1) Mystery of “Pheretima koellikeri” distributed only in Kyushu by Kobayashi 1941:Pheretima koellikeri” – Amynthas vesiculata the distribution of Amynthas vesiculata

2) Pheretima koellikeri again Dr. Blakemore's aims on New genus proposal Re-description of "Pheretima koellikeri" (Manus koellikeri) by Blakemore 2010 Comparison Pheretima koellikeri, Manus koellikeri, Metaphire vesiculata

3) Metaphire hilgendorfi / Amynthas tokioensis species complex

4) Slide of addition 10 or more specimens of Metaphire ryukyusensis sent from Dr. Hamada to me

I ask Dr. Blakemore “Blakemore “Pheretima koellikeriPheretima koellikeri””

Lecture date 20x14 /5x /1x ; Beginning day :2014/4/30 The description will be corrected in the future.

Metaphire hilgendorfi / Amynthas tokioensis specise complex in Kobayashi (1941)

沖縄

大島

屋久島

甑島

五島

壱岐

対馬

九州

隠岐

四国

中国

近畿

中部

関東

東北

北海道

朝鮮

済州島

台湾

南シナ

中シナ

北シナ

満州国

樺太

Okinaw

a (Island)

Am

amioshim

a (Island)

Yakusim

a (Island)

Kosikijim

a (Island)

Gotou Fukue (Island)

Iki (Island)

Tusima (Island)

Kyushu

Oki (Island)

Shikoku

Chu-goku

Kinki

Chu-bu

Kantou

Touhoku

Hokkaido

korea

Cheju Island

Taiwan; Form

osa

South Chaina

Mid C

haina

North C

haina

North east part of C

haina (Manchuria)

Saghalien

Disappearing Ph. koellikeri + + + +Ph. agrestis (Goto et Hatai) + + + + + + + + + + + + + +Ph. communissima (G.et.H.) + + + + +Ph. glandularis (G. et. H.) +Ph. gomejimensis Ohfuchi + + +Ph. hilgendorfi (Mihaelsen) + + + + + + + + + + +Ph. irregularis (Goto et Hatai) + + + + + + + + + +Ph. parvicystis (G.et.H.) +Ph . soulensis Kobayashi + + + + + + + +Ph . tappensis (Ohfuchi) + + +Ph. vittata (Goto et Hatai) + + + + + + + +Ph. yunoshimensis Hatai + + + +Ph. yamadai Hatai + + + + +

Metaphire hilgendorfi / Amynthas tokioensis species complex in Kyushu, Sikoku, Chu-goku, Kinki,Chu-bu and its surrouding

Blakemore (03-11)

Pheretima (Parapheretima) koellikeri Michaelsen, 1928 is herein considered synonymous with the prior Metaphire vesiculata (Goto & Hatai, 1899), thereby removing the genus Pheretima sensu stricto from the Japanese list.

Pheretima koellikeri again

dr=fragliche acanthodriline Prostata→Secretory diverticulum duct [fragliche→questionable]

Pr=Pheretima-Prostata→Prostatic duct

Pb=Penial borste →Penial setae

Figure above was quoted from Record of the Australian Museum (2000) Vol. 52: 187-222 Native Earthworms (Oligochaeta) from Southeastern Australia, with the Description of fifteen New Species by R.J. Blakemore

as Stephenson (1930) remarked, normal glandular prostates with the usual relations are also present, thus duplicated or as Stephenson (1930) remarked, normal glandular prostates with the usual relations are also present, thus duplicated or transitional functionality is unlikely.transitional functionality is unlikely. (Blakemore 2013) (Blakemore 2013)

I might have misunderstood the penial setae.

I might have misunderstood the penial setae.

boxed sketch: lhs spermathecae, 18 rhs prostate and manicate caecum of the immature second type specimen

Ventral side (6/7/8) of Metaphire vesiculata

6/ 7/ 8 17/18/19 6/ 7 / 8 /9 16/17/18/19 /20 　　　　 　 24/25 /26/27

Metaphire vesiculata

Pheretima koellikeri Manus koellikeri Metaphire vesiculataOligochaeta. Das Tierreich,10. xxixMichaelsen, W. 1900a.

