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Page 1: WETLAND BIRDS - Sympatico · 2006. 11. 7. · Wetland birds of North America : a guide to observation, understanding and conservation / Scott Leslie. Includes bibliographical references

WETLAND BIRDS

of North America

Page 2: WETLAND BIRDS - Sympatico · 2006. 11. 7. · Wetland birds of North America : a guide to observation, understanding and conservation / Scott Leslie. Includes bibliographical references
Page 3: WETLAND BIRDS - Sympatico · 2006. 11. 7. · Wetland birds of North America : a guide to observation, understanding and conservation / Scott Leslie. Includes bibliographical references

WETLAND BIRDS

of North America

A Guide to Observation,

Understanding and Conservation

Scott Leslie

Page 4: WETLAND BIRDS - Sympatico · 2006. 11. 7. · Wetland birds of North America : a guide to observation, understanding and conservation / Scott Leslie. Includes bibliographical references

Copyright © 2006 by Scott Leslie

All rights reserved. No part of this work covered by the copyrights hereon may be reproducedor used in any form or by any means—graphic, electronic or mechanical, including photocopy-ing, recording, taping or information storage and retrieval systems—without the prior writtenpermission of the publisher, or, in case of photocopying or other reprographic copying, a licencefrom Access Copyright, the Canadian Copyright Licensing Agency, One Yonge Street, Suite1900, Toronto, Ontario, M6B 3A9.______________________________________________________________________Library and Archives Canada Cataloguing in Publication

Leslie, Scott, 1963 –Wetland birds of North America : a guide to observation, understanding

and conservation / Scott Leslie.

Includes bibliographical references and index.ISBN 1-55263-722-0

1. Water birds—North America—Identifiication. 2. Bird watching—North America—Guidebooks. I. Title.

QL681.L47 2006 598’.176’097 C2005-906161-8________________________________________________________________________

The publisher gratefully acknowledges the support of the Canada Council for the Arts and theOntario Arts Council for its publishing program. We acknowledge the support of the Governmentof Ontario through the Ontario Media Development Corporation’s Ontario Book Initiative.

We acknowledge the fiinancial support of the Government of Canada through the BookPublishing Industry Development Program (BPIDP) for our publishing activities.

Conservation status maps courtesy of NatureServe. 2004. NatureServe Explorer: An onlineencyclopedia of life [web application]. Version 4.1. NatureServe, Arlington, Virginia. Availableat www.natureserve.org/explorer.

Range maps courtesy of Ridgely, R.S., T.F. Allnutt, T. Brooks, D.K. McNicol, D.W. Mehlman,B.E. Young, and J.R. Zook. 2003. Digital Distribution Maps of the Birds of the WesternHemisphere, version 1.0. NatureServe, Arlington, Virginia, USA.

Key Porter Books LimitedSix Adelaide Street East, Tenth FloorToronto, OntarioCanada M5C 1H6

www.keyporter.com

Text design and formatting: Marijke FriesenPhotographs on pages 11 and 19: Paula Leslie

Printed and bound in China

05 06 07 08 09 6 5 4 3 2 1

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For Paula, whose love, support and wisdom have made so many things possible.

Page 6: WETLAND BIRDS - Sympatico · 2006. 11. 7. · Wetland birds of North America : a guide to observation, understanding and conservation / Scott Leslie. Includes bibliographical references

Foreword . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .9

Wonderful Wetland Birds . . . . . . . . . . . . . . . . . . . . .11Wetlands: A Diversity of Homes for Birds . . . . . . . .17Wetlands: An Intricate Web of Life . . . . . . . . . . . . . .25

Introduction to the Species Accounts . . . . . . . . . . . .35

WATERFOWLCanada Goose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .39Wood Duck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .43Mallard . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .46American Black Duck . . . . . . . . . . . . . . . . . . . . . . . .49Northern Pintail . . . . . . . . . . . . . . . . . . . . . . . . . . . .53Northern Shoveler . . . . . . . . . . . . . . . . . . . . . . . . . . .56Blue-Winged Teal . . . . . . . . . . . . . . . . . . . . . . . . . . .59Green-Winged Teal . . . . . . . . . . . . . . . . . . . . . . . . . .62Ring-Necked Duck . . . . . . . . . . . . . . . . . . . . . . . . . .65Bufflehead . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .69Hooded Merganser . . . . . . . . . . . . . . . . . . . . . . . . . .72

