiscidae), precopulatory guarding of tritonymph by male. F—in the office at the University of Michigan, Ann Arbor, USA (September 2003).

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advisor—scientific interests of Andrei have ex-panded to several groups of mites parasitic on birds; namely, the families Syringophilidae and Harpi-rhynchidae (Prostigmata). Based on his long-term study of myobiids, in 1999, Dr. Bochkov success-fully defended his Candidate of Sciences thesis (Russian equivalent of PhD) “Mites of the family Myobiidae (Acari: Prostigmata) and their position in the system”. His thesis work resulted in a pub-lication titled “New classification of myobiid mites (Acari, Acariformes)” (Bochkov, 1997c), which is considered a significant contribution to this sub-field of acarology.

Foreign postdoc research (2000–2006). Post-doc fellowships at foreign museums and universi-ties played a huge role in Andrei’s subsequent development as a world-class scientist. They al-lowed him to work under the mentorship of world leading acarologists and to gain access to rich materials deposited in some of the largest world collections. It is necessary to give Andrei credit for his determination and perseverance: his self-admit-ted less-than-mediocre knowledge of English did not deter him from undertaking his first postdoc.

In 2000, Andrei participated in a study under the supervision of Professor Alex Fain (the Institut royal des Sciences naturelles de Belgique, Brussels, Belgium)—the greatest taxonomist-acarologist of the 20th century. The project was titled “Phylogeny and systematics of the family Cheyletidae (Acari: Cheyletoidea),—Grant from the Belgian Office for Scientific, Technical and Cultural Affairs”. For the first time, Andrei had access to the richest collec-tions of parasitic mites stored in Brussels and Tervuren (at the Musee royale de l’Afrique central, Belgium). As a result of this and other subsequent collaborative works with Professor Fain, 49 joint publications were produced, among which was the reconstruction of the phylogenetic system of predatory and parasitic mites of the family Chey-letidae (Bochkov and Fain 2001).

From 2003 to 2006, Dr. Bochkov held a three-year postdoc position at the Museum of Zoology of the University Michigan, Ann Arbor, USA, in the lab of Professor Barry M. OConnor—the great-est expert on systematics of astigmatan mites. There, Andrei participated in the project titled “Phylogeny and Historical Ecology of Mite Para-sites of Vertebrates (Acari: Dermanyssoidea and Sarcoptoidea) (PEET grant from NSF)”, in the course of which he excellently mastered the use of cladistics methods in systematics. Being an open

and communicative person, Andrei has established scientific contact with many researchers world-wide. His interests have expanded to include such mites as bee-associated cheyletids (Cheletophyes), skin-dwelling feather mites of the families Derma-tionidae and Epidermoptidae (Astigmata) and even gamasid mites (Mesostigmata: Laelapidae). The period of postdoc studies was crucial for Andrei’s development, when in less than a decade he became a world expert in the systematics and phylogeny of parasitic acariform mites. Among his most im-portant generalization works are those on the clas-sification and phylogeny of mites of the superfam-ily Cheyletoidea (Bochkov 2002); on the phylog-eny and host associations of the fur mite subgenus Listrophoroides (Atopomelidae) (Bochkov and OConnor 2005b); and, in collaboration with S.V. Mironov and A. Fain, on the phylogeny and evolu-tion of parasitism in feather mites of the families Epidermoptidae and Dermationidae (Analgoidea) (Mironov et al. 2005).

Scientific peaks (2007–2012). After the post-doc period of his life—being full of energy, enthu-siasm and creative plans—Andrei decisively fo-cused on generalization studies and the preparation of his Doctor of Sciences thesis (equivalent of a Habilitation Degree in many European countries). During six subsequent years, he published 64 works on astigmatan and prostigmatan mites associated with mammals, including three monographs: on the phylogeny and systematics of the endoparasitic astigmatids of the families Gastronyssidae, Lemur-nyssidae and Pneumocoptidae (Bochkov et al. 2008); a review of mites of the parvorder Eleuther-engona, permanent parasites of mammals (Bochkov 2009a); and a monograph on host-parasite relation-ships between the mite subgenus Microtimyobia (Myobiidae) and cricetid rodents (Bochkov 2011a). Among the most important works of this period are generalization papers on the origin and evolution of parasitism in mites of the infraorder Eleutheren-gona (Bochkov 2008c, 2009a); works on the phy-logeny and evolution of parasitism in mammal-associated psoroptidian mites (Astigmata) (Boch-kov 2011b; Bochkov and Mironov 2011); and works on the use of phylogeny and coevolutionary rela-tionships of parasites in the study of the evolution of hosts, e.g., the use of mange mites (Psoroptidae) to study strepsirrine primates (Bochkov et al. 2011). Additionally, in collaboration with S.V. Mironov, Dr. Bochkov has studied the phenomenon of syn-hospitality in parasitic acariform mites (Bochkov

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and Mironov 2008). The two authors have also generated an original macrophylogenetic system of the order Acariformes (Mironov and Bochkov 2009). In general, this period of Andrei’s career has proven to be most fruitful. Finally, in 2011, Andrei successfully defended his Doctor of Sciences thesis titled “Acariform mites, permanent parasites of mammals: phylogeny, systematic and host-parasite interactions”.

Acknowledged acarologist (2013–2018). Be-ing at the top of the acarology profession, Dr. Bochkov did not decrease the intensity of his re-search, continuing his studies in various fields until his last days. He continued working on the phylogeny and the taxonomic revisions of a practi-cally important family of mange mites, Psoroptidae, which he started back in 2009. A series of taxo-nomic works, devoted to the mites of the family Harpirhynchidae parasitizing on birds in the New World, were finalized in a taxonomic monograph (Bochkov et al. 2015).

During this period, Andrei successfully col-laborated with colleagues from the A. Mickiewicz University (Poznan, Poland) and, in particular, with a prominent expert on quill mites, Dr. M. Skoracki. Two major investigatory projects, notably, have been carried out: “Host-parasite relationships and biodiversity of prostigmatan mites (Acari: Acari-formes), permanent parasites of passeriform birds (Aves: Passeriformes) of Holarctic region” (2011–2013) and “Acariform mites (Acari: Acariformes) permanently parasitizing warm-blooded vertebrates in Sub-Saharan Africa and Madagascar: fauna genesis, phylogeny, and host-parasite relation-ships” (2013–2017). As a result, 18 works on mites of the families Syringophilidae, Myobiidae and Cheyletidae have been published. Andrei’s gen-eral investigations during this period include the works on: the species delimitation and morpho-logical diagnostics of Chorioptis mange mites (Bochkov et al. 2014); the cophylogeny of harpy-rhynchid mites and birds (Bochkov et al. 2017); and the phylogeny of house dust mites of the subfamily Guatemalichinae (Klimov et al. 2015a). Two works dating to this period present a special interest. One is his controversial review on the reversion of parasitism “Is parasitism of Metazoa a one-way ticket?” (Bochkov and Mironov 2013). The other is his latest generalizing work “Host-parasite interactions of acariform mites (Acari-formes)—permanent parasites of mammals” (Bochkov 2016).

Scientific heritage. The scientific heritage of Andrei Bochkov is rich and diverse, in terms of both mite taxa and the general scope of research topics. He is the author of 220 scientific works, including 5 large monographs (see the List of pub-lications; monographs are marked with *). In the course of his taxonomic studies, Dr. Bochkov has established 496 new mite taxa, including 7 family-group names, 43 genus-group names and 445 new mite species (see the List of taxa and Table 1). It is important to emphasize that in the overwhelming majority of his scientific publications, Andrei was the leading author, who performed essentially the entire work. But, being a polite person, Andrei often acknowledged the participation of senior and respectful co-workers by placing them ahead of him in the author list.

In the last decade, about two dozen of his articles were devoted to the reconstruction of the phylogeny of various family groups of parasitic acariform mites, including Cheyletoidea and Psoroptidia, as well as to major questions in parasitology and general biol-ogy. Unfortunately, some monographic works, such as the revision of the scabies mites of the family Psoroptidae and of the hair mites of the family Atopomelidae, were never finished.

Because of his efforts, a sizeable representa-tive world-class collection of astigmatan and prostigmatan mites parasitizing on mammals has accumulated in the Department of Parasitology of the Zoological Institute RAS. The collection contains 26 mite families, over 10,000 slides and about 900 species, including many type speci-mens. Since the early 1990s, Andrei has partici-pated in many expeditions to Central Asia, Ka-zakhstan and Kamchatka, becoming an extreme-ly active collector of valuable research material. His last expedition in 2016 was to Ethiopia, where unique material for morphological and molecular studies was collected.

Simultaneously with his intensive research, Andrei took the time to conduct other work for the benefit of science. He actively worked on editorial boards of a number of national and in-ternational scientific journals, such as Acarina, Proceedings of the Zoological Institute of the Russian Academy of Sciences, International Jour-nal of Acarology, Zootaxa and Zookeys. Largely due to his efforts as the deputy editor, the Russian journal Acarina has become one of the leading research journals in the field of acarology in the world.

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Being an extremely sociable person, Andrei has established contacts with many zoologists engaged in the collection of host-animals of his mites not only in Russia, but also in other countries. Despite his challenging character, straightforwardness and at times quickness of judgment, Andrei was an extremely benevolent and responsive person both in professional and personal life. A man of sharp intellect, Andrei was fiery and at times harsh in his scientific argumentation. However, he could hon-estly and openly admit his errors. He was ex-tremely communicative and easily made contacts with people of all ages, positions and social circles. Known for his cheerfulness, optimism and (at times rough) sense of humor, Andrei could liven up any company, be it in or outside of the workplace.

It is safe to say that Andrei was one of the most “famous” employees of the Zoological Institute. In addition to his exceptional sociability and unique personal charm, Andrei possessed solid authority on many scientific topics that went beyond the mite systematics. In early 2000s, he became one of the most active specialists who contributed to the de-velopment of cladistic methods in systematics. Andrei even organized a short course of lectures on his approach at the Zoological Institute.

Andrei left us in the prime of his scientific career, maintaining youthful enthusiasm and admi-rable work ethic until his very last days, despite the severe illness. We will keep the memory of Andrei in our hearts—an extraordinary bright person, a special friend and an outstanding scientist.

BIBLIOGRAPHY LIST OF SCIENTIFIC WORKS BY A.V. BOCHKOV

1989

1. Dubinina, H.V. and Bochkov, A.V. 1989. [New data on ectoparasites of the Royle’s pika in Kirghizia]. Entomologicheskie issledovania v Kirghizii, 20: 107–116. [In Russian]

1990

2. Bochkov, A.V., Dubinina, H.V. and Chirov, P.A. 1990. [A first record of mites of the subgenus Aus-tromyobia (Acariformes, Myobiidae) in the USSR fauna]. Izvestiya AN Khirgizskoy SSR (khimiko-tekhnologicheskie nauki), 1: 55–66. [In Russian]

3. Efremova, G.A. and Bochkov, A.V. 1990. [Mites of families Cheyletidae and Cheyletiellidae (Acari-formes: Trombidiformes) of the Byelorussian fau-na]. Vesti AN BSSR, 6: 98–103. [In Byelorussian]

1991

4. Mironov, S.V., Bochkov, A.V. and Chirov, P.A. 1991. [Mites of the genus Cheletopsis (Acariformes, Chey-letidae) from the quills of charadriiform birds from Middle Asia]. Izvestiya AN Khirgizskoy SSR (khimiko-tekhnologicheskie nauki), 4: 50–56. [In Russian]

1992

5. Bochkov, A.V. and Rybin, S.N. 1992. [A new species Ewingana teniotis (Acariformes: Myobiidae) from Tadarida teniotis from Kirghizia]. Parazitologiya, 26 (4): 333–338. [In Russian with English summary]

1993

6. Dubinina, E.V., Bochkov, A.V. and Bobrovskaja, V.I. 1993. [Notes on systematics of mites of the genus Schizocarpus (Acariformes: Chirodiscidae)]. Parazitologiya, 27 (6): 450–453. [In Russian with English summary]

1994

7. Bochkov, A.V. 1994a. [Data on mites of the genus Protomyobia (Acariformes: Myobiidae) of Russia and neighboring countries]. Parazitologiya, 28 (2): 137–146. [In Russian with English summary]

8. Bochkov, A.V. 1994b. [Mites of the subgenus Graphiurobia (Myobiidae, Radfordia) of the fauna of Russia and surrounding states]. Parazitologiya, 28 (5): 421–428. [In Russian with English summary]

9. Bochkov, A.V., Mironov, S.V. and Gorgol, V.T. 1994. Ornithocheyletia phylloscopi sp. n. (Acariformes: Cheyletidae), new species of parasitic mites from the willow warbler. Acarina, 2 (1–2): 73–80.

10. Kivganov, D.A. and Bochkov, A.V. 1994. [A new mite species of the genus Cheletopsis (Acari, Cheyletidae)]. Vestnik Zoologii, 2: 39–43. [In Rus-sian with English summary]

1995

11. Bochkov, A.V. 1995. [Mites of the subgenus Mi-crotimyobia (Acariformes, Myobiidae, Radfordia) found of the Russia and surrounding countries]. Parazitologiya, 29 (6): 480–492. [In Russian with English summary]

12. Dubinina, H.V., Bochkov, A.V. and Javrujan, J. G. 1995. Mites of the family Myobiidae (Acari-formes)—ectoparasites of bats of Armenia. Aca-rina, 3 (1–2): 99–104.

1996

13. Bochkov, A.V. 1996a. Hesperomyobia (Acari: Myobiidae: Radfordia), a new subgenus of myo-

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biid mites from rodents of the family Hesperomy-idae (Rodentia). Acarina, 4 (1–2): 39–42.

14. Bochkov, A.V. 1996b. Mites of the family Myobi-idae (Acariformes: Prostigmata) parasitizing on bats (Mammalia: Chiroptera) of the fauna of the former USSR. Acarina, 4 (1–2): 3–26.

15. Bochkov, A.V. and Malikov, V.G. 1996. Eucheyle-tiella faini sp. n. (Acari: Cheyletidae), a new spe-cies of parasitic mites from Ochotona rufescens (Gray) (Lagomorpha, Ochotonidae). Acarina, 4 (1–2): 43–48.

