Bulletin 77

New Mexico Museum of Natural History & Science

A Division of the

DEPARTMENT OF CULTURAL AFFAIRS

Carboniferous-Permian transition in Socorro County, New Mexico

Edited by

Spencer G. Lucas, William A. DiMichele and Karl Krainer

Albuquerque,2017

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Bulletin 77

New Mexico Museum of Natural History & Science

A Division of the DEPARTMENT OF CULTURAL AFFAIRS

Carboniferous-Permian transition in Socorro County, New Mexico

Edited by Spencer G. Lucas, William A. DiMichele and Karl Krainer

New Mexico Museum of Natural History & Science Albuquerque, 2017

UNIVERSITY OF NEW MEXICO

STATE OF NEW MEXICO Department of Cultural Affairs Veronica Gonzales, Secreta,y

NEW MEXICO MUSEUM OF NATURAL HISTORY AND SCIENCE Margaret Marino, Executive Director

BOARD OF TRUSTEES Susanna Martinez, Governor, State of New Mexico, ex officio

Margaret Marino, Executive Director, ex officio Gary Friedman, President

Leonard Duda Maya Elrick, Ph.D.

Peter F. Gerity, Ph.D. Deron Knoner

Laurence Lattman, Ph.D. Viola Martinez

Mike Mertz Marvin Moss

John Montgomery, Ph.D. Jennifer Riordan

Laura Smigielski-Garcia Steve West

Cover illustration: Pinna of the medullosan pteridosperm Neuropteris ovata from the Tinajas black shale,

Missourian age, Socorro County, New Mexico. Neuropteris ova/a was a geographically widespread, temporally

long-ranging species that typically grew in wetland habitats across the Euramerican portion of the Pangean

equatorial region. New Mexico Museum ofNatural History and Science specimen P-40166.

Original Printing lSSN: 1524-4156

Available from the New Mexico Museum of Natural History and Science, 180 I Mountain Road NW, Albuquerque, NM 87104; Telephone (505) 841-2800; Fax (505) 841-2866; www.nmnaturalhistory.org

NMMNH Bulletins online at: http://nmnaturalhistory.org/bullctins and on Google Books

BULLETIN OF THE NEW MEXICO MUSEUM

OF NATURAL HISTORY AND SCIENCE

Spencer G. Lucas

Robert Sullivan Lawrence H. Tanner

Asher J . Lichtig

Guillermo Alvarado Marco Avanzini David Berman Brent Breithaupt William DiMichele John R. Foster Gerard Gierlinski Jean Guex Jerald D. Harris Andrew 8. Heckert Adrian P. Hunt Hendrik Klein Karl Krainer Martin G. Lockley Claudia Marsicano Gary S. Morgan Donald R. Prothero Silvio Renesto Joerg W. Schneider Jingeng Sha Sebastian Voigt

Ralf Werneburg

Richard S. White, Jr.

EDITORS

New Mexico Museum of Natural History and Science, Albuquerque, NM, USA (NMMNHS) NMMNHS Le Moyne College. Syracuse, NY, USA

MANAGING EDITOR

NMMNHS

ASSOCIATE EDITORS

Asociacion Costarricense de Gcotccnica, San Jose, Costa Rica Musco Tridcntino di Scicnzc Naturali, Trento, Italy Carnegie Museum of Natural History, Pittsburgh, PA. USA Laramie, WY. USA National Museum orNatural History, Washington. D.C .. USA Museum of Moab, Moab, Utah Polish Geological Institute, Warsaw, Poland University of Lausanne. Lausanne. Switzerland Dixie State College, Sl. George. UT. USA Appalachian State University, Boone, NC, USA Flying Heritage Collection, E,·crctl. WA, USA Neumarkl, Germany Univt.:rsity of Innsbruck. Innsbruck. Austria University of Colorado at Denver, Denver, CO, USA Univcrsidad de Buenos Aires, Buenos Aires, Argentina

