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APG-III
2009
18S rDNA
rbcL
atpB
atp1
matR
+ 61 genes de 45
táxons
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
APG-III
2009
www.mobot.org/MOBOT/research/APweb (Stevens 2010)
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
APG-III
2009
ANITA Qiu et al. 1999
18S rDNA
rbcL, atpB
atp1, matR
+ 61 marcadores
de 45 táxons
Moore et al. 2007
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Qiu et al. 1999
INDEL matR mitocondrial
A
N
I
T
A
GIMNO
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Amborella trichopoda
AMBORELLACEAE
Philipson 1993
Judd et al. Plant sytematics 2006
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NYMPHAEACEAE
Heywood 1974
J R Pirani
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Barclaya
NYMPHAEACEAE
Heywood 1974
rizoma
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NYMPHAEACEAE Judd et al. 2002
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NYMPHAEACEAE
Victoria amazonica
Heywood 1974
J R Pirani
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NYMPHAEALES
CABOMBACEAE
Cabomba aquatica
Rudall et al. 2009
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NYMPHAEALES
Trithuria submersa
Hydatellaceae
Rudall et al. 2009
Stevens 2010
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Crisp & Cook 2005
Interpretação de árvores de acordo com a topologia
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Crisp & Cook 2005
Interpretação de árvores de acordo com a topologia
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Muitas características ditas “basais” em grupos-irmãos
podem ser independentemente derivadas
e altamente especializadas.
Exemplo:
As adaptações morfológicas e ecofisiológicas de
Nymphaeales ao ambiente aquático.
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
APG-III
2009
www.mobot.org/MOBOT/research/APweb (Stevens 2010)
Cabombaceae
Nymphaeaceae
Hydatellaceae
Austrobaileyaceae
Trimeniaceae
Illiciaceae
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Austrobaileya
scandens
AUSTROBAILEYALES
Austrobaileyaceae Endress 1993
Trimenia papuana
Trimeniaceae Philipson 1993
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Illicium floridanum, Illiciaceae
AUSTROBAILEYALES
Cronquist 1981
Keng 1993
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AUSTROBAILEYALES
Illicium floridanum
Illiciaceae
Thien et al. 2009
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VASOS
Meylan &
Butterfield
1972
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
VASOS
Traqueídes (simplesiomorfia)
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Williams &
Friedman
2004
AUSTROBAILEYALES (grado ANITA)
Maioria das angiospermas
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Williams & Friedman 2004
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
Endosperma
diplóide
Endosperma
triplóide
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Endress 1997
Fechamento
carpelar em
Laurus nobilis
(Lauraceae)
Fusão posgênita
das margens
carpelares
ANGIOSPERMIA
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Endress & Igersheim 2000
ANITA
Tipos de
angiospermia
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Endress & Igersheim 2000
Magnoliídeas
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
Angiospermia
por secreção
fusão
posgênita
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
APG-III
2009
Hedyosmum
brasiliense
Ascarina
flor masculina
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Ceratophyllales
APG-II
2009
www.mobot.org/MOBOT/research/APweb (Stevens 2010)
Magnoliídeas 2%
alcalóides
benzilisoquinolínicos;
céls antipodais efêmeras
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MAGNOLIALES 6 famílias
3000 spp.
MAGNOLIACEAE -folhas alternas
- estípulas
-células oleíferas
Heywood 1976
J R Pirani
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Liriodendron tulipifera
MAGNOLIACEAE
K3 C3+3 A∞ G∞ *
Judd et al. 2002
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MAGNOLIALES
MAGNOLIACEAE
Magnolia grandiflora
Flor na abertura
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MAGNOLIALES
MAGNOLIACEAE
Magnolia grandiflora
Flor após fecundação
A.B. Joly
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MAGNOLIACEAE
Magnolia grandiflora
FRUTO
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Pólen
Fruto polifolículo;
sementes pêndulas
por longo funículo;
zoocoria
Cronquist 1978
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Talauma ovata
MAGNOLIACEAE
J.R. Pirani
A.B. Joly
Heywood 1976
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Michelia figo
Michelia champaca
MAGNOLIACEAE
A. B. Joly
A. B. Joly
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MAGNOLIALES
ANNONACEAE
Heywood 1976
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MAGNOLIALES
ANNONACEAE Xylopia aromatica
K3 C3+3 A∞ G1-∞ J. R. Pirani
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ANNONACEAE Guatteria
Annona
A. B. Joly
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Gottsberger & Gottsberger 1988
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ANNONACEAE
frutos
Judd et al. 2002
A. B. Joly
Strassburger 1956
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Duguetia marcgraviana
Annona muricata A. B. Joly
J. R. Pirani
Strassburger 1956
Heywood 1974
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MAGNOLIALES
diagrama floral
-flor estrobilóide (estames e carpelos espiralados)
- cantarofilia
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Donoghue & Doyle 1989 Morfologia
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Amborellaceae
Nymphaeaceae
Austrobaileyales
Chloranthaceae
Monocotiledôneas
Eudicotiledôneas
Magnoliideae
Ceratophyllales
exina
granular
exina
columelada
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Ceratophyllales
APG-II
2009
www.mobot.org/MOBOT/research/APweb (Stevens 2010)
Magnoliídeas 2%
alcalóides
benzilisoquinolínicos
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LAURALES 7 famílias
2850 spp.