16. Dubinina, H.V. and Bochkov, A.V. 1996. Pleisto-cene mites (Acariformes) from the narrow-skulled vole Microtus gregalis egorovi Feigin (Rodentia: Arvicolinae). Acarina, 4 (1–2): 35–38.

1997

17. Bochkov, A.V. 1997a. [Description of the female of the mite Radfordia (Austromyobia) aegyptica (Acariformes: Myobiidae)]. Parazitologiya, 31 (5): 458–461. [In Russian with English summary]

18. Bochkov, A.V. 1997b. [Myobiid mites (Acari-formes: Myobiidae) parasiting a rodent family Muridae (Rodentia) in Russia and neighbouring countries]. Parazitologiya, 31 (3): 201–209. [In Russian with English summary]

19. Bochkov, A.V. 1997c. [New classification of myobiid mites (Acari, Acariformes)]. Entomo-logicheskoe obozrenie, 76 (4): 938–951. [In Rus-sian with English summary] [More correct trans-lation of the title: new classification of mites of the family Myobiidae (Acari, Acariformes)]

20. Bochkov, A.V. 1997d. [Two new subgenera of myobiid mites of the genus Radfordia Ewing (Acariformes, Myobiidae)]. Entomological Re-view, 76 (3): 684–704. [In Russian with English summary]

21. Bochkov, A.V. 1997e. Mites of the family Myo-biidae (Acariformes: Prostigmata) parasitizing on Lipotyphla (Mammalia: Insectivora) of the fauna of former USSR. Acarina, 5 (1–2): 45–62.

22. Bochkov, A.V. and Fain, A. 1997. On a systemat-ics of mites of the subgenus Radfordia s.str. (Acari: Myobiidae: Radfordia). Acarina, 5 (1–2): 19–28.

23. Bochkov, A.V. and Mironov, S.V. 1997. On tax-onomy of predatory mites of the genus Neoeuchey-la Radford, 1950 and related genera (Acari: Cheyletidae). Acarina, 5 (1–2): 73–78.

24. Chirov, P.A., Bochkov, A.V. and Rybin, S.N. 1997. [The mites of the family Myobiidae from Kir-ghizia]. Entomologicheskie issledovaniya v Kir-gizii, 21: 42–50. [In Russian]

1998

25. Bochkov, A.V. and Mironov, S.V. 1998a. [On a systematics of mites of the subgenus Microtimyo-bia (Acariformes: Myobiidae: Radfordia) and their distribution on voles (Rodentia: Cricetidae: Arvi-colinae)]. Parazitologiya, 32 (4): 300–316. [In Russian with English summary]

26. Bochkov, A.V. and Mironov, S.V. 1998b. Quill mites of the family Syringophilidae Lavoipierre, 1953 (Acariformes: Prostigmata) parasitizing on birds (Aves) of the fauna of the former USSR. Acarina, 6 (1–2): 3–16.

27. Bochkov, A.V. and Mironov, S.V. 1998c. Samsina-kia trilobitus sp. n., a new cheyletid mite from South India (Acari, Cheyletidae). Entomologische Mitteilungen aus dem Zoologischen Museum, Hamburg, 12: 265–268.

28. Chirov, P.A. and Bochkov A.V. 1998. Ornithochey-letia mironovi sp. n. (Acari: Cheyletidae), a new species of parasitic mite from the bank swallow from Kirghizia. Acarina, 6 (1–2): 35–36.

1999

29. Bochkov A.V. 1999a. [A new myobiid mites spe-cies Radfordia cricetuliphila sp. n. (Acari: Myobi-idae) from the striped hamster Cricetulus baraben-sis (Rodentia: Cricetidae) from Buryatia]. Parazi-tologiya, 33 (4): 358–363. [In Russian with English summary]

30. Bochkov, A.V. 1999b. [The system and host-para-sites relationships of the mites of the subfamily Myobiinae (Acariformes: Myobiidae)]. Entomo-logicheskoe obozrenie, 78 (3): 764–776. [In Rus-sian with English summary; also translated into English as: Classification and host-parasite rela-tions of mites of the subfamily Myobiinae (Acari-formes, Myobiidae), Entomological Review, 79 (6): 666–675.]

31. Bochkov A. V. 1999c. [Mites of the family Myo-biidae (Acari: Prostigmata) and their position in the system]. Candidate of Science thesis. Zoo-logical Institute RAS, Saint Petersburg. 480 pp. [Manuscript, in Russian]

32. Bochkov, A.V. and Haustov, A.A. 1999. Proso-cheyla villosa sp. n., a new mite species from Crimea (Prostigmata: Cheyletidae). Genus, 10 (1): 151–154.

33. Bochkov, A.V. and Khaustov, A.A. 1999. [Descrip-tion of Cheyletus kuznetzovi sp. n. (Acariformes: Cheyletidae) from Turkmenia]. Zoologicheskii Zhurnal, 78 (1): 115–116. 302 [In Russian with English summary; also published in English in Entomological Review, 79 (4): 482–483.]

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34. Bochkov, A.V., Khaustov, A.A. and Kuznetzov, N.N. 1999. [A new mite species of the genus Chelacheles (Acariformes: Cheyletidae) from the Crimea]. Zoologicheskii Zhurnal, 78 (1): 117–120. [In Russian with English summary; also published in English in Entomological Review, 79 (4): 484–487.]

35. Bochkov, A., Malikov, V. G. and Arbobi, M. 1999. Trichoecius calomysci sp. n. (Acari: Myocoptidae), a new mite species from Iran. Folia Parasitologi-ca, 46: 316–318.

36. Bochkov, A.V. and Mironov, S.V. 1999a. A system-atic review of the parasitic mite genus Eucheyletia Volgin, 1969 (Prostigmata: Cheyletidae). Mit-teilungen aus dem Hamburgischen zoologischen Museum und Institut, 96: 141–155.

37. Bochkov, A.V. and Mironov, S.V. 1999b. New quill species of the family Syringophilidae (Acari: Cheyletoidea) from the European part of Russia. Acarina, 7 (1): 35–45.

38. Bochkov, A.V., Mironov, S.V. and Fain, A. 1999. Phylogeny and host–parasite relationships of the family Harpirhynchidae (Acari, Prostigmata). Acarina, 7 (2): 69–87.

39. Fain, A., Bochkov, A.V. and Mironov, S.V. 1999. A contribution to the systematics of the mite fam-ily Harpirhynchidae (Acari: Cheyletoidea). Aca-rologia, 40 (1): 35–52.

2000

40. Bochkov, A.V. 2000a. [A first record of the mites of genus Neharpyrhynchus Fain, 1972 (Acari: Harpi-rhynchidae) in Russian fauna]. Parazitologiya, 34 (6): 534–538. [In Russian with English summary]

41. Bochkov, A.V. 2000b. A new harpirhynchid mite Harpyrhynchus alaudinus sp. n. (Acari: Harpirhyn-chidae) from Alauda arvensis (Passeriformes: Alaudidae) from Russia. Acarina, 8 (2): 91–93.

42. Bochkov, A.V. 2000c. Syringophilus numidiae sp. n., a new species of quill mites (Acarina: Syringophili-dae) ex Numida meleagris (Galliformes: Phasianidae) from South Africa. Acarina, 8 (2): 103–104.

43. Bochkov, A. V., Arbobi, M. and Malikov, V. G. 2000. Notes on mites of the family Myobiidae (Acari: Prostigmata) parasitizing rodents (Mammalia: Ro-dentia) in Iran. Folia Parasitologica, 47: 73–77.

44. Bochkov, A.V. and Mironov, S.V. 2000. Two new species of the genus Geckobia (Acari: Pterygoso-matidae) from geckos (Lacertilia: Gekkonomor-pha) with a brief review of host–parasite associa-tions of the genus. Russian Journal of Herpetol-ogy, 7 (1): 61–68.

45. Bochkov, A. V., Mironov, S. V. and Kravtsova, N. T. 2000. Two new syringophilid mites (Acari: Syringophilidae) from the Greenfinch Carduelis chloris (Passeriformes: Fringillidae) from Kir-ghizia. Genus, 11 (2): 351–358.

46. Fain, A., Bochkov, A. V. and Mironov, S. V. 2000. New genera and species of quill mites of the fam-ily Syringophilidae (Acari: Prostigmata). Bulletin de l’Institut royal des Sciences naturelles de Bel-gique (Entomologie), 70: 33–70.

2001

47. Bochkov, A.V. 2001a. [On systematics of mites of the genus Syringophiloidus (Acari: Syringophilidae) from the European part of Russia]. Parazitologiya, 35 (2): 149–153. [In Russian with English summary]

48. Bochkov, A.V. 2001b. [Parallel evolution of mites of the family Myobiidae (Acari: Prostigmata) and jerboas (Rodentia: Dipodidae)]. Parazitologiya, 35 (1): 9–18. [In Russian with English summary]

49. Bochkov, A.V. and Apanaskevich, D. A. 2001. Two new species of the family Syringophilidae (Acari: Cheyletoidea) from passeriform birds collected in Altai. Folia Parasitologica, 48: 321–325.

50. Bochkov, A.V. and Chistyakov, D.V. 2001. De-scription of Charadriphilus lyudmilae gen. et sp. n. (Acari: Syringophilidae) from Scolopax rustica (Aves: Scolopacidae) in NW Russia. Zoosystem-atica Rossica, 9: 295–297.

51. Bochkov, A.V. and Fain, A. 2001. Phylogeny and system of the Cheyletidae (Acari: Prostigmata) with special reference to their host–parasite asso-ciations. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, 71: 5–36.

52. Bochkov, A.V., Fain, A. and Ardeshir, F. 2001. Redescription of Nodele calamondin Muma, 1964 (Acari Cheyletidae). Bulletin de la Société Royale Belge d’Entomologie, 137: 123–126.

53. Bochkov, A.V. and Galloway, T. D. 2001. The parasitical cheyletoid mites (Acari: Cheyletoidea) associated with passeriform birds (Aves: Passeri-formes) from Canada. Canadian Journal of Zool-ogy, 79: 2014–2028.

54. Bochkov, A.V. and Mironov, S.V. 2001. New re-cords of fur-mites of the subgenus Listrophoroides s.str. (Acari: Atopomelidae) from rats of Vietnam. Mitteilungen aus dem Hamburgischen zoolo-gischen Museum und Institut, 13: 279–285.

55. Bochkov, A.V., Mironov, S.V. and Skoracki, M. 2001. Four new quill mite species of the genus Syringophilopsis Kethley, 1970 (Acariformes: Prostigmata). Acarina, 9 (2): 233–239.

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56. Bochkov, A.V., Mironov, S.V. and Stekolnikov, A.A. 2001. [Acariform mites (Acariformes)]. In: S.A. Fetisov and G.Yu. Konechnaya (Eds.). Bioraz-noobraziye i redkie vidy natsionalnogo parka “Sebezhskii” [Biodiversity and rare species of the national park “Sebezhskii”]. Saint Petersburg State University (Publications of the St. Petersburg Society of Naturalists, Series 6, Vol. 4), Saint Pe-tersburg, pp. 109–112. [In Russian]

57. Fain, A. and Bochkov, A.V. 2001a. A new genus and species of mite (Acari: Epidermoptidae) from the ear of a South American Dove. Bulletin de la So-ciété Royale Belge d’Entomologie, 137: 117–122.

58. Fain, A. and Bochkov, A.V. 2001b. A new species of the genus Epimyodex Fain and Orts, 1969 (Acari: Cloacaridae: Epimyodicinae) parasitizing Sorex trowbridgii (Soricidae) from the U.S.A. International Journal of Acarology, 27: 221–223.

59. Fain, A. and Bochkov, A.V. 2001c. A review of some cheyletid genera (Acari : Prostigmata) with descriptions of new species. Acarina, 9 (1): 47–95.

60. Fain, A. and Bochkov, A.V. 2001d. A review of the genus Cheyletus Latreille, 1776 (Acari: Cheyleti-dae). Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, 71: 83–114.

61. Fain, A. and Bochkov, A.V. 2001e. Observations on the taxonomic status of some cheyletid genera (Acari Cheyletidae). Belgian Journal of Entomol-ogy, 3: 291–301.

2002

62. Balashov Yu.S., Bochkov, A.V., Vashchonok, V.S., Grigoreva, L.A. and Tretyakov, K.A. 2002. [Struc-ture and seasonal dynamics of the community ectoparasites in the bank vole in the Ilmen-Volkhov lowland]. Parazitologiya, 36 (6): 433–446. [In Russian with English Summary]

Bochkov, A.V. 2002. [Classification and phylog-eny of mites of the superfamily Cheyletoidea (Acari: Prostigmata)]. Entomologicheskoe Oboz-renie, 81: 488–513. [In Russian with English Sum-mary; also published in English in Entomological Review, 82 (6): 641–664.]

63. Bochkov, A.V., Fain, A. and Dabert, J. 2002. A revision of the genus Cheletopsis (Acari Chey-letidae). Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, 72: 5–26.

64. Bochkov, A.V. and Perez, T.M. 2002. New quill mites of the family Syringophilidae (Acari: Chey-letoidea) parasitizing Mexican parrots. Belgian Journal of Entomology, 4: 145–159.

65. Fain, A. and Bochkov, A.V. 2002a. A new genus and species of cheyletid mite (Acari: Cheyletidae)

from a cave in Western Australia. International Journal of Acarology, 17: 37–40.

66. Fain, A. and Bochkov, A.V. 2002b. A new fur mite Lynxacarus (Lutracarus) visoni sp. n. (Acari, Lis-trophoridae) from Mustela vison (Carnivora, Mustelidae) in North America. Acta Parasitolog-ica, 47 (1): 47–50.

67. Fain, A. and Bochkov, A.V. 2002c. A new species of the genus Eudusbabekia Jameson, 1971 (Acari: Myobiidae) parasitizing Rhinophylla pumilio (Chiroptera: Phyllostomatidae) from Ecuador. Systematic Parasitology, 52: 167–169.