WADING BIRDSAmerican Bittern . . . . . . . . . . . . . . . . . . . . . . . . . . . .77Great Blue Heron . . . . . . . . . . . . . . . . . . . . . . . . . . .80Great Egret . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .84Snowy Egret . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .87Reddish Egret . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .90Little Blue Heron . . . . . . . . . . . . . . . . . . . . . . . . . . . .93Tri-Colored Heron . . . . . . . . . . . . . . . . . . . . . . . . . .96Green Heron . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .99Yellow-crowned Night Heron . . . . . . . . . . . . . . . . .102Glossy Ibis . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .105White Ibis . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .108Roseate Spoonbill . . . . . . . . . . . . . . . . . . . . . . . . . .111Wood Stork . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .114

TABLE OF

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Limpkin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .117Sandhill Crane . . . . . . . . . . . . . . . . . . . . . . . . . . . . .120

BIRDS OF PREYOsprey . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .125Northern Harrier . . . . . . . . . . . . . . . . . . . . . . . . . .129Red-Shouldered Hawk . . . . . . . . . . . . . . . . . . . . . .132Bald Eagle . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .135Peregrine Falcon . . . . . . . . . . . . . . . . . . . . . . . . . . .138Short-Eared Owl . . . . . . . . . . . . . . . . . . . . . . . . . . .141

RAILSClapper Rail . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .147Sora . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .150Common Moorhen . . . . . . . . . . . . . . . . . . . . . . . . .154American Coot . . . . . . . . . . . . . . . . . . . . . . . . . . . .157

SHOREBIRDS, GULLS AND TERNSBlack-Bellied Plover . . . . . . . . . . . . . . . . . . . . . . . . .163Killdeer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .166Semi-palmated Plover . . . . . . . . . . . . . . . . . . . . . . .169Piping Plover . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .172Snowy Plover . . . . . . . . . . . . . . . . . . . . . . . . . . . . .175Wilson’s Plover . . . . . . . . . . . . . . . . . . . . . . . . . . . .178American Oystercatcher . . . . . . . . . . . . . . . . . . . . .181American Avocet . . . . . . . . . . . . . . . . . . . . . . . . . . .184Greater Yellowlegs . . . . . . . . . . . . . . . . . . . . . . . . .187Lesser Yellowlegs . . . . . . . . . . . . . . . . . . . . . . . . . . .190Willet . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .193Pectoral Sandpiper . . . . . . . . . . . . . . . . . . . . . . . . .196Semi-palmated Sandpiper . . . . . . . . . . . . . . . . . . . .199Least Sandpiper . . . . . . . . . . . . . . . . . . . . . . . . . . . .203Short-Billed Dowitcher . . . . . . . . . . . . . . . . . . . . . .206Wilson’s Snipe . . . . . . . . . . . . . . . . . . . . . . . . . . . . .209

CONTENTS

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Ring-Billed Gull . . . . . . . . . . . . . . . . . . . . . . . . . . .212Herring Gull . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .215Caspian Tern . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .218Black Skimmer . . . . . . . . . . . . . . . . . . . . . . . . . . . .221

DIVING BIRDSCommon Loon . . . . . . . . . . . . . . . . . . . . . . . . . . . .227Pied-Billed Grebe . . . . . . . . . . . . . . . . . . . . . . . . . . .231American White Pelican . . . . . . . . . . . . . . . . . . . . .234Brown Pelican . . . . . . . . . . . . . . . . . . . . . . . . . . . . .237Anhinga . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .240Double-Crested Cormorant . . . . . . . . . . . . . . . . . . .243

PERCHING BIRDSBelted Kingfisher . . . . . . . . . . . . . . . . . . . . . . . . . . .249Alder Flycatcher . . . . . . . . . . . . . . . . . . . . . . . . . . .252Tree Swallow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .255Yellow Warbler . . . . . . . . . . . . . . . . . . . . . . . . . . . .259Common Yellowthroat . . . . . . . . . . . . . . . . . . . . . .262Savannah Sparrow . . . . . . . . . . . . . . . . . . . . . . . . .265Song Sparrow . . . . . . . . . . . . . . . . . . . . . . . . . . . . .268Swamp Sparrow . . . . . . . . . . . . . . . . . . . . . . . . . . .271Bobolink . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .274Red-Winged Blackbird . . . . . . . . . . . . . . . . . . . . . .277Common Grackle . . . . . . . . . . . . . . . . . . . . . . . . . .280