Novidades evolutivas:
Hipanto (flores períginas a epíginas)
Estaminódios
Pólen inaperturado
Cinnamomum
camphora Erdtman 1952
Kubitzki 1990
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LAURALES
LAURACEAE 2500 spp.
Heywood 1976
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LAURACEAE
Cinnamomum
Umbellularia
A. B. Joly
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LAURACEAE
Cinnamomum camphora
Endress 1994
Sassafras
Umbellularia
Judd et al. 2002
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LAURACEAE
Kubitzki 1990
hipanto
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LAURACEAE Heywood 1974
Cinnamomum
Ocotea
Persea
J>R. Pirani
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LAURALES
LAURACEAE
Cassytha
holoparasita
Não confundir
com cipó-chumbo
(Cuscuta
Convolvulaceae,
Asterídeas)
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Amborellales
Nymphaeales
Austrobaileyales
Chloranthales
Monocotiledôneas
Eudicotiledôneas
Ceratophyllales
APG-II
2009
www.mobot.org/MOBOT/research/APweb (Stevens 2010)
Magnoliídeas 2%
alcalóides
benzilisoquinolínicos
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Piper nigrum
Novidade evolutiva:
câmbio pouco ativo
(lianas ou ervas)
PIPERALES 4 famílias
4090 spp.
Heywood 1976
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http://pt.scribd.com/doc/7229952/Angiospermas-Basais
PIPERALES PIPERACEAE
Piper
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PIPERALES
PIPERACEAE
Peperomia
polybotrya
Cronquist 1988
Piper
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PIPERACEAE
Pólen
A. Peperomia
B. Heckeria
C. Piper
Erdtman 1952
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Figueiredo & Sazima
2000
PIPERACEAE
polinização
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PIPERALES
Heywood 1976 A.B. Joly
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ARISTOLOCHIACEAE Heywood 1976
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Megaselia Phoridae PIPERALES - ARISTOLOCHIACEAE
Aristolochia
gigantea
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ARISTOLOCHIACEAE
A.B. Joly
J.R.Pirani
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Donoghue & Doyle 1989 Morfologia
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Donoghue & Doyle 1989 Morfologia
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Donoghue & Doyle 1989 Morfologia
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Amborellales
Nymphaeales
Austrobaileyales
APG III (2009)
sistema
caulinar
atactostélico Chlorophytum
Piper
Piperales
Chloranthaceae
Canellales Aristolochiaceae
Hydnoraceae
Piperaceae
Saururaceae
Laurales
Magnoliales
MONOCOTILEDÔNEAS
Ceratophyllales
EUDICOTILEDÔNEAS
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Prosopanche
Hydnora
africana
Prosopanche
HYDNORACEAE: endoparasaitas, inclusão
ecente na ordem
2 gen/ 7 spp.
PIPERALES
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Amborellaceae
Nymphaeaceae
Austrobaileyales
Chloranthaceae
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Ceratophyllales
alcalóides
benzilisoquinolínicos
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CANELLALES - 2 fam. 93 spp. - sinapom.= K e C distintos
WINTERACEAE
Drymis
Judd et al. 2002 plant systematics
A.B. Joly
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Doyle & Endress 2000
VASOS
ANITA
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Amborellales
Nymphaeales
Austrobaileyales
Monocotiledôneas
Magnoliídeas
Eudicotiledôneas
(ou Tricolpadas)
APG III 2009
Chloranthales
Ceratophyllales
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Amborellales
Nymphaeales
Austrobaileyales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
APGIII
2009
Chloranthaceae
Ceratophyllales
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Amborellales
Nymphaeales
Austrobaileyales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
APG-III
2009
Chloranthales
? Ceratophyllales
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Endress
1994
Austrobaileya
Austrobaileyaceae
Exospermum
Winteraceae
Kitaibelia
Malvaceae
EUDICOT.
Annona
Annonaceae
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Amborellaceae
Nymphaeaceae
Austrobaileyales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Chloranthaceae
androceu e
gineceu cíclicos
Ceratophyllales
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Aquilegia atrata
(Ranunculaceae)
Drymis winteri
(Winteraceae)
APOCARPIA
Leins 2000
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Hypericum hookerianum
(Guttiferae)
SINCARPIA - conação possibilitada pela
80% das angiospermas organização cíclica
Leins 2000
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Amborellaceae
Nymphaeaceae
Austrobaileyales
Monocotiledôneas
Eudicotiledôneas
Magnoliídeas
Chloranthaceae
Ceratophyllales
SINCARPIA
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