68. Fain, A. and Bochkov, A.V. 2002d. On some little known and a new species of Myobiidae (Acari) associated with rodents. Bulletin de la Société Royale Belge d’Entomologie, 138: 95–105.

69. Fain, A. and Bochkov, A.V. 2002e. On some new or little known species of parasitic Cheyletidae (Acari: Prostigmata). Acarologia, 17 (2): 145–160.

70. Fain, A., Bochkov, A.V. and Corpuz-Raros, L.A. 2002. A revision of the Hemicheyletia generic group (Acari: Cheyletidae). Bulletin de l’Institut royal des Sciences naturelles de Belgique, 72: 27–66.

71. Skoracki, M. and Bochkov A.V. 2002. A new quill mite species Bubophilus asio sp. n. (Acari: Syrin-gophilidae) from the long-eared owl Asio otus (Strigiformes: Strigidae). Genus, 13 (1): 149–152.

2003

72. Balashov, Yu.S., Bochkov, A.V., Vashchonok, V.S. and Tret’yakov, K.A. 2003. [Structure and sea-sonal dynamics of the community ectoparasites on the common shrew (Sorex araneus) in the Ilmen-Volkhov Lowland]. Parazitologiya, 37 (6): 441–454. [In Russian with English Summary]

73. Bochkov, A.V. and Fain, A. 2003a. Revision of the subgenus Marquesania (Acari, Atopomelidae). Invertebrate Systematics, 17 (4): 575–604.

74. Bochkov, A.V. and Fain, A. 2003b. [A new mite species Radfordia sigmomys sp. n. (Acari: Myobi-idae) parasitizing Sigmodon alstoni (Rodentia: Sigmodontidae) from Central America]. Parazi-tologiya, 37 (2): 107–112. [In Russian with English summary]

75. Bochkov, A.V. and Fain, A. 2003c. New and little known species of the family Syringophilidae (Acari: Cheyletoidea) from parrots (Aves: Psit-taciformes). Acarina, 11: 37–44.

76. Bochkov, A.V. and Fain, A. 2003d. New observa-tions of mites of the family Myobiidae Megnin, 1778 (Acari: Prostigmata) with special reference to their host-parasite relationships. Bulletin de

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l’Institut Royal des Sciences Naturelles de Bel-gique, 73: 5–50.

77. Bochkov, A.V. and Labrzycka, A.A. 2003. Revision of the European species of the genus Myobia Van Heyden, 1826 (Acari: Myobiidae). Acta Parasito-logica, 48 (1): 24–40.

78. Bochkov, A.V. and OConnor, B.M. 2003a. New cheyletid mites (Acari, Cheyletidae) associated with birds. Acta Parasitologica, 48 (4): 265–279.

79. Bochkov, A.V. and OConnor, B.M. 2003b. Two new species of the genus Cheyletus Latreille, 1796 (Acari: Cheyletidae) associated with Philippine vertebrates. International Journal of Aca rology, 29: 327–330.

80. Fain, A. and Bochkov, A.V. 2003. A new species of the genus Metacheyletia Fain, 1972 (Acari: Cheyletidae) parasitizing Serinus mozambicus (Passeriformes: Fringillidae) in Central Africa. International Journal of Acarology, 29: 119–121.

81. Fain, A. and Bochkov, A.V. 2003a. A review of the genus Asiochirus Fain, 1970 (Acari Listrophoridae). Bulletin de la Societe Royale Belge d’Entomologie, 139: 227–232.

82. Fain, A. and Bochkov, A.V. 2003b. New notes on the family Atopomelidae (Acari: Listrophoroidea). Annales du Musée Royal de l’Afrique Centrale, series Zoologie, 291: 1–32.

83. Fain, A. and Bochkov, A.V. 2003c. New species of mites parasitic on the skin of birds (Acari: Epider-moptidae and Dermationidae). Bulletin de la So-ciete Royale Belge d’Entomologie, 139: 121–149.

84. Fain, A. and Bochkov, A.V. 2003d. Observations on the Pyroglyphidae (Acari: Astigmata) with description of a new genus and species from Cyg-nus melanocoryphus (Molina) (Aves: Anatidae). International Journal of Acarology, 29: 123–126.

2004

85. Bochkov, A.V. 2004a. A new finding of Teinochey-lus gundii Fain and al., 1982 (Acari: Cheyletidae) from a ctenodactylid rodent (Rodentia: Ctenodac-tylidae). Zoosystematica Rossica, 13 (1): 4.

86. Bochkov, A.V. 2004b. [Mites of the family Chey-letidae (Acari: Prostigmata): phylogeny, distribu-tion, evolution and analysis of host-parasite rela-tionships]. Parazitologiya, 38 (2): 122–138. [In Russian with English summary]

87. Bochkov, A.V., Fain, A. and Skoracki, M. 2004. New quill mites of the family Syringophilidae (Acari: Prostigmata). Systematic Parasitology, 57: 135–150.

88. Bochkov, A.V. and Galloway, T.D. 2004. New species and records of cheyletoid mites (Acari:

Cheyletoidea) from birds in Canada. Journal of the Kansas Entomological Society, 77: 26–44.

89. Bochkov, A.V. and Klimov, P.B. 2004. A new predaceous mite Nodele (Aztecocheyletus) con-quistador subgen. n., sp. n. (Acari: Cheyletidae) phoretic on Aztecontidium tenochtitlanicum (Hy-menoptera: Megachilidae). Acarina, 12 (1): 23–27.

90. Bochkov, A.V. and OConnor, B.M. 2004a. Phylog-eny, systematics and biology of mites of the genera Chelacheles and Neochelacheles (Acari: Chey-letidae). Invertebrate Systematics, 18: 547–592.

91. Bochkov, A.V. and OConnor, B.M. 2004b. System-atics of the genus Sciurocheyla (Acari: Cheyletidae), parasites of tropical squirrels (Rodentia: Sciuridae). Journal of Medical Entomology, 41: 308–320.

92. Bochkov, A.V., OConnor, B.M. and Gorog, A. 2004. New species and records of fur mites (Ac-ari: Astigmata: Atopomelidae) from Southeast Asian rodents (Rodentia: Muridae). Acarina, 12 (2): 67–86.

93. Fain, A. and Bochkov, A.V. 2004a. A new species of the genus Myocoptes Claparede, 1896 (Acari: Myocoptidae) parasitizing Pitymys pinetorum (Rodentia: Cricetidae) from the USA. Interna-tional Journal of Acarology, 30: 37–39.

94. Fain, A. and Bochkov, A.V. 2004b. Listrophoroides (Afrolistrophoroides) prionomys n.sp. (Acari, Atopomelidae) parasitic on Prionomys batesi (Ro-dentia, Dendromurinae) from Republique Cen-trafricaine. Journal of Afrotropical Zoology, 1: 5–8.

95. Skoracki, M., Bochkov, A.V. and Sikora, B. 2004. New records of the cheyletoid mites (Acari: Chey-letoidea) from passeriform birds in Poland. Belgian Journal of Entomology, 6: 83–90.

96. Skoracki, M., Bochkov, A.V. and Wauthy, G. 2004. Revision of the quill mites of the genus Picobia Haller, 1878 (Acari: Syringophilidae) with notes on their host-parasites relationships. Insect System-atics and Evolution, 35: 155–176.

2005

97. Bochkov, A.V., Hakimitabar, M. and Saboori, A. 2005. A review of the Iranian Cheyletidae (Acari Prostigmata). Belgian Journal of Entomology, 7: 99–109.

98. Bochkov, A.V. and Klimov, P.B. 2005. Three new species of the predaceous Cheyletidae (Acari: Prostig-mata) phoretic on insects. Acarina, 13 (1): 15–22.

99. Bochkov, A.V., Klimov A.B. and OConnor, B.M. 2005. Fur-mites of the genus Atopomelus Troues-sart, 1918 (Acari: Atopomelidae): life-cycle, phy-logeny, and host parasite-associations. Acarologia, 45: 207–241.

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100. Bochkov, A.V. and Ochoa, R. 2005. Three new cheyletid species (Acari: Cheyletidae) from the National Museum of Natural History Entomo-logical collection, USA. Acarina, 13 (2): 137–144.

101. Bochkov, A.V. and OConnor, B.M. 2005a. Ob-servations on the listrophorid (Acari: Listro-phoridae) fauna of Pakistan. Acarina, 13 (2): 165–171.

102. Bochkov, A.V. and OConnor, B.M. 2005b. Phy-logeny and host associations of the fur-mite subgenus Listrophoroides Hirst (Acari: Atopo-melidae), with an intrigue example of synhospi-tality on rats of the genus Maxomys. Invertebrate Systematics, 19: 437–498.

103. Bochkov, A.V. and OConnor, B.M. 2005c. The life-cycle of Hemigalichus chrotogale sp. nov. (Acari: Listrophoridae), with comparative obser-vations on listrophorid morphology. Journal of Natural History, 39: 3811–3832.

104. Bochkov, A.V., Williams, G. and Proctor, H. 2005. Record of Picobia zumpti (Acari: Syringophili-dae) from quills of the rock pigeon in North America. Belgian Journal of Entomology, 7: 121–127.

105. Mironov, S.V., Bochkov, A.V. and Fain, A. 2005. Phylogeny and evolution of parasitism in feather mites of the families Epidermoptidae and Der-mationidae (Acari: Analgoidea). Zoologischer Anzeiger, 243: 155–179.

2006

106. Bochkov, A.V., Mironov, S.V. and OConnor, B.M. 2006. Revision of the genus Galagocheles Fain (Acari: Cheyletidae) parasites of galagos (Pri-mates: Galagonidae). Systematic Parasitology, 63: 181–200.

107. Bochkov, A.V. and Literak, I. 2006. A review of the European Harpirhynchidae (Acari: Prostig-mata) with description of a new species. Acta Parasitologica, 51: 136–142.

108. Bochkov, A.V. and OConnor, B.M. 2006a. A re-view of the external morphology of the mite family Pterygosomatidae and its systematic posi-tion within the Prostigmata (Acari: Acariformes). Parazitologiya, 40: 201–214. [In Russian with English summary]

109. Bochkov, A.V. and OConnor, B.M. 2006b. Fur-mites of the family Atopomelidae (Acari: Astig-mata) of the Philippines: systematics, phylogeny, and host-parasite relationships. Miscellaneous Publications. Museum of Zoology, University of Michigan, 196: 1–62.

110. Bochkov, A.V. and OConnor, B.M. 2006c. Hylo-myobia (Acari: Myobiidae), a new genus of mites

parasitic on gymnures of the genus Hylomys (Eulipotyphla: Erinaceidae). Folia Parasitologi-ca, 53: 302–310.

111. Bochkov, A.V. and OConnor, B.M. 2006d. Revi-sion of the fur-mite family Listrophoridae (Aca-ri: Astigmata) associated with Philippine mam-mals. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Entomologie, 76: 5–24.

112. Bochkov, A.V. and OConnor, B.M. 2006e. Revi-sion of the genus Gaudalges (Acari: Psoroptidae), parasites of Malagasy lemurs. Acarina, 14: 3–20.

113. Klimov, P.B., Bochkov, A.V. and OConnor, B.M. 2006. Mites of the genus Cheletophyes (Acarina: Cheyletidae) associated with large African car-penter bees (Hymenoptera: Apidae: Xylocopa), with multivariate analyses of sibling species. Bio-logical Journal of Linnaean Society, 87: 45–58.

2007

114. Balashov, Yu.S., Bochkov, A.V., Vashenok, V.S., Grigoreva, L.A., Stanukovich M.K. and Tretya-kov K.A. 2007a. [Segregation of communities of ectoparasites of small forest mammals among ecological niches]. Doklady Akademii Nauk, 415 (2): 277–280. [In Russian; also publised in Eng-lish in Doklady Biological Sciences, 415 (2): 280–283]

115. Balashov, Yu.S., Bochkov, A.V., Vashenok, V.S., Grigoreva, L.A., Stanukovich M.K. and Tretya-kov K.A. 2007b. [Structure of populations and ecological niches of ectoparasites in parasite communities of small forest mammals]. Parazi-tologia, 41 (5): 329-347. [In Russian with English summary]

116. Bochkov, A.V. 2007. [Morphological adapta-tions of acariform mites (Acari: Acariformes) to permanent parasitism on mammals.] Parazi-tologiya, 41 (6): 428–458. [In Russian with English summary]

117. Bochkov, A.V., Literak, I. and Capek, M. 2007. Neharpyrhynchus baile n.sp. (Prostigmata: Harpi-rhynchidae) parasitizing Turdus leucomelas Viel-lot (Aves: Turdidae) from Brazil. International Journal of Acarology, 33: 35–39.

118. Bochkov, A.V. and Walter, D.E. 2007. The life-cycle of Pomerantzia philippina sp. n. (Prostig-mata: Pomerantziidae) described from the Philip-pines. Acarina, 15 (2): 159–170.

119. Dowling, A.P.G., Bochkov, A.V. and OConnor, B.M. 2007. Revision of the genus Andreacarus (Acari: Laelapidae) with description of seven new species and a new genus for Australian species formerly placed in Andreacarus. Journal of Medical Entomology, 44: 405–421.

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2008

120. Bochkov, A.V. 2008a. A review of the mite fam-ily Stigmocheylidae Berlese (Acari: Prostigmata). Annales Zoologici, 58: 311–325.

121. Bochkov, A.V. 2008b. New observations on phy-logeny of cheyletoid mites (Acari: Prostigmata: Cheyletoidea). Proceedings of the Zoological Institute RAS, 312 (1–2): 54–73.

122. Bochkov, A.V. 2008c. Origin and evolution of parasitism in mites of the infraorder Eleutheren-gona (Acari: Prostigmata). Report I. Lower Raphignathae. Parazitologiya, 42 (5): 337–359. [In Russian with English Summary]

123. Bochkov, A.V., Klompen, H. and OConnor, B.M. 2008. A review of the external morphology of the family Chirorhynchobiidae (Acari: Sarcoptoidea) with description of a new species. Journal of Medical Entomology. 45: 193–202.