A New Earthworm Genus(Oligochaeta, Megadrili,Megascolecidae) from JapanRobert J. Blakemore1

Several taxa belonging toMetaphire hilgendorfi/Amynthastokioensis species complex havingfull sexual organ in FukuokaKyushu Japan Y. Sugi

Caeca 5 oder 6 schlanke fingerformige, ///(Manicate) Manicate Manicate

Spermatheca 2 pair in 6/7/8 2 pair in 6/7/8 2 pair in 6/7/8

Diverticulum Michaelsen shows one withzig-zagging diverticular bulb;

in contrast, the current lectotypehad flat, rather compact bulbs onshort only slightly wavy stalks

� iverticular bulb is straight.

　 Other features　 1

An attached gland is not described. There is an attached gland of a

long handle near the base ofeach spermathea.

Spermatheca surrounding

Genital markings absent except forpads around spermathecal pores

pads around spermathecal poreswas signs.

Prostatic duct on 18 on 18

Penial seta (Penial　 borste)

Present (shown in Fig. 1)

attached glands

An attached gland is not described. There are two attached glandswith a long handle near the baseof prostatic duct.

I made an effort to understand the paper of manus by Dr. Blalemore. However, I amuncertain whether to have understood the paper enough.

Further reading solving of "Manus"one a posterior amputee mature, previously dissected with most ofinternal organs removed such as gizzard, one a posterior amputeemature,内臓の大部分がgizzard both lhs spermathecae, and primary maleorgans except for part of 18lhs prostate glandなど取り除かれている状態で、以前に、解剖されるProstates racemose in 17.22 each ducting to penis within copulatory pouch that is confined to bodywall with accessory secretory diverticulum opening anterior to male pore and associated with alarge penial seta (as shown in Michaelsen’s fig. 1). These di

Prostates racemose in 17.22 accessory secretory diverticulum始まりがmale poreに前方であって大きいpenial setaで関連していた状態で体壁に限定されるcopulatory pouchの中の陰茎への各ダクト(Michaelsenの図1で示されるように。) これらのdiverticulaはflattened tubular prostatesに類似して、segments 16 or 17に達する

the current type has large flat pads around the spermathecal pores, possibly these have somecorollary function to the diverticula and penial setae of the male pores.

the current typeがthe spermathecal poresの周りに大きいflat padsを持っているということである、ことによるとこれらはthe male poresのthe diverticulaとpenial setaeに何らかの

Previously thought to resemble parts of the M. hilgendorfi species-group (Blakemore 2003, 2010), it is now clearly separated.以前M. hilgendorfi species-group(Blakemore 2003、2010)の部分に類似するように思われて、それは現在、明確に切り離される。

Previously thought to resemble parts of the M. hilgendorfi species-group (Blakemore 2003, 2010), it is now clearly separated.

Manus by Dr. Blakemore

INTRODUCTION: This paper aims to establish it firmly within bounds of ICZN (http://iczn.org/) stated aims for standard, sense and stability for animal names in science.

A New Earthworm Genus (Oligochaeta, Megadrili, Megascolecidae) from Japan

Robert J. Blakemore1

Abstract Genus Manus is newly proposed with type and only species Pheretima koellikeri

Michaelsen, 1928. Type specimens of the type species preserved in the Zoological Museum Hamburg

are inspected and sketched. Distribution of the species is Japan (Kyushu?) with an unconfirmed Korean

record. The new genus is diagnosed by a secretory diverticulum opening adjacent to each prostate

gland and associated with a penial seta internally and, secondarily, on its manicate intestinal caeca.

Key words: Pheretima, new genus, Asiatic, prostates, conservation status, von Kölliker.

manus1 -ūs, f

1 （人間の）手．

ad manum esse (Liv) 手近にある

per manūs tradere (Caes) 手渡しする

ad meam manum (Cic) 手ずから , 自ら

羅和辞典から

Appendix slide