NMMNIIS Los Angeles, CA. USA Universita dcgli Studi dcll'Insubria. Varese, Italy Technical University BergAkadcmie of Freiberg, Freiberg, Germany Nanjing Institute of Geology and Palaeontology, Nanjing, China Urwcltmuscum GEOSKOP/Burg Lichtenburg. Thalliclltenbcrg, Germany Naturhistorischcs Museum Schloss Bcrtholdsburg, Schleusingcn, Germany International Wildlife Museum, Tucson, AZ. USA

NEW MEXICO MUSEUM OF NATURAL HISTORY AND SCIENCE BULLETINS

-10. l riassic of the American West, 2007. edited b) Spencer(,. Lucas and Justin \. Spielmann. 2-17 pp . ..\I. The Global Triassic. 2007. edited by Spt:ncer Ci. Luca� and Justm A Sp1clma11n . ..\ 15 pp. -11. Ccno;,oic Vertebrate Tracb and Traces. 2007. edited by Spencer(., Lucas. Justin A. Spielmann and Martin u. Lockley. 330 pp 43. The Late Triassic archosaummorph Trifop'10.1u11m1. 2008. by Justin,\ Spielmann. Spencer G. Lucas. Larry r. Rinehart and

/\nJrC\\ B. 1 leckert, 177 pp . ..\..\. Neogene Mammals, 2001:t. edited by Spencer G. Lucas. C.,ary S. \forgan. Justin A. Spielmann and Donald R. Prothero . ..\-12 pp. 45. The Paleohiology of C'oclophysis bauri (Cope) from the Upper lnassic ( 1\ pachcan) Whit11kcr quarry, Nc11 Me:-.ico, 11 ith deta lied

analysis of a singlt: quarry block. 2009. by Larry F. Rinehart. Spencer Ci. Lucas. Andre\\ 13. Heckert. Justin A. �pidmann and Mauhe11 D. Ccleskey. 260 pp.

46. The taxonomy and paleobiology of the Late Triassic (Cam1an-Norian: Atlamantan-Apachean) drepanosaurs (Diapsida: Archosauromorpha: Drcpanosauromorpha. 20 I 0. by Sill 10 Rcnc�lo, Justin A. <:;p,clmann. Spencer G. Lucas and Giorgio Tarditi Spagnoli. 8 t pp.

4 7. lchnology or the Upper Triassic (Apachcan) Redonda Formation. cast-central Ne,, Mexico. 20 I 0. by Spencer G. Lucas. Justin A. Spielmann, Hendrik Klein and Allan J Lerner. 75 pp.

48. New Smithian (Early Triassic) ammonoids from Crillendcn Springs. Elko Count). Ne, ada: Implications for taxonomy. biostratigraphy and biogeography, 20 I 0. by James F . Jenks. Arnaud Brayarcl, Thomas Briihwiler and I lugo Bucher . ..\ I pp.

49. Carboniferous-Permian transition in Caiion dcl Cobrc, northern 1\c" Mexico, 20 I 0. edited by Spencer G. Lucas. Jorg W. Schneider and Justin A. Spielmann. 229 pp.

50. Re, icw of the tetrapod ichnofauna of the Moenkopi Formation Group ( l:arly-Middle Triassic) or the American S0t1th1\ est. 20 I 0. b)- 1 lendrik Klein and Spencer G. Lucas. 6 7 pp.

51. Crocodyle tracks and trace�. 20 I 0. edited by Je�pcr Milan, Spencer (i. Luca�. Martin G. Lockley and Jw,tin A. Spielmann. 24-1 pp. 52. Sclachians from the Upper Cretaceous (Santonian) 1 losta Tongue of the Pomt Loo�out ,andswne. central Nc11 Mexico. 2011. by

Jim Bourdon. Keith Wright, Spencer G. Lucas, Justin A. Spielmann and Randy Pence. 54 pp. 53. Fossil Record J. 2011. edited by Robert M. Sullivan. Spencer G. Lucas and Justin A. Spielmann. 736 pp. 54. lchnology of the Mississippian Mauch Chunk Formation, eastern Pennsylvania. 2012. by Da, id L. r illmorc, Spencer G. Luca, and

Edward L. Simpson. 136 pp. 55. letrapod fauna of the Upper Triassic Rcdonda Formation. east -central New Mc,1co: The characteristic a�semhlage ul'the