124. Bochkov, A.V. and Literak, I. 2008. Description of the life-stages of Harpyrhynchoides rubeculi-nus (Černý and Sixl, 1971) (Prostigmata: Harpi-rhynchidae). International Journal of Acarology, 34: 227–236.

125. Bochkov, A.V. and Mironov, S.V. 2008. [The phenomenon of “phylogenetic synhospitality” in acariform mites (Acari: Acariformes)—the permanent parasites of vertebrates]. Parazi-tologiya, 42 (2): 81–100. [In Russian with English Summary]

126. Bochkov, A.V. and OConnor, B.M. 2008a. A new mite superfamily Cloacaroidea and its position within the Prostigmata (Acariformes). Journal of Parasitology, 94: 335–344.

127. Bochkov, A.V. and OConnor, B.M. 2008b. The external morphology of the mite subfamily Dro-micioptinae (Acari: Myocoptidae), evidences for and against the myocoptid monophyly. Annals of the Entomological Society of America, 101: 289–296.

128. Bochkov, A.V., OConnor, B.M. and Wauthy, G. 2008. Phylogenetic position of the family Myo-biidae within the Prostigmata (Acari: Acari-formes). Zoologischer Anzeiger, 247: 15–45.

129. *Bochkov, A.V., Zabludovskaya, S.A. and OCon-nor, B.M. 2008. Phylogeny and systematics of the endoparasitic astigmatid mites (Acari: Sar-coptiformes) of mammals: families Gastronys-sidae, Lemurnyssidae, and Pneumocoptidae. Zootaxa, 1951: 1–152.

2009

130. *Bochkov, A.V. 2009a. A review of mites of the parvorder Eleutherengona (Acariformes: Pro-

stigmata)—permanent parasites of mammals. Acarina, supplement, 149 pp.

131. Bochkov, A.V. 2009b. Mites of the family Myobi-idae (Acari: Prostigmata) parasitizing rodents of the former USSR. Acarina, 17 (2): 109–169.

132. Bochkov, A.V. 2009c. [Origin and evolution of parasitism in mites of the infraorder Eleutheren-gona (Acari: Prostigmata). Report II. Superfam-ily Cheyletoidea]. Parazitologiya, 43 (2): 97–117. [In Russian with English Summary]

133. Bochkov, A.V., Flannery, M.E. and Spicer, G. 2009. Mites of the genus Torotrogla (Prostig-mata: Syringophilidae) from North American passerines. Journal of Medical Entomology, 46: 183–197.

134. Bochkov, A.V., Melnikov, D.A. and Nazarov, R.A. 2009. Pterygosoma (Pterygosoma) pseudo-trapelus sp. nov. (Acariformes: Pterygosomati-dae)—ectoparasite of Pseudotrapelus sinaitus (Squamata: Agamidae) from Jordan. Zootaxa, 2232: 61–68.

135. Bochkov, A.V. and OConnor, B.M. 2009. A new mite genus Caenolestomyobia gen. nov. (Acari-formes: Myobiidae) from marsupials of the genus Caenolestes (Pauctituberculata: Caenolestidae). Proceedings of the Zoological Institute RAS, 313 (4): 140–151.

136. Bochkov, A.V. and Wauthy, G. 2009. A review of the genus Listropsoralges (Acari, Psoroptidae) with the descriptions of two new species. Acta Parasitologica, 54: 269–275.

137. Literak, I. Bochkov, A.V., Dusbabek, F., Martinu, J., Podzemny, P. and Sychra, O. 2009. Málo známí cizopasníci ptáků—roztoči čeledi Harpirhynchi-dae. Ziva, 3: 126–128. [In Czech]

138. Mironov, S.V. and Bochkov, A.V. 2009. [Modern conceptions of macrophylogeny in acariform mites (Chelicerata: Acariformes)]. Zoologiches-kiy Zhurnal, 88: 922–937. [In Russian with English summary; also translated into English as: Modern conceptions concerning macrophylogeny of acariform mites (Chelicerata: Acariformes), Entomological Review, 89 (8): 975–992.]

139. Whitaker, J.O. Jr., Ruckdeschel, C. and Bochkov, A.V. 2009. Species of the genus Schizocarpus Trouessart, 1896 (Acari: Chirodiscidae) from Florida and Georgia beavers. Florida Scientist, Biological Sciences, 72: 18–21.

2010

140. Bochkov, A.V. 2010. A review of mammal as-sociated Psoroptidia (Acariformes: Astigmata). Acarina, 18 (2): 99–260.

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141. Bochkov, A.V. and Mertins, J.W. 2010. Harpi-rhynchus quasimodo n.sp. (Acariformes: Harpi-rhynchidae), a new species parasitizing Molothrus ater (Passeriformes: Icteridae) in Florida. Inter-national Journal of Acarology, 36 (1): 83–87.

142. Bochkov, A.V. and OConnor, B.M. 2010. Exter-nal morphology and systematics of the family Lobalgidae (Acari: Astigmata) with resolution of the question of family monophyly. Insect System-atics and Evolution, 41: 187–229.

143. Bochkov, A.V. and OConnor, B.M. 2010. Nidi-colocheyletus kennedyae n.gen., n.sp. (Prostig-mata: Cheyletidae) from nests of passerine birds in Michigan, USA. International Journal of Aca-rology, 36 (1): 59–71.

144. Bochkov, A.V., OConnor, B.M. and Wauthy, G. 2010. A review of the mite subfamily Makialginae (Acari: Psoroptidae)—permanent parasites of strepsirrhine primates. Tijdschrift voor Ento-mologie, 153 (2): 217–237.

145. Bochkov, A.V. and Otto, J.S. 2010. Five new species of predaceous cheyletid mites (Acari: Cheyletidae). Zootaxa, 2727: 1–20.

146. Skoracki, M. and Bochkov, A.V. 2010. Syringo-philid mites (Acari: Syringophilidae) of Kazakh-stan. Zootaxa, 2546: 52–68.

2011

147. *Bochkov A.V. 2011a. Mites of the subgenus Microtimyobia (Acariformes: Myobiidae: Rad-fordia) and their host-parasite relationships with cricetid rodents (Cricetidae). Zootaxa, 2954: 1–86.

148. Bochkov, A.V. 2011b. [Evolution of parasitism in mammal-associated mites of the group Psoroptidia (Acari: Astigmata).] Parazitologiya, 45 (4): 257–272. [In Russian with English summary, also trans-lated in English: Evolution of parasitism in mam-mal-associated mites of the Psoroptidia group (Acari: Astigmata), Entomological Review, 91 (9): 1206–1215.]

149. Bochkov, A. V. 2011c. [Acariform mites perma-nent parasites of mammals: phylogeny, system-atic and host-parasite interactions]. Doctor of Sciences thesis. Zoological Institute RAS, Saint-Petersburg. 783 pp. [Manuscript, in Russian]

150. Bochkov, A.V., Abramov, A.V., Durden, L.A., Apanaskevich, D.A., Stekolnikov, A.A., Stanyu-kovich, M.K., Gnophanxay, S. and Tikhonov, A.N. 2011. Arthropod symbiotes of Laonastes aenigmamus (Rodentia: Diatomyidae). Journal of Parasitology, 97 (2): 352–353.

151. Bochkov, A.V., Abramov, A.V. and Hugot, G.P. 2011. Five new species of fur mites (Acariformes: Sarcoptoidea) from small mammals in Laos. Zootaxa, 2980: 1–22.

152. Bochkov, A.V. and Dubinina, H.V. 2011. Mites of the genus Schizocarpus (Acariformes: Chiro-discidae) parasitizing the Eurasian beaver Castor fiber (Rodentia: Castoridae) in the Voronezh National Reserve. Acarina, 19 (1): 53–66.

153. Bochkov, A.V., Klimov, P.B. and Wauthy, G. 2011. Phylogeny and coevolutionary associations of makialgine mites (Acari, Psoroptidae, Maki-alginae) provide insight into evolutionary history of their hosts, strepsirrhine primates. Zoology Journal of the Linnean Society, 162: 1–14.

154. Bochkov, A.V. and Literak, I. 2011. Mites of the genus Neharpyrhynchus Fain (Acariformes, Harpirhynchidae) from Neotropical birds. Zoo-Keys, 89: 15–31.

155. Bochkov, A. V. and Medvedev, S.G. 2011. [Ev-erything appeared to be not so dreadful]. Priroda, 8: 75–80. [In Russian]

156. Bochkov A.V. and Mironov S.V. 2011. Phylogeny and systematics of mammal-associated psoroptid-ian mites (Acariformes: Astigmata: Psoroptidia) derived from external morphology. Invertebrate Systematics, 25: 22-59.

157. Bochkov A.V. and Skoracki M. 2011. A new cheyletid mite Metacheyletia ngaii n.sp. (Acari-formes: Cheyletidae) from quills of Corythaixo-des leucogaster (Musophagidae) from Tanzania. Acarologia, 51 (1): 93–97.

158. Bochkov, A.V. and Skirnisson, K. 2011. Descrip-tion of the life stages of quill mite Mironovia lagopus sp. nov. (Acari: Syringophilidae) parasit-izing the rock ptarmigan Lagopus muta (Pha-sianidae) from Iceland. Parasitology Research, 108: 715–722.

159. Bochkov, A.V., Skoracki, M., Hendricks, S.A. and Spicer, G.S. 2011. Further investigations of the mite genus Syringophiloidus (Acariformes: Syringophilidae) from North American Passer-ines. Systematic Parasitology, 79: 201–211.

160. Skoracki, M., Bochkov, A.V. and OConnor, B.M. 2011. Notes on quill mites (Acariformes, Syrin-gophilidae) from aquatic birds in North America. Acta Parasitologica, 56 (3): 325–330.

161. Zhang, Z.-Q., Fan, Q.-H., Pesic, V., Smit, H., Bochkov, A.V., Khaustov, A.A., Baker, A., Wohlt-mann, A., Wen, T., Amrine, J.W., Beron, P., Lin, J., Gabrys, G. and Husband, R. 2011. Order Trombidiformes Reuter, 1909. In: Z.-Q. Zhang (Ed.). Animal biodiversity: an outline of higher-

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level classification and survey of taxonomic rich-ness. Zootaxa, 3148: 129–138.

2012

162. Bochkov, A.V. 2012. Macropodicoptes mironovi n.g., n.sp. (Acariformes: Sarcoptidae), a new mange mite from Wallabia bicolor (Desmarest) (Diprotodonia: Macropodidae). Systematic Para-sitology, 83 (1): 29–38.

163. Bochkov, A.V. 2012. Schizocarpus saveljevi sp. nov. (Acariformes: Chirodiscidae) parasitizing the Eurasian beaver—Castor fiber Linnaeus, 1758 (Rodentia: Castoridae) from Leningrad Province (Russia). Proceedings of the Zoological Institute RAS, 316 (2): 166–171.

164. Bochkov, A.V. and Mironov, S.V. 2012. Mites of the genus Passeroptes Fain (Acariformes: Dermationidae) from passerines (Aves: Pas-seriformes) of North-Western Russia. Zootaxa, 3563: 43–57.

165. Bochkov, A.V. and Skoracki, M. 2012. New and poorly known predaceous cheyletids (Acari-formes: Cheyletidae) from feather quills. Inter-national Journal of Acarology, 38 (6): 486–496.

166. Bochkov, A.V., Labrzycka, A., Skoracki M. and Saveljev, A.P. 2012. Fur mites of the genus Schizocarpus Trouessart (Acari: Chirodiscidae) parasitizing the Eurasian beaver Castor fiber belorussicus Lavrov (Rodentia: Castoridae) in NE Poland (Suwałki). Zootaxa, 3152: 39–59.

167. Literak, I., Bochkov A.V., Cárdenas-Callirgos, J. and Capek, M. 2012. The first records of mites of the genus Neharpyrhynchus (Acariformes: Harpirhynchidae) from birds in Peru. Neotropical Helminthology, 6 (1): 109–114.

168. Sikora, B. and Bochkov, A.V. 2012. Fur mites of the family Listrophoridae (Acariformes: Sarcop-toidea) associated with South American sigmo-dontine rodents (Cricetidae: Sigmodontinae). Acta Parasitologica, 57 (4): 388–396.

169. Skoracki, M., Zabludovskaya, S. and Bochkov, A.V. 2012. A review of Prostigmata (Acariformes: Trombidiformes) permanently associated with birds. Acarina, 20 (2): 67–107.

2013

170. Bochkov, A.V. and Galloway, T.D. 2013. New records and new species of mites of the subfam-ily Harpirhynchinae (Acariformes: Harpirhynchi-dae) infesting birds in Manitoba, Canada. Acta Parasitologica, 58 (4): 405–419.

171. Bochkov, A.V. and Mironov S.V. 2013. [Is para-sitism of Metazoa “a one-way ticket?”] Parazi-

tologiya, 47(3): 212–227. [In Russian with English summary; also published in English in Entomological Review, 93 (9): 1196–1206]

172. Bochkov, A.V. and OConnor, B.M. 2013. Mites of the Subfamily Harpirhynchinae (Acariformes: Harpirhynchidae) from North American Birds. Journal of Medical Entomology, 50 (3): 501–509.

173. *Bochkov, A.V., OConnor, B.M. and Grootaert, P. 2013. Revision of the family Listropsoralgidae Fain, 1965 (Acariformes: Sarcoptoidea)—skin parasites of marsupials and rodents. Zootaxa, 3611: 1–69.

174. Skoracki, M., Glowska, E. and Bochkov, A.V. 2013. Phylogeny of quill mites of the family Syringophilidae (Acari: Prostigmata) based on their external morphology. European Journal of Entomology, 110 (4): 663–675.

2014

175. Bochkov A.V., Barreto M. 2014. New species of the subgenus Radfordia (Hesperomyobia) (Acari-formes, Myobiidae)—parasites of the subfamily Sigmodontinae (Rodentia: Cricetidae: Sigmodon-tinae). International Journal of Acarology, 40 (1): 37–42.

176. Bochkov, A., Lareschi, M. and Barreto, M. 2014. New species and records of the mite genus Pro-listrophorus (Acariformes: Listrophoridae) from rodents of the subfamily Sigmodontinae (Roden-tia: Cricetidae). Acta Parasitologica, 59 (4): 586–595.

