botany lecture - chapter 5 and 8

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    STEMS AND FUNCTIONS

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    provide a framework for leaves, flowers

    and seeds.

    carry water and minerals from the soil, and

    sugars manufactured in leaves throughout

    the plant.

    MAJOR FUNCTIONS OF THE STEM

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    ECONOMIC IMPORTANCE

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    Horticulture ,

    landscaping and bonsai

    making

    History,Culture and

    Economics

    ECONOMIC IMPORTANCE

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    (Curcuma longa)-luyang dilaw

    ECONOMIC IMPORTANCE

    hops (Humulus lupu lus).-

    Beer flavor

    Cinnamon bark

    cinnamon bread

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    Parts of a Stem

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    Parts of a Stem

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    STEM GROWTH AND DIFFERENTIATION

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    STEM GROWH AND DIFFERENTIATION

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    The three sections in young dicot stem are dermal, cortex

    and stele.

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    Cross section of

    young dicot stem

    A young dicot stem has eustele a true stele.

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    Monocot stem shows dermal and ground tissue sections.

    The arrangement of vascular bundles is scattered.

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    Monocot stem

    Vascular bundle in monocot stem is enclosed by bundle

    sheath

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    Dicot stem has eustele

    Monocot stem has dispersed bundles

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    EUSTELE

    With loose ring

    of vascular

    bundles

    From Root Protostele to stem eustele

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    VASCULAR CAMBIUM

    o Initiation of the vascular cambium

    VASCU AR CAMBIUM

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    VASCULAR CAMBIUM

    Fusiform initials

    Ray initials

    Ray initials

    Fusiform initials

    1. Storage parenchyma2. Albuminous cells

    1. tracheids, vessel

    elements and fibers

    2. Sieve cells, sieve tube

    members, companion

    cells, fibers

    S d G th

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    Presence of secondary growth

    Modern ferns Absent in all

    species

    Gymnosperms Present in all

    species

    Dicots Present in manyspecies, other

    species are

    herbs

    Monocots Ordinary type is

    absent in many

    species, somehave anomalous

    secondary

    growth

    Secondary Growth

    VASCULAR CAMBIUM PRODUCES SECONDARY XYLEM

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    VASCULAR CAMBIUM PRODUCES SECONDARY XYLEM

    AND PHLOEM

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    Region of secondary vascular tissues

    Old di d l i

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    Vascular cambium

    Secondary xylem

    Primary xylem

    pith

    Secondary phloem

    Primary phloem

    Old dicot stem secondary vascular tissues

    S d X l T f W d C ll

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    Secondary Xylem Types of Wood Cells

    Axial System from fusiform initials

    Radial System ray initials

    Fibers and vessels in Wood

    SECONDARY XYLEM

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    SECONDARY XYLEM

    1. Types of Wood Cells derived from fusiform initials and ray initials

    fusiform initialsaxial system

    ray initialsradial system

    SECONDARY XYLEM

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    SECONDARY XYLEMGrowth Rings a.k.a. Annual Ring

    1. Early Woodspring wood

    2. late woodsummer wood

    SECONDARY XYLEM

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    Darkening of the Heartwood

    1. Breakage of water columns in

    tracheary elements (freezing, windvibration, etc)

    2. Fungal attack

    3. Formation of tylosis

    4. Metabolic changes that produce

    phenolic compounds, lignins,aromatic compounds

    SECONDARY XYLEM

    HEARTWOOD AND SAPWOOD

    SECONDARY XYLEM

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    SECONDARY XYLEM

    REACTION WOOD

    Angiosperm: Tension Wood

    Upper side of stem produces many

    special gelatinous fibers whose walls

    are rich in cellulose

    Gymnosperm : Compression Wood

    Underside of the branch produces

    fibers whose walls are rich in lignin.

    SECONDARY PHLOEM

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    SECONDARY PHLOEM

    INNER BARK

    OUTER BARK or PERIDERM

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    OUTER BARK or PERIDERM

    Cork and the Cork Cambium

    OUTER BARK or PERIDERM

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    lenticel

    phellemPhellogen or

    Cork cambium

    Phelloderm orCork

    parenchyma

    OUTER BARK or PERIDERM

    Cork and the Cork Cambium

    VARIABILITY OF THE OUTER BARK

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    VARIABILITY OF THE OUTER BARK

    LENTICELS AND OXYGEN DIFFUSION

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    LENTICELS AND OXYGEN DIFFUSION

    Pine stems have resin ducts

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    Pine stems have resin ducts

    ANOMALOUS FORMS OF GROWTH

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    ANOMALOUS FORMS OF GROWTH

    1. Anomalous Secondary Growth in potato

    Formation of cambia

    around the vesselsAdvantage: speed the production of storage

    capacity in storage roots and stems

    ANOMALOUS FORMS OF GROWTH

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    ANOMALOUS FORMS OF GROWTH

    Included phloemtype of secondary phloem located between two bands

    of xylem

    Selective advantage may be protection of phloem from insects by one to

    several layers of wood

    ANOMALOUS FORMS OF GROWTH

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    ANOMALOUS FORMS OF GROWTH

    3. Secondary Growth in Monocots

    Joshua tree (Yucca brevifolia)

    Outermost cells

    of secondary

    vascular bundles

    develop into

    fibers with thick

    secondary walls.

    Modified stems

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    Modified stems

    Onion

    Bulb short stem covered with fleshy blades

    Function storage and reproduction

    Modified stems

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    Cactus

    Cladodes- green stemPhotosynthesis

    Reproduction

    storage

    Rhizome- scaly underground stem

    Reproduction

    storage

    Modified stems

    Modified stems

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    Tubers

    swollen outgrowths of stolon for

    storage and reproduction

    Hooks for protection in stem of rose

    Modified stems

    Modified stems

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    Stolons-horizontal stems with

    adventitious buds

    for reproduction of new plants

    Modified stems

    Modified stems

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    Gabi or taro

    Corm

    vertical and shortunderground stem

    reproduction

    storage

    Twinerclimbing/twining

    stem support

    Modified stems