Apachean land-vcrtehrak faunachron, 2012. by Justin A Spielmann and Spencer G. Lucas. 119 pp. 56. Revision of the Lower Triassic tetrapod ichnol'auna from Wiory. Holy Cross Mountains. Poland. 2012. by llendrik Klem and

Gr1:egorz Nied,:11 iedzl-i, 62 pp. 57. Vcrtl.!hratc Coprolites.2012. edited by /\drian P . I lunt, Jesper Milan, Spencer U. Lucas mid Justin A. Spielmann. 387 pp. 58. A new archaic basking shark (Lamniformcs: Cctorhinidae) from the late C:ocenc ot',\estcm Oregon, U.S.A., and description of the

dentition, gill rakcrs and vertebrae of the recent basking sh,1rk Ce10rh111us maximus (Gunncrus), 2013. by Brncc J. Welton. 4K pp 59. The Carboniferous-Permian transition in central Ne11 Mexico.2013. ed11cd by Spencer G. Lucas. W . John Nelson. William A.

l)iMichelc. Justin/\. Spielmann, Karl Krainer. James f:.. Barrick. Sco11 l:lrick and St'.bastian Voigt. 389 pp. 60. The Carhoniferous-Permiun transition. 2013. edited hy Spencer (1 1 ucas. William A. DiMichcle. James E. Barrick. Joerg W.

Schneider and Justin A. Spielmann. 465 pp. 61. TI1e Triassic System: ew Developments 111 Stratigraph)- and Paleontology, 2013. edited hy Ln,uence 11. Tanner, Justin A.

Spielmann and Spencer U. Lucas, 612 pp. 62. Fossil Footprints of Western North America. 2014. edited by Martin G. Locke) and Spencer G. Lucas. 508 pp. 63. Variation in the Dentition of Coefophysis houri, 2014. by Li�a G. Buckley and Philip J. Currie, 73 pp. 64. Conodonts from the Carnian-Norian Boundary ( Upper Triassic) of Black Bear Ridge. ortheastcrn British Columbia. Canada.

2014. by Michael J. Orchard. 139 pp. 65. Carboniferous-Pern1ian Transition in the Robledo Mountains Southern Ne1� Mexico. 2015. edited by Spencer G. Lucas and

William A. DiMichele. 167 pp. 66. The Marine Fish Fauna of the Middle Pleistocene Port Orford Formation and flk River Beds, ('ape Blanco, Oregon. 2015, by

Bruce J. Welton, 45 pp. 67. Fossil Record 4.2015. edited by Robert M. Sullivan and Spencer G. Lucas, 332 pp. 68. Fossil Vertebrates in New Mexico, '.1015. edited by Spencer G. Lucas and Robert M. Sullivan. -B8 pp. 69. The Pennsylvanian System in the Mud Springs Mol!ntains. Sicm1 County. Ne11 Mexico. USA.2016. by Spencer G. Lucas, Karl

Krainer, James E. Barrick and Daniel Vachard. 58 pp. 70. Eocycforo.1·wir11s appe10/at11.1. a Middle Triassic Amphibian.2016, by Rmehart and Lucas, 118 pp. 71. Cretaceous Period: Biotic Di, ersity and Biogeography. 2016. edited by Khosla and Lucas, 330 pp. 72. Rotten Hill: a Late Triassic Bonebed in the Texas Panhandle. USA. 2016. b} S.G. Lucas. L.F. Rinehart, A.f3. lleckcrt A.P. 1 lunl

and J.A. Spielmann. 97 pp. 73. The Red Corral ( Proctor Ranch) Local Fauna ( Pliocene, Blancan) of Oldlrnm Count). Texas.2016. b) G.E. Schult,:, 74. Fossil Record 5.2016, edited by Robert M. Sullivan and Spencer G. I uca�. 352 pp. 75. New Well Peak, 2017, by Karl Kraincr, Daniel Vachard and Spencer G. Lucas, 163 pp. 76. The Lockatong Formation, 2017. by David L. Fi II more, Michael .I. SzaJna. Spencer U. Lucas. f3rian W. I larthnc and Ech, ard L.

Simpson, 107 pp.