177. Bochkov, A.V. 2014. Two new species of the subfamily Harpirhynchinae (Acari: Harpirhyn-chidae) from African birds. Annales Zoologici, 64: 479–483.

178. Bochkov, A.V. and Abramov, A.V. 2014. Acari-form mites (Acariformes)—permanent symbionts of Hapalomys delacouri Thomas (Rodentia, Muridae) in Vietnam. ZooKeys, 459: 137–145.

179. Bochkov, A.V. and Grootaert, P. 2014. Mites of the genus Paracoroptes Lavoipierre, 1955 (Acari-formes: Psoroptidae)—skin parasites of the Af-rican monkeys of the family Cercopithecidae (Primates). Zootaxa, 3887 (2): 225–238.

180. Bochkov, A.V. and Guzman-Cornejo, C. 2014. Mites of the subgenus Neotomobia n.subg. (Acariformes: Myobiidae: Radfordia), parasites of the subfamily Neotominae (Rodentia: Criceti-dae). Systematic Parasitology, 89 (2): 91–100.

181. Bochkov, A.V. and Klompen, H. 2014. A review of the subfamily Harpypalpinae Fain, 1972 (Acariformes: Harpirhynchidae)—parasites of passerine birds. Zootaxa, 3857 (4): 451–477.

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182. Bochkov, A.V. and Klompen, H. 2014. A review of the zumpti species group of the genus Harpy-rhynchoides (Acariformes: Harpirhynchidae)—ectoparasites of passerines. Zootaxa, 3884 (5): 401–418.

183. Bochkov, A.V. and Klompen, H. 2014. New Harpirhynchinae Dubinin (Acariformes: Harpi-rhynchidae)—intracutaneous and feather-base parasites of birds. Zootaxa, 3860 (4): 301–324.

184. Bochkov, A.V. and OConnor, B.M. 2014. Mites of the genus Trichorhynchiella (Acariformes: Harpirhynchidae): A new example of ovovivipar-ity in acariform mites. Journal of Medical Ento-mology, 51 (2): 323–330.

185. Bochkov, A.V. and Saveljev, A.P. 2014. Mites of the genus Schizocarpus Trouessart, 1896 (Acari-formes: Chirodiscidae) from the North American beavers (Castor canadensis) in Russia. Parazi-tologiya, 48 (6): 430–436. [In Russian with English summary]

186. Bochkov, A.V., Klimov, P.B. and Grootaert, P. 2014. Revision of the subfamily Onychalginae Fain, 1988 (Acariformes: Pyroglyphidae)—ectoparasites of passerine birds. Zootaxa, 3785(2): 175–200.

187. Bochkov, A.V., Klimov, P.B., Hestvik, G. and Saveljev, A.P. 2014. Integrated Bayesian species delimitation and morphological diagnostics of chorioptic mange mites (Acariformes: Psoropti-dae: Chorioptes). Parasitology Research, 113 (7): 2603–2627.

188. Literák, I, Bochkov, A. and Franco, S. 2014. The first record of mites of the genus Neharpyrhyn-chus (Acariformes: Harpirhynchidae) from birds in Paraguay. Neotropical Helminthology, 8 (1): 183–188.

189. Mertins, J.W. and Bochkov, A.V. 2014. Key to species of the genus Neocheyletiella (Acari-formes: Cheyletidae), with description of a new species. Journal of Medical Entomology, 51 (6): 1116–1121.

2015

190. Bochkov, A.V. 2015. Comparison of external morphology of tritonymphs in myobiid mites (Myobiidae) associated with marsupials (Marsu-pialia). International Journal of Acarology, 41: 590–599. DOI: 10.1080/01647954.2015.1084046

191. Bochkov, A.V. 2015. Mites of the genus Harpy-rhynchiella Fain, 1972 (Acariformes: Harpirhyn-chidae), an example of an unusual mating strat-egy. International Journal of Acarology, 41: 31–40. DOI: 10.1080/01647954.2014.988644

192. Bochkov, A.V. and Klompen, H. 2015. Two new mite species of the subfamily Harpirhynchinae

Dubinin, 1957 (Acariformes: Harpirhynchidae), parasites of the passerine birds (Aves: Passeriformes) in Australia and South Asia. Systematic Parasitol-ogy, 92: 23–29. DOI: 10.1007/s11230-015-9

193. Bochkov, A.V. and Mirolubov, A.A. 2015. Descrip-tion of postembrionic immature stages of Protomyo-bia onoi Jameson & Dusbabek, 1971 (Acariformes: Myobiidae) and comparative analysis of the juvenile external morphology in the tribe Protomyobiini. International Journal of Acarology, 41: 115–127. DOI: 10.1080/01647954.2014.1003966

194. Bochkov, A.V. and Mironov, S.V. 2015. A new species of the genus Neodermation Fain (Acari-formes: Dermationidae) and the first record of the precopulatory mate-guarding among dermatio-nids. Systematic and Applied Acarology, 20: 612–628. DOI: 10.11158/saa.20.6.4

195. Bochkov, A.V., Klompen, H., Junge, R.E. and Wiliams, C.V. 2015. Lemuralges propithecus sp. n. (Acariformes: Psoroptidae), an ectoparasite of the diademed sifaka Propithecus diadema (Primates: Indriidae). Folia Parasitologica, 62: 11. DOI: 10.14411/fp.2015.011

196. *Bochkov, A.V., OConnor, B.M. and Klompen, H. 2015. A review of the mite subfamily Harpi-rhynchinae (Acariformes: Harpirhynchidae)—parasites of New World birds (Aves: Neognathae). Zootaxa, 4023: 1–130. DOI: 10.11646/zoo-taxa.4023.1.1.

197. Hernandes, F.A., Pedroso, L.G.A. and Bochkov, A.V. 2015. Carcinopodacarus polymorphus gen. n. et sp. n. from Guira guira (Cuculiformes: Cuculidae) in Brazil: a first example of male polymorphism in the family Dermationidae (Acariformes: Analgoidea). Folia Parasitologica, 62: 9. DOI: 10.14411/fp.2015.009.

198. Klimov, P.B., Bochkov, A.V. and OConnor, B.M. 2015a. Phylogenetic position of the house dust mite subfamily Guatemalichinae (Acariformes: Pyroglyphidae) based on integrated molecular and morphological analyses and different mea-sures of support. Cladistics, 32 (3): 261–275. DOI: 10.1111/cla.12126

199. Klimov, P.B., Bochkov, A.V. and OConnor, B.M. 2015b. Tachornithoglyphus gen. nov.—a new genus of nidicolous Pyroglyphidae (Acariformes: Astigmata). Zootaxa, 3956: 97–112. DOI: 10.11646/zootaxa.3956.1.5

2016

200. Bochkov, A.V. 2016. A new mite species Schizo-coptes daberti sp. n. (Acariformes: Chirodiscidae) from Chrysochloris stuhlmanni Matsche (Af-rosoricida: Chrysochloridae) in the Democratic

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Republic of the Congo. Folia Parasitologica, 63: 005. DOI: 10.14411/fp.2016.005

201. Bochkov, A.V. 2016. Myocoptid mites (Acari-formes: Myocoptidae) of the fauna of the former USSR. Zootaxa, 4193 (3): 451–485. DOI: 10.11646/zootaxa.4193.3.2

202. Bochkov, A.V. and Abramov, V.V. 2016. To fauna of the free-living Cheyletidae (Acari-formes: Cheyletoidea) of the European part of Russia. Systematic and Applied Acarology, 21 (3): 335–346. DOI: 10.11158/saa.21.3.8

203. Bochkov, A.V. and Sidorchuk, E.A. 2016. A new Eocene free-living cheyletid mite from Baltic amber. Acta Palaeontologica Polonica, 61 (4): 869–874. DOI: 10.4202/app.00244.2016

204. Bochkov, A.V. and Valim, M.P. 2016. New spe-cies and records of mites of the superfamily Sarcoptoidea (Acariformes: Psoroptidia) from mammals in Brazil. Acta Parasitologica, 61 (1): 22–41. DOI: 10.1515/ap-2016-0003

205. Bochkov, A.V., Mbalitini, S.G. and Verheyen, E. 2016. Crocidurobia faini sp. n. (Acariformes: Myobiidae)—a new mite species parasitizing shrews of the genus Crocidura (Soricimorpha: Soricidae) in DR Congo. Systematic Parasitology, 93: 493–499. DOI: 10.1007/s11230-016-9623-2

206. Bochkov, A.V., Mbalitini, S.G. and Verheyen, E. 2016. Review of the fur-mite genus Soricilichus Fain, 1970 (Acariformes: Chirodiscidae)—sym-bionts of the African shrews of the subfamily Crocidurinae (Soricomorpha: Soricidae). Zoo-taxa, 4072 (2): 235–253. DOI: 10.11646/zoo-taxa.4072.2.6

207. Bochkov, A.V., Valim, M.P., Ochoa, R., OConnor, B.M. and Averianov, A.O. 2016. Origin of host-parasite associations of Marsupialges misonnei (Acariformes: Psoroptidae)—a parasitological detective story. Parasitology Research, 115 (10): 3695–3698. DOI: 10.1007/s00436-016-5192-1

208. Bochkov, A.V. 2016. [Chapter IV. Host-parasite relationships of acariform mites (Acariformes), permanent parasites of mammals]. In: K.V. Galak-tionov (Ed.). [Coevolution of parasites and hosts]. Proceedings of the Zoological Institute RAS, Sup-plement 4: 160–263. https://www.zin.ru/journals/trudyzin/eng/supplements.html?year=2016 [In Russian with English summary].

2017

209. Bochkov, A.V., Klimov, P.B. and Skoracki, M. 2017. Morphological phylogenetic conflict in the parasitic mite family Harpirhynchidae (Acari-formes: Cheyletoidea) correlates with host as-sociations. Zoologischer Anzeiger, 271: 33–48.

210. Bochkov A.V., OConnor B.M., Skoracki, M. 2017. New species and new records of mites of the fam-ily Myocoptidae (Acariformes: Sarcoptoidea) from African rodents. Journal of Medical Entomology, 54 (3): 587–596.

211. Bochkov, A.V. 2017. A review of fur-mites of the genus Dubininetta Fain & Lukoschus, 1978 (Acari-formes: Listrophoridae), ectoparasites of moles (Soricomorpha: Talpidae). Systematic Parasitology, 94: 635–647. DOI: 10.1007/s11230-017-9732-6

212. Bochkov, A.V. 2017. Crocidurobia lukoschusi sp.nov. (Acariformes: Myobiidae) parasitizing Crocidura tanakae Kuroda, 1938 (Soricimorpha: Soricidae) in Vietnam, the first record of the genus from oriental white-toothed shrews. Interna-tional Journal of Acarology, 43: 212–216. DOI: 10.1080/01647954.2016.1268204

213. Bochkov, A.V. 2017. To the systematics of the subgenus Hesperomyobia Bochkov, 1997 (Acari-formes: Myobiidae: Radfordia). Systematic and Applied Acarology, 22 (6): 785–788. DOI: 10.11158/saa.22.6.4.

214. Bochkov, A.V. and OConnor, B.M. 2017. A new species of myocoptid mites Myocoptes (Comyo-coptes) nesokia sp.nov. (Acariformes: Myocop-tidae) parasitizing Nesokia indica (Rodentia: Muridae) in Pakistan. Acta Parasitologica, 62: 46–49. DOI: 10.1515/ap-2017-0005

2018

215. Bochkov A.V. 2018. External morphology of postembrionic stages of Lutrilichus javanicus (Acariformes: Chirodiscidae) from Melogale moschata (Carnivora: Mustelidae) from Vietnam. Acarologia, 58 (4): 763–772. DOI: 10.24349/acarologia/20184271

216. Bochkov A.V. 2018. Gaudalges averianovi sp.nov. (Acariformes: Psoroptidae)—a new parasite of Malagasy lemurs. Systematic and Ap-plied Acarology, 23 (4): 593–596. DOI http://doi.org/10.11158/saa.23.4.1

217. Bochkov, A. V., Klimov, P. B., Da-Hee, K. and Skoracki, M. 2018. Validation of the status of a species with high CO1 and low nuclear genetic divergences: the scab mite Caparinia ictonyctis stat. res. (Acariformes: Psoroptidae) parasitizing the African hedgehog Atelerix albiventris. Zootaxa, 4544 (4): 523–547. DOI: 10.11646/zoo-taxa.4544.4.4

2019

218. Klimov, P. B., Skoracki, M. and Bochkov, A. V. 2019. Cox1 barcoding versus multilocus species delimitation: validation of two mite species with

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contrasting effective population sizes. Parasites and Vectors, 12 (1): 8. DOI: 10.1186/s13071-018-3242-5

219. Sidorchuk, E. A., Bochkov, A. V., Weiterschan, T. and Chernova, O. F. 2019. A case of mite-on-mammal ectoparasitism from Eocene Baltic amber (Acari: Prostigmata: Myobiidae and Mam-malia: Erinaceomorpha). Journal of Systematic Paleontology, 17 (4): 331–347. DOI: 10.1080/ 14772019.2017.1414889

*—monographs.

LIST OF MITE TAXA ESTABLISHED BY A.V. BOCHKOV

Andrei Bochkov personally and with co-au-thors established 7 family-group names (3 sub-families and 4 tribes), 45 genus-group names (34 genera and 11 subgenera) and 445 species-group names (1 fossil).

Family-group names

SubfamiliesApocnemidocoptinae Mironov, Bochkov et

Fain, 2005 (Dermationidae) Otocoptoidinae Fain et Bochkov, 2001 (Der-

mationidae)Protomyobiinae Bochkov, 1997 (Myobiidae)

TribesAcanthophthiriini Bochkov, 1997 (Myobiidae)Elephantulobiini Bochkov, 1997 (Myobiidae)Phyllostomonyssini Bochkov, Zabludovskaya

et OConnor, 2008 (Gastronyssidae)Protomyobiini Bochkov, 1997 (Myobiidae)

Genus-group names

GeneraApocalypsis Bochkov, 2010 (Myocoptidae),

nom. preocc., Rothschild 1903 (Lepidoptera, Sphingidae)

Apocalyptoides Bochkov, 2016 (Myocoptidae), nom. nov. pro Apocalypsis Bochkov, 2010

Archemyialges Mironov, Bochkov et Fain, 2005 (Epidermoptidae)

Caenolestolichus Fain et Bochkov, 2003 (Atopo-melidae)

Caenolestomyobia Bochkov et OConnor, 2009 (Myobiidae)

Carcinopodacarus Hernandes, Pedroso et Boch-kov, 2015 (Dermationidae)

Castosyringophilus Bochkov et Perez, 2002 (Syringophilidae)

Charadriaulobia Bochkov, Fain et Skoracki, 2004 (Syringophilidae)

Charadriphilus Bochkov et Chistyakov, 2001 (Syringophilidae)

Corvitorotroglus Skoracki et Bochkov, 2010 (Syringophilidae)

Crassacarus Bochkov, OConnor et Klompen, 2015 (Harpirhynchidae)

Cygnocoptes Fain et Bochkov, 2003 (Pyro-glyphidae; presently in Psoroptoididae)

Didelphialges Bochkov, OConnor et Grootaert, 2013 (Listropsoralgidae)

Fainharpirhynchus Bochkov et Galloway, 2013 (Harpirhynchidae)

Fritschisyringophilus lonchurae Bochkov, Fain et Skoracki, 2004 (Syringophilidae)

Granulocheyletus Fain et Bochkov, 2002 (Cheyletidae)

Hemimyalges Mironov, Bochkov et Fain, 2005 (Epidermoptidae)

Hylomysobia Bochkov et OConnor, 2006 (Myobiidae)

Juxtalaelaps Dowling, Bochkov et OConnor, 2007 (Laelapidae)

Kenyacheylus Fain et Bochkov, 2001 (Chey-letidae)

Macropodicoptes Bochkov, 2012 (Sarcoptidae)Megasyringophilus Fain, Bochkov et Mironov,

2000 (Syringophilidae)Neoaulobia Fain, Bochkov et Mironov, 2000

(Syringophilidae)Nidocheyletus Bochkov et OConnor, 2010

(Cheyletidae)Oconnoricheylus Bochkov et Otto, 2010

(Cheyletidae)Otocoptoides Fain et Bochkov, 2001 (Epider-

moptidae)Phalarophilus Skoracki, Bochkov et OConnor,

2011 (Syringophilidae)Picocheyletus Bochkov et OConnor, 2003

(Cheyletidae) Protohylomysobia Sidorchuk et Bochkov, 2018

(2019) (Myobiidae)Pseudoopsonyssus Bochkov, Zabludovskaya

et OConnor, 2008 (Gastronyssidae)Psittaciphilus Fain, Bochkov et Mironov, 2000

(Syringophilidae)

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Tachornithoglyphus Klimov, Bochkov et OConnor, 2015 (Pyroglyphidae)

Terratosyringophilus Bochkov et Perez, 2002 (Syringophilidae)

Trochiloptes Mironov, Bochkov et Fain, 2005 (Dermationidae)

SubgeneraAcomyobia Bochkov et Fain, 2003 (Myobiidae:

Radfordia)Aztecocheyletus Bochkov et Klimov, 2004

(Cheyletidae: Nodele)Dendromyobia Bochkov, 1997 (Myobiidae:

Radfordia)Dipodomyobia Bochkov, 1997 (Myobiidae:

Radfordia)Emballomyobia Bochkov et Fain, 2003 (Myo-

biidae: Ugandobia)Hesperomyobia Bochkov, 1996 (Myobiidae:

Radfordia)Macroscelistrophoroides Fain et Bochkov,

2003 (Atopomelidae: Listrophoroides)Neotomobia Bochkov et Guzman-Cornejo,

2014 (Myobiidae: Radfordia)Otomyobia Bochkov et Fain, 2003 (Myobiidae:

Myobia)Petromyscobia Bochkov et Fain, 2003 (Myo-

biidae: Radfordia)Pseudoharpirhynchus Fain, Bochkov et

Mironov, 1999 (Harpirhynchidae: Harpirhynchus), synonymized with Harpyrhynchoides Fain, 1972 in: Bochkov et al. 2015

Species-group names

ACARIFORMESASTIGMATASuperfamily AnalgoideaFamily Dermationidae

Carcinopodacarus polymorphus Hernandes, Pedroso et Bochkov, 2015

Neodermation actitis Bochkov et Mironov, 2015

Passeroptes ampeliceps Fain et Bochkov, 2003Passeroptes hirundiphilus Fain et Bochkov, 2003Passeroptes hippolais Bochkov et Mironov, 2012Passeroptes lamprotornis Fain et Bochkov, 2003Passeroptes oenanthe Fain et Bochkov, 2003Passeroptes sylviae Bochkov et Mironov, 2012Passeroptes turdoides Fain et Bochkov, 2003Paradermation colii Fain et Bochkov, 2003

Psittophagoides brotogeris Fain et Bochkov, 2003

Family Epidermoptidae

Otocoptoides mironovi Fain et Bochkov, 2001

Myialges (Metamicrolichus) pelecani Fain et Bochkov, 2003

Rallepidermoptes rallicola Fain et Bochkov, 2003

Family Psoroptoididae

Cygnocoptes prasadi Fain et Bochkov, 2003 [in content of Pyroglyphidae]

Superfamily PsoroptoideaFamily Lobalgidae

Echimytricalges lonchothrix Bochkov et OConnor, 2010

Echimytricalges proechimys Bochkov et OConnor, 2010

Family Psoroptidae

Chorioptes sweatmani Bochkov, Klimov, Hes-tvik et Saveljev, 2014

Gaudalges averianovi Bochkov, 2018Gaudalges brevisetosus Bochkov et OConnor,

2006

Lemuralges propithecus Bochkov, Klompen, Junge et Wiliams, 2015

Paracoroptes miopithecus Bochkov et Groot-aert, 2014

Paracoroptes piliocolobus Bochkov et Groot-aert, 2014

Superfamily SarcoptoideaFamily Atopomelidae

Atopomelus hylomys Bochkov, Klimov et OCon-nor, 2005

Atopomelus megalotis Bochkov, Abramov et Hugot, 2011

Atopomelus priapus Bochkov, Klimov et OCon-nor, 2005

Austrochirus (Austrochiroides) dubininae Fain et Bochkov, 2003

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134

Austrochirus (Austrochirus) peroryctes Fain et Bochkov, 2003

Bathyergolichus cryptomys Fain et Bochkov, 2003

Caenolestolichus lukoschusi Fain et Bochkov, 2003

Didelphoecius micoureus Bochkov et Valim, 2016

Listrophoroides (Afrolistrophoroides) colomys Fain et Bochkov, 2003

Listrophoroides (Afrolistrophoroides) myomys-cus Fain et Bochkov, 2003

Listrophoroides (Afrolistrophoroides) paraleg-gada Fain et Bochkov, 2003

Listrophoroides (Afrolistrophoroides) priono-mys Fain et Bochkov, 2005

Listrophoroides (Alistrophoroides) aethomys Fain et Bochkov, 2003

Listrophoroides (Lemurlistrophoroides) lepil-emur Fain et Bochkov, 2003

Listrophoroides (Listrophoroides) batomys Bochkov et OConnor, 2008

Listrophoroides (Listrophoroides) bellulus Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) cambodien-sis Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) chrootomys Bochkov et OConnor, 2008

Listrophoroides (Listrophoroides)crunomys Bochkov et OConnor, 2008

Listrophoroides (Listrophoroides) femoratus Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) gorogae Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) helenae Bochkov et Mironov, 2001

Listrophoroides (Listrophoroides) inas Boch-kov et OConnor, 2005

Listrophoroides (Listrophoroides) inflatus Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) insu1aris Bochkov et OConnor, 2008

Listrophoroides (Listrophoroides) khmeri Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) maxomys Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) megaphallus Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) moi Boch-kov et OConnor, 2005

Listrophoroides (Listrophoroides) musschen-broekii Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) musseri Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) oconnori Bochkov, Abramov et Hugot, 2011

Listrophoroides (Listrophoroides) parafemo-ratus Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) pseudoan-cophallus Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) pseudorajah Bochkov et OConnor, 2005

Listrophoroides (Listrophoroides) rapit Boch-kov, OConnor et Gorog, 2004

Listrophoroides (Listrophoroides) surifer Bochkov et OConnor, 2005

Listrophoroides (Macroscelistrophoroides) petrodromi Fain et Bochkov, 2003

Listrophoroides (Marquesania) apomys Boch-kov et OConnor, 2008

Listrophoroides (Marquesania) bullimus Boch-kov et OConnor, 2008

Listrophoroides (Marquesania) corpirzvarosae Bochkov et OConnor, 2008

Listrophoroides (Marquesania) faini Bochkov et OConnor, 2008

Listrophoroides (Marquesania) heaneyi Boch-kov et OConnor, 2008

Listrophoroides (Marquesania) 1imnomys Bochkov et OConnor, 2008

Listrophoroides (Marquesania) luzonicus Bochkov et OConnor, 2008

Listrophoroides (Marquesania) melomys Boch-kov et Fain, 2003

Listrophoroides (Marquesania) mordax Boch-kov et Fain, 2003

Listrophoroides (Marquesania) echiothrix Bochkov, OConnor et Gorog, 2004

Listrophoroides (Marquesania) leneothrix Bochkov, OConnor et Gorog, 2004

Listrophoroides (Marquesania) paracucullatus Bochkov et OConnor, 2008

Listrophoroides (Marquesania) pseudolativen-tris Bochkov, OConnor et Gorog, 2004

Listrophoroides (Olistrophoroides) oenomiph-ilus Fain et Bochkov, 2003

Listrophoroides (Olistrophoroides) thallomys Fain et Bochkov, 2003

Family Chirodiscidae

Schizocarpus anomalus Bochkov, 1993

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Schizocarpus azasicus Bochkov et Saveljev, 2012Schizocarpus daberti Bochkov et Saveljev, 2012Schizocarpus heatherae Bochkov, Labrzycka,

Skoracki et Saveljev, 2012Schizocarpus heideckei Bochkov et Saveljev, 2012Schizocarpus faini Bochkov, Labrzycka, Skoracki

et Saveljev, 2012Schizocarpus gozdziewskii Bochkov, Labrzyc-

ka, Skoracki et Saveljev, 2012Schizocarpus klompeni Bochkov, Labrzycka,

Skoracki et Saveljev, 2012Schizocarpus lavrovi Bochkov et Saveljev, 2012Schizocarpus parahumilis Bochkov, Labrzyc-

ka, Skoracki et Saveljev, 2012Schizocarpus pseudonumerosus Bochkov,

Labrzycka, Skoracki et Saveljev, 2012Schizocarpus saveljevi Bochkov, 2012Schizocarpus testiculatus Bochkov, Labrzycka,

Skoracki et Saveljev, 2012Schizocarpus tuvinicus Bochkov et Saveljev, 2012Schizocarpus unzhakovi Bochkov et Saveljev,

2012Schizocarpus zurowskii Bochkov, Labrzycka,

Skoracki et Saveljev, 2012Schizocoptes daberti Bochkov, 2016

Soricilichus sylvisorex Bochkov, Mbaltini et Verheyen, 2016

Family Chirorhynchobiidae

Chirorhynchobia glossophaga Bochkov, Klom-pen et OConnor, 2008

Family Dromiciocoptidae

Dromiciocoptes caenolestes Bochkov et OCon-nor, 2008

Dromiciocoptes marmosops Bochkov et OCon-nor, 2008

Family Gastronyssidae

Eidolonyssus myersi Bochkov et OConnor, 2008

Gastronyssus philippinensis Bochkov et OCon-nor, 2008

Mycteronyssus haplonycteri Bochkov et OCon-nor, 2008

Opsonyssus klompeni Bochkov et OConnor, 2008

Opsonyssus macroglossus Bochkov et OCon-nor, 2008

Opsonyssus pseudoindicus Bochkov et OCon-nor, 2008

Opsonyssus pteropodi Bochkov et OConnor, 2008

Rodhainyssus saccopteryx Bochkov et OCon-nor, 2008

Yunkeracarus apomys Bochkov et OConnor, 2008

Yunkeracarus hylomyscus Bochkov et OCon-nor, 2008

Yunkeracarus limnomys Bochkov et OConnor, 2008

Yunkeracarus lophuromys Bochkov et OCon-nor, 2008

Yunkeracarus otomys Bochkov et OConnor, 2008

Yunkeracarus rattus Bochkov et OConnor, 2008

Family Listrophoridae

Aeromychirus fimbriatus Bochkov et OConnor, 2005

Aeromychirus petinomys Bochkov et OConnor, 2006

Afrolistrophorus (Afrolistrophorus) laonastes Bochkov, Abramov et Hugot, 2011

Afrolistrophorus (Afrolistrophorus) hapalomys Bochkov et Abramov, 2014

Afrolistrophorus (Afrolistrophorus) saxatilo-mys Bochkov, Abramov et Hugot, 2011

Amlistrophorus geoxus Sikora et Bochkov, 2012

Asiochirus nepalensis Fain et Bochkov, 2003Asiochirus soriculus Fain et Bochkov, 2003

Dubininetta abramovi Bochkov, 2017Dubininetta euroscaptor Bochkov, 2017

Hemigalichus chrotogale Bochkov et OCon-nor, 2005

Lynxacarus (Lutracarus) visoni Fain et Boch-kov, 2002

Lynxacarus (Lynxacarus) serrafreirei Bochkov et Valim, 2016

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136

Prolistrophorus (Aprolistrophorus) parabiden-tatus Bochkov, Lareschi et Barreto, 2014

Prolistrophorus (Aprolistrophorus) tylomys Bochkov, Lareschi et Barreto, 2014

Prolistrophorus (Prolistrophorus) hylaeamys Bochkov et Valim, 2016

Pteromychirus menetes Bochkov, Abramov et Hugot, 2011

Family Listropsoralgidae

Didelphialges metachirus Bochkov, OConnor et Grootaert, 2013

Listropsoralges brevisetosus Bochkov, OCon-nor et Grootaert, 2013

Listropsoralges caenolestes Bochkov, OCon-nor et Grootaert, 2013

Listropsoralges caluromys Bochkov et Wau-thy, 2009

Listropsoralges faini Bochkov et Wauthy, 2009Listropsoralges similis Bochkov, OConnor et

Grootaert, 2013Listropsoralges thylamys Bochkov, OConnor

et Grootaert, 2013Listropsoralges vossi Bochkov, OConnor et

Grootaert, 2013

Family Myocoptidae

Myocoptes (Comyocoptes) nesokia Bochkov et OConnor, 2017

Myocoptes (Myocoptes) lophuromys Bochkov, OConnor et Scoracki, 2016 (2017)**

Myocoptes (Myocoptes) meriones Bochkov, 2016Myocoptes (Myocoptes) pitymys Fain et Boch-

kov, 2004

Trichoecius calomysci Bochkov, Malikov et Arbobi, 1999

Trichoecius dubininae Bochkov, 2016Trichoecius faini Bochkov, OConnor et

Skoracki, 2016 (2017)**Trichoecius hylomyscus Bochkov, OConnor et

Skoracki, 2016 (2017)**Trichoecius lemmus Bochkov, 2016Trichoecius meriones Bochkov, 2016

Family Sarcoptidae

Macropodicoptes mironovi Bochkov, 2012

PROSTIGMATASuperfamily CheyletoideaFamily Cheyletidae

Apodicheles collocalia Bochkov et OConnor, 2003

Bak indonesiensis Bochkov et Otto, 2010Bak nadchatrami Fain et Bochkov, 2001

Chelacaropsis kenyensis Fain et Bochkov, 2001Chelacheles aigamuxa Bochkov et OConnor, 2004Chelacheles algericus Fain et Bochkov, 2001Chelacheles apanaskevichi Bochkov et Abramov,

2016Chelacheles bacchusi Bochkov, Khaustov et

Kuznetzov, 1999Chelacheles indra Bochkov et Otto, 2010Chelacheles klimovi Bochkov et OConnor, 2004 Chelacheles seminole Bochkov et OConnor, 2004Chelacheles temoak Bochkov et OConnor, 2004Chelacheles thomasi Bochkov et Otto, 2010

Cheletomimus (Hemicheyletia) ascutatus Fain et Bochkov, 2002

Cheletomimus (Hemicheyletia) crinitus Bochkov et Sidorchuk, 2016

Cheletomimus (Hemicheyletia) gracilis Fain et Bochkov, 2002

Cheletomimus (Hemicheyletia) greenwoodi Fain et Bochkov, 2002

Cheletomimus (Hemicheyletia) ochoai Fain et Bochkov, 2002

Cheletomimus (Philippicheyla) crowei Bochkov et Otto, 2010

Cheletophyes vespae Fain et Bochkov, 2001Cheletophyes decorus Bochkov et Klimov, 2005Cheletophyes mbomba Bochkov, Klimov et

OConnor, 2006Cheletophyes occisor Bochkov et Klimov, 2005Cheletophyes torridae Bochkov, Klimov et OCon-

nor, 2006

Cheletopsis charadrii Mironov, Bochkov et Chi-rov, 1991

Cheletopsis daberti Kivganov et Bochkov, 1994Cheletopsis larosterna Bochkov et Skoracki, 2012Cheletopsis limnodromi Bochkov, Fain et Dabert,

2002Cheletopsis mariae Mironov, Bochkov et Chirov,

1991

S.V. Mironov et al.

137

Cheletopsis prosobonialis Bochkov, Fain et Da-bert, 2002

Cheletopsis rynchops Bochkov, Fain et Dabert, 2002

Cheletopsis thalasseus Bochkov et OConnor, 2003

Cheletosoma tauraco Bochkov et Skoracki, 2012

Cheyletus ascutatus Bochkov et OConnor, 2003Cheyletus batomysi Bochkov et OConnor, 2003Cheyletus kuznetzovi Bochkov et Khaustov, 1999Cheyletus volgini Fain et Bochkov, 2001

Eucheyletiella daurica Bochkov et Mironov, 1999Eucheyletiella faini Bochkov et Malikov, 1996Eucheyletiella pallasius Bochkov et Mironov,

1999Eucheyletiella pusillinus Bochkov et Mironov,

1999

Galagocheles lawrencei Bochkov, Mironov et OConnor, 2005

Granulocheyletus corpuzrarosae Fain et Boch-kov, 2002

Hoffmannita gersoni Fain et Bochkov, 2001

Kenyacheylus troglodytes Fain et Bochkov, 2001

Ker afrotropicalis Fain et Bochkov, 2001

Metacheyletia amazonae Bochkov et OCon-nor, 2003

Metacheyletia degenerata Fain et Bochkov, 2003Metacheyletia ngaii Bochkov et Skoracki, 2011

Mexecheles shiva Bochkov et Ochoa, 2005

Neochelacheles corpuzrarosae Bochkov et OConnor, 2004

Neochelacheles mendicus Bochkov et OCon-nor, 2004

Neocheyletiella aegithali Bochkov et OConnor, 2003

Neocheyletiella ardeola Bochkov et Ochoa, 2005

Neocheyletiella athene Fain et Bochkov, 2002Neocheyletiella lonchurae Fain et Bochkov, 2002Neocheyletiella nectarinia Bochkov et OCon-

nor, 2003

Neocheyletiella parvisetosa Mertins et Boch-kov, 2014

Neocheyletiella queleae Fain et Bochkov, 2002

Nidocheyletus kennedyae Bochkov et OCon-nor, 2004

Nodele (Aztecocheyletus) conquistador Boch-kov et Klimov, 2004

Oconnoricheylus chimaera (Bochkov et Ochoa, 2005)

Oconnoricheylus speciosus Bochkov et Otto, 2010

Ornithocheyletia columbigallinae Fain et Boch-kov, 2002

Ornithocheyletia mironovi Bochkov et Chirov, 1998

Ornithocheyletia orioli Fain et Bochkov, 2002Ornithocheyletia prognae Bochkov et Gallo-

way, 2001Ornithocheyletia turdi Bochkov et Galloway, 2001Ornithocheyletia wauthyi Fain et Bochkov, 2002

Picocheyletus tricholaemae Bochkov et OCon-nor, 2003

Prosocheyla (Prosocheyla) ripkai Fain et Boch-kov, 2001

Prosocheyla (Prosocheyla) villosa Bochkov et Khaustov, 1999

Samsinakia trilobitus Bochkov et Mironov, 1998Samsinakia tilae Bochkov et Klimov, 2005

Sciurocheyla funisciurus Bochkov et OConnor, 2004

Thryonomycheyla angolensis Fain et Bochkov, 2002

Family Cloacaridae

Caminacarus dawsoni Bochkov et OConnor, 2008

Family Epymyodicidae

Epimyodex soricis Fain et Bochkov, 2001

Family Syringophilidae

Aulobia galinago Bochkov et Mironov, 1998Aulobia sylviae Bochkov et Mironov, 1998

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138

Aulonastus prunellae Bochkov et Mironov, 1999

Castosyringophilus forpi Bochkov et Perez, 2002

Corvitorotroglus alpha Skoracki et Bochkov, 2010

Bubophilus asiobius Skoracki et Bochkov, 2002

Charadriphilus ralli Skoracki et Bochkov, 2010Charadriaulobia vanelli Bochkov, Fain et

Skoracki, 2004

Charadriphilus lyudmilae Bochkov et Chistya-kov, 2001

Chenophila kanduli Bochkov, 1998

Fritschisyringophilus lonchurae Bochkov, Fain et Skoracki, 2004

Megasyringophilus cyanocephala Fain, Boch-kov et Mironov, 2000

Megasyringophilus dubinini Bochkov et Fain, 2003

Megasyringophilus kethleyi Fain, Bochkov et Mironov, 2000

Megasyringophilus platycercus Bochkov et Fain, 2003

Megasyringophilus rhynchopsitta Bochkov et Perez, 2002

Megasyringophilus trichoglossus Fain, Boch-kov et Mironov, 2000

Mironovia coturnae Bochkov, Fain et Skoracki, 2004

Mironovia lagopus Bochkov et Skirnisson, 2011

Neoaulobia agapornis Fain, Bochkov et Mironov, 2000

Neoaulobia aratingae Fain, Bochkov et Mironov, 2000

Neoaulobia mexicana Bochkov et Perez, 2002Neoaulobia mironovi Bochkov et Perez, 2002Neoaulobia psittaculae Fain, Bochkov et Mironov,

2000

Niglarobia calidridis Bochkov et Mironov, 1998Niglarobia chirovi Bochkov et Mironov, 1998Niglarobia rhinoptili Fain, Bochkov et Mironov,

2000Niglarobia skorackii Bochkov et Galloway, 2004

Phalarophilus fulicarius Skoracki, Bochkov et OConnor, 2011

Philoxanthornia dubinini Bochkov et Mironov, 1998

Picobia alectoris Fain, Bochkov et Mironov, 2000Picobia biarmicus Skoracki, Bochkov et Wau-

thy, 2004Picobia brotogeris Fain, Bochkov et Mironov,

2000Picobia chloris Bochkov, Mironov et Kravtso-

va, 2000Picobia cissa Skoracki, Bochkov et Wauthy,

2004Picobia locustella Skoracki, Bochkov et Wau-

thy, 2004Picobia phoeniculi Fain, Bochkov et Mironov,

2000Picobia ramphastos Fain, Bochkov et Mironov,

2000Picobia sturni Skoracki, Bochkov et Wauthy,

2004

Psittaciphilus amazonae Fain, Bochkov et Mironov, 2000

Psittaciphilus fritschi Fain, Bochkov et Mironov, 2000

Stibarokris langei Bochkov et Mironov, 1999

Syringophiloidus agelaius Bochkov, Skoracki, Hendricks et Spicer, 2011

Syringophiloidus carpodaci Bochkov et Ap-anaskevich, 2001

Syringophiloidus cypsiuri Fain, Bochkov et Mironov, 2000

Syringophiloidus daberti Bochkov Bochkov, Fain et Skoracki, 2004

Syringophiloidus delichorum Bochkov, 2001Syringophiloidus dendrocittae Fain, Bochkov

et Mironov, 2000Syringophiloidus graculae Fain, Bochkov et

Mironov, 2000Syringophiloidus jackowiaki Bochkov, Skoracki,

Hendricks et Spicer, 2011Syringophiloidus klimovi Skoracki et Bochkov,

2010Syringophiloidus motacillae Bochkov et Mironov,

1998Syringophiloidus serini Bochkov, Fain et Skoracki,

2004Syringophiloidus xanthocephalus Bochkov,

Skoracki, Hendricks et Spicer, 2011Syringophiloidus zonotrichia Bochkov, Skoracki,

Hendricks et Spicer, 2011

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Syringophilopsis aegythali Bochkov, Mironov et Skoracki, 2001

Syringophilopsis borini Bochkov et Mironov, 1999

Syringophilopsis dendroicae Bochkov et Gal-loway, 2001

Syringophilopsis emberizae Fain, Bochkov et Mironov, 2000

Syringophilopsis faini Bochkov et Apanaskev-ich, 2001

Syringophilopsis garrulaxi Bochkov, Mironov et Skoracki, 2001

Syringophilopsis icteri Bochkov et Galloway, 2001

Syringophilopsis kirgizorum Bochkov, Mironov et Kravtsova, 2000

Syringophilopsis phylloscopi Bochkov, Mironov et Skoracki, 2001

Syringophilopsis porzanae Bochkov et Gal-loway, 2004

Syringophilopsis stachyris Bochkov, Mironov et Skoracki, 2001

Syringophilopsis sylviettae Fain, Bochkov et Mironov, 2000

Syringophilopsis tyrrani Bochkov et Galloway, 2004

Syringophilus numidae Bochkov, 2000

Terratosyringophilus lorricinus Bochkov et Fain, 2002

Terratosyringophilus pioni Bochkov et Perez, 2002

Torotrogla aphelocoma Bochkov, Flannery et Spicer, 2009

Torotrogla cardinalis Bochkov, Flannery et Spicer, 2009

Torotrogla cardueli Bochkov et Mironov, 1999Torotrogla coccothraustes Bochkov, Flannery

et Spicer, 2009Torotrogla cyanocitta Bochkov, Flannery et

Spicer, 2009Torotrogla gaudi Bochkov et Mironov, 1998

Torotrogla piranga Bochkov, Flannery et Spicer, 2009

Family Harpirhynchidae

Anharpyrhynchus apodus Bochkov et Klom-pen, 2014

Anharpyrhynchus elizae Bochkov et Klompen, 2014

Anharpyrhynchus lukoschusi Bochkov et Klompen, 2014

Crassacarus alexfaini Bochkov, OConnor et Klompen, 2015

Crassacarus fritschi Bochkov, OConnor et Klompen, 2015

Crassacarus melanerpes Bochkov, OConnor et Klompen, 2015

Crassacarus sialia Bochkov, OConnor et Klompen, 2015

Crassacarus tinae Bochkov, OConnor et Klom-pen, 2015

Fainharpirhynchus contopus Bochkov et Gal-loway, 2013

Fainharpirhynchus legatus Bochkov et Klom-pen, 2014

Fainharpirhynchus mossi Bochkov et Klom-pen, 2014

Harpirhynchus dusbabeki Bochkov et Literak, 2006

Harpirhynchus galleridae Fain, Bochkov et Mironov, 1999

Harpirhynchus quasimodo Bochkov et Mer-tins, 2010

Harpyrhynchoides accipiter Bochkov et OCon-nor, 2013

Harpyrhynchoides actitis Bochkov, OConnor et Klompen, 2015

Harpyrhynchoides aegolius Bochkov et OCon-nor, 2013

Harpyrhynchoides alaudinus Bochkov, 2000Harpyrhynchoides artamus Bochkov et Klom-

pen, 2015Harpyrhynchoides athene Bochkov, OConnor

et Klompen, 2015Harpyrhynchoides botaurus Bochkov et Gal-

loway, 2013Harpyrhynchoides bubulcus Bochkov, OCon-

nor et Klompen, 2015Harpyrhynchoides calidris Bochkov, OConnor

et Klompen, 2015Harpyrhynchoides charadrius Bochkov,

OConnor et Klompen, 2015Harpyrhynchoides clamator Bochkov, 2014Harpyrhynchoides coccyzus Bochkov, OCon-

nor et Klompen, 2015Harpyrhynchoides crotophaga Bochkov, OCon-

nor et Klompen, 2015Harpyrhynchoides heatherae Bochkov et Gal-

loway, 2013

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Harpyrhynchoides gallowayi Bochkov, OCon-nor et Klompen, 2015

Harpyrhynchoides ixobrychus Bochkov, OCon-nor et Klompen, 2015

Harpyrhynchoides kirgizorum Fain, Bochkov et Mironov, 1999, syn. of H. zumpti (Fain, 1972)

Harpyrhynchoides megascops Bochkov, OCon-nor et Klompen, 2015

Harpyrhynchoides parazumpti Fain, Bochkov et Mironov, 1999

Harpyrhynchoides pluvialis Bochkov, OCon-nor et Klompen, 2015

Harpyrhynchoides setophaga Bochkov et Klom-pen, 2014

Harpyrhynchoides spizella Bochkov et Klom-pen, 2014

Harpyrhynchoides vulgaris Bochkov et Gal-loway, 2004

Harpyrhynchoides xanthocephalus Bochkov et Klompen, 2014

Harpyrhynchoides zenaida Bochkov, OConnor et Klompen, 2015

Harpypalpus lonchura Bochkov et Klompen, 2014

Harpypalpus pyrrhula Bochkov et Klompen, 2014Harpypalpus spermestes Fain, Bochkov et

Mironov, 1999Harpypalpus sturnus Bochkov et Klompen,

2014Harpypalpus taeniopygia Bochkov et Klom-

pen, 2014

Harpypalpoides hirundinoides Bochkov et Klompen, 2014

Harpypalpoides johnstoni Bochkov et Klom-pen, 2014

Harpypalpoides lesickii Skoracki, Bochkov et Sikora, 2004

Harpypalpoides namibiensis Fain, Bochkov et Mironov, 1999

Harpypalpoides regulus Bochkov et Klompen, 2014

Harpypalpoides sitta Bochkov et Klompen, 2014

Harpypalpoides sylvia Bochkov et Klompen, 2014

Harpyrhynchiella apus Bochkov, 2014

Neharpyrhynchus agelaius Bochkov, OConnor et Klompen, 2015

Neharpyrhynchus baile Bochkov et Literak, 2007

Neharpyrhynchus bombycilla Bochkov, OCon-nor et Klompen, 2015

Neharpyrhynchus campylorhynchus Bochkov, OConnor et Klompen, 2015

Neharpyrhynchus chlorospingus Bochkov et Literak, 2011

Neharpyrhynchus domrowi Bochkov et Klom-pen, 2015

Neharpyrhynchus hippolae Bochkov, 2000Neharpyrhynchus icterus Bochkov et OConnor,

2013Neharpyrhynchus loxia Bochkov et Galloway,

2013Neharpyrhynchus mironovi Bochkov et Liter-

ak, 2011Neharpyrhynchus oenanthe Bochkov, 2014Neharpyrhynchus picidarum Bochkov, OCon-

nor et Klompen, 2015Neharpyrhynchus quiscalus Bochkov, OCon-

nor et Klompen, 2015Neharpyrhynchus spizella Bochkov, OConnor

et Klompen, 2015Neharpyrhynchus tangara Bochkov et Literak,

2011Neharpyrhynchus turdus Bochkov, OConnor

et Klompen, 2015Neharpyrhynchus vireo Bochkov, Oconnor et

Klompen, 2015

Perharpyrhynchus caprimulgus Bochkov et Klompen, 2014

Perharpyrhynchus charadrius Bochkov, OCon-nor et Klompen, 2015

Perharpyrhynchus elseyornis Bochkov et Klom-pen, 2014

Trichorhynchiella myiarchus Bochkov et Klom-pen, 2014

Trichorhynchiella phleocryptes Bochkov et OConnor, 2014

Superfamily MyobioideaFamily Myobiidae

Caenolestomyobia faini Bochkov et OConnor, 2009

Caenolestomyobia lukoschusi Bochkov et OCon-nor, 2009

Crocidurobia (Crocidurobia) dusbabeki Bochkov et Fain, 2003

Crocidurobia (Crocidurobia) lasiurae Bochkov, 1997

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Crocidurobia (Crocidurobia) leucodoni Boch-kov, 1997

Crocidurobia (Crocidurobia) lukoschusi Boch-kov, 2017

Eudusbabekia (Eudusbabekia) rhinophylla Fain et Bochkov, 2002

Ewingana (Ewingana) teniotis Bochkov et Rybin, 1992

Hylomysobia chinensis Bochkov et OConnor, 2006

Hylomysobia mikhailzaitzevi Bochkov et OCon-nor, 2006

†Protohylomysobia erinaceophilus Sidorchuk et Bochkov, 2018 (2019)**

Protomyobia (Protomyobia) caecutiens Boch-kov, 1994

Protomyobia (Protomyobia) jamesoni Boch-kov, 1994

Radfordia (Austromyobia) dubinini Bochkov, Dubinina et Chirov, 1990

Radfordia (Austromyobia) merioni Bochkov, Dubinina et Chirov, 1990

Radfordia (Austromyobia) persica Bochkov, Arbobi et Malikov, 2000

Radfordia (Dipodomyobia) alactaguli Boch-kov, 1997

Radfordia (Dipodomyobia) allactodipi Boch-kov, 1997

Radfordia (Dipodomyobia) baranovae Boch-kov, 1997

Radfordia (Dipodomyobia) dipi Bochkov, 1997Radfordia (Dipodomyobia) majori Bochkov, 1997Radfordia (Dipodomyobia) paradipi Bochkov,

1997Radfordia (Dipodomyobia) paralactaga Boch-

kov, 1997Radfordia (Dipodomyobia) pygeretmusi Boch-

kov, 1997Radfordia (Dipodomyobia) stylodipi Bochkov,

1997Radfordia (Graphiurobia) myomimusi Boch-

kov, 1994Radfordia (Graphiurobia) selevinia Bochkov,

2009Radfordia (Hesperomyobia) akodon Bochkov,

2017Radfordia (Hesperomyobia) burbanoae Boch-

kov et Barreto, 2014

Radfordia (Hesperomyobia) rhipidomys Boch-kov et Barreto, 2014

Radfordia (Microtimyobia) abramovi Bochkov et Mironov, 1998

Radfordia (Microtimyobia) alticolae Bochkov, 1995

Radfordia (Microtimyobia) cricetuliphila Bochkov, 1999

Radfordia (Microtimyobia) dinaromys Boch-kov, 2011

Radfordia (Microtimyobia) golenishchevi Bochkov, 2011

Radfordia (Microtimyobia) myopusi Bochkov et Mironov, 1998

Radfordia (Microtimyobia) pitymys Bochkov, 2011

Radfordia (Microtimyobia) rufocani Bochkov, 1995

Radfordia (Microtimyobia) stekolnikovi Bochkov et Mironov, 1998, syn. of R. lemnina (Koch, 1841)

Radfordia (Microtimyobia) stenocrani Bochkov et Mironov, 1998, syn. of R. lemnina (Koch, 1841)

Radfordia (Microtimyobia) synaptomysi Boch-kov et Mironov, 1998

Radfordia (Neotomobia) megadontomys Boch-kov et Guzman-Cornejo, 2014

Radfordia (Neotomobia) onychomys Bochkov et Guzman-Cornejo, 2014

Radfordia (Neotomobia) peromyscus Bochkov et Guzman-Cornejo, 2014

Radfordia (Radfordia) berylmysi Bochkov et Fain, 1997

Radfordia (Radfordia) colomys Bochkov et Fain, 2003

Radfordia (Radfordia) delectori Bochkov et Fain, 2003

Radfordia (Radfordia) dephomys Bochkov et Fain, 2003

Radfordia (Radfordia) eremici Fain et Boch-kov, 2002

Radfordia (Radfordia) lukoschusi Bochkov et Fain, 1997

Radfordia (Radfordia) mirabilis Bochkov et Abramov, 2014

Radfordia (Radfordia) mironovi Bochkov, 1997

Radfordia (Radfordia) myomysci Bochkov et Fain, 2003

Radfordia (Radfordia) niviventris Bochkov et Fain, 1997

Radfordia (Radfordia) sigmomys Bochkov et Fain, 2002

In memoriam of Andrei V. Bochkov

142

Superfamily ParatydeoideaFamily Stigmocheylidae

Stigmocheylus americanus Bochkov, 2008Stigmocheylus oberon Bochkov, 2008

Superfamily PomerantzioideaFamily Pomerantziidae

Pomerantzia philippina Bochkov et Walter, 2007

Superfamily PterygosomatoideaFamily Pterygosomatidae

Geckobia anocellatus Bochkov et Mironov, 2000

Geckobia hirsti Bochkov et Mironov, 2000

Pterygosoma pseudotrapelus Bochkov, Mel-nikov et Nazarov, 2009

PARASITIFORMESMESOSTIGMATASuperfamily DermanyssoideaFamily Laelapidae

Andreacarus brachyuromys Dowling, Bochkov et OConnor, 2007

Andreacarus eliurus Dowling, Bochkov et OConnor, 2007

Andreacarus galidia Dowling, Bochkov et OConnor, 2007

Andreacarus gymnuromys Dowling, Bochkov et OConnor, 2007

Andreacarus nesomys Dowling, Bochkov et OConnor, 2007

Andreacarus tenrec Dowling, Bochkov et OConnor, 2007

Andreacarus voalavo Dowling, Bochkov et OConnor, 2007

**—printed version appeared the next year after online version; †—fossil species.

MITE TAXA NAMED IN HONOR OF A.V. BOCHKOV

Family-group names

Bochkovocoptidae OConnor, 2019 fam.n. (Astig-mata, Sarcoptoidea), present issue

Genus-group names

Andrebochkovia Khaustov et Frolov, 2019 gen. n. (Prostigmata, Neopygmephoridae), present issue

Bochkovlaster Khaustov, 2019 gen. n. (Prostig-mata, Pygmephoridae), present issue

Bochkovocoptes OConnor, 2019 gen. n. (Astig-mata, Bochkovocoptidae), present issue

Species-group names

Allogalumna bochkovi Ermilov et Starý, 2018 (Oribatida, Galumnidae)

Calcealges bochkovi Dabert, 2019 sp. n. (Astig-mata, Trouessartiidae), present issue

Cunaxa bochkovi Kaustov et Kuznetzov, 1998 (Prostigmata, Cunaxidae)

Eustigmaeus bochkovi Khaustov 2019 (Prostig-mata, Stigmaeidae)

Fringillosphaera bochkovi Mironov, 2001 (Astig-mata, Apionacaridae)

Gaeolaelaps bochkovi Jocharchi, Khaustov et Ermilov, 2019 sp. n. (Mesostigmata, Laelapidae), present issue

Galliphilopsis bochkovi Skoracki et Sikora, 2004 (Prostigmata, Syringophilidae)

Haemaphysalis bochkovi Apanaskevich et Tomlinson, 2019 (Ixodida, Ixodidae)

Hypozetes andreii Ermilov, Hugo-Coetzee, Khaus-tov et Kontschán, 2019 sp. n. (Oribatida, Tegoribatidae), present issue

Ipsoglyphus bochkovi Klimov et Khaustov, 2018 (Astigmata, Acaridae)

Leptotrombidium bochkovi Stekolnikov, 2013 (Astigmata, Trombiculidae)

Meristacarus bochkovi Ermilov et Kalúz, 2019 sp. n. (Oribatida, Lohmanniidae), present issue

Neharpyrhynchus bochkovi Martinu, Dusba-bek et Literak, 2008 (Prostigmata, Harpyrhyn-chidae)

Neoeucheyla bochkovi Fain, 2003 (Prostig-mata, Cheyletidae)

Proctophyllodes bochkovi Mironov 2019 sp. n. (Astigmata, Proctophyllodidae), present issue

Ramusella (Dosangoppia) bochkovi Ermilov et Frolov, 2019 (Oribatida, Oppiidae)

Syringophilopsis bochkovi Skoracki, Mironov et Bermúdez, 2019 sp. n. (Prostigmata, Syringo-philidae), present issue

Trouessartia bochkovi Mironov et Galloway 2019 (Astigmata, Trouessartiidae)

S.V. Mironov et al.

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In memoriam of Andrei V. Bochkov

Table 1. Mite taxa described by A.V. Bochkov and coauthors.

Family Subfamily Tribe Genus Subgenus Species TotalASTIGMATAAnalgoidea

1 Dermationidae 2 2 11 152 Epidermoptidae 3 3 63 Psoroptoididae 1 1 24 Pyroglyphidae 1 1

Psoroptoidea

5 Lobalgidae 2 26 Psoroptidae 6 6

Sarcoptoidea

7 Atopomelidae 1 1 53 558 Chirodiscidae 18 189 Chirorhynchobiidae 1 110 Dromiciocoptidae 2 211 Gastronyssidae 1 1 14 1612 Listrophoridae 17 1713 Listropsoralgidae 1 8 914 Myocoptidae 1 10 1115 Sarcoptidae 1 1 2

PROSTIGMATACheyletoidea

16 Cheyletidae 5 1 76 8217 Cloacaridae 1 118 Epymyodicidae 1 119 Harpirhynchidae 2 1 74 7720 Syringophilidae 10 78 88

Myobioidea

21 Myobiidae 1 3 3 8 54 69Paratydeoidea

22 Stigmocheylidae 2 2Pomerantzioidea

23 Pomerantziidae 1 1Pterygosomatoidea

24 Pterygosomatidae 3 3MESOSTIGMATADermanyssoidea

25 Laelapidae 1 7 8Total 3 4 33 11 445 496

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