Wetland Birds: The Future . . . . . . . . . . . . . . . . . . .283

Selected Bibliography . . . . . . . . . . . . . . . . . . . . . . .289Selected Important Wetlands in Canada . . . . . . . . .292Selected Important Wetlands in the United States . .300Wetland Birds Internet Resources . . . . . . . . . . . . . .306Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . .307Index . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .309

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FOREWORD

Of the Earth’s wild creatures, birds are the ones we have the mostcontact with. Unlike mammals, which tend to be shy and retiring, andare generally nocturnal or active at twilight, most birds are out andabout during daylight hours. As the naturalist David Attenboroughsays, “They are lively; they are lovely; they are everywhere.” Theyare present in our daily lives. The same is true of wetlands.

Many cities and towns that are found along lakeshores, sea-coasts and rivers are adjacent to what remains of natural wetlands.In fact, millions of North Americans pass by wetlands on theirdaily commute, many of them unaware of the stunning wealth oflife right under their noses. Such proximity and visibility is whyboth wetlands and birds are vital reminders of a parallel worldmade up of animals, plants and habitats, completely independentof our human existence.

Despite disappearing from the continent at a rate of about 150acres every hour, gobbled up by urban development, agriculture,roads and the like, many wetlands hang on, providing homes forhundreds of millions of birds and other organisms across NorthAmerica. Some are close at hand and accessible, others are remote.They are truly amazing places to visit and to know; they are therichest ecosystems on the continent, seething with life, brimmingwith beauty and complex beyond comprehension. Our lives areenhanced by the existence of wetlands as they, among other things,supply and purify our drinking water, regulate flooding and bufferagainst coastal erosion.

I hope this book succeeds in illustrating some of the wonderfuldiversity of wetland birds while helping you identify and under-stand them a little better. As well, it is my sincere wish that it conveysthe need to protect their habitats. But I suppose my ultimate aim isto inspire you to visit a local wetland and see for yourself just howfascinating this important part of natural world is.

9

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for many species of rare plants in particular. The high populationof insects attracts insectivorous birds such as swallows and fly-catchers and the abundant frogs attract green herons.

Woodland vernal pools are wetlands that form after rainwateror snowmelt fills pockets on the forest floor. They are anywherefrom a few square yards to pond size. Because they generally lackany fish that would act as predators, insects are able to reproducefreely, creating abundant food for the insectivorous birds that mayuse woodland pools such as the yellow-rumped warbler, thecommon yellowthroat and the alder flycatcher. These birds willoften hunt by “hawking,” where they make short flights from aperch to catch an insect.

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Temperate zone lakes and fens are key habitats for many species of wetlandbirds, but have been hit hard by acid rain, which can result in the depletion of

vital fish populations

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WETLANDS:

AN INTRICATE

WEB OF LIFE

I look with keen anticipation through the gauze of oscillatingmayflies to the cove beyond. It is covered with water lilies andbrimming with life. A beaver dam has blocked the flow of the littlebrook that runs from the cove, flooding the area and transformingit into a marshy, languid pool.

As I emerge from the trees that edge the cove, a startled ring-necked duck patters across the pond before finally taking flight.Several painted turtles sunning themselves on a log plop into thewater. It’s surprising how acute the senses of these little turtlesare—they contradict our usualstereotypes of reptiles. A woodduck hen and her train of youngquietly glide by, unaware of mypresence. Overhead, an osprey ison a keen lookout for fish for itshungry young, while on a tinyexposed bar at the edge of thecove, a loon carefully rearrangesthe eggs on her muddy nest.

Secret lives such as these arelived in this little wetland andthousands of others across NorthAmerica. The best-kept secret ofall, however, is just how complex awetland ecosystem is. To trulyappreciate its intricate nature, an

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Insects, such as these matingdamselflies, are a vital strand in the

complex food web of a wetland

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amphibious approach is needed. So, once I’ve surveyed the worldabove with its birds and damselflies, I don a mask and snorkel andslip under the cove’s surface into the cool, softly lit water beneath.

Magical underwater groves of pond lilies spring from themuddy bottom a few feet below to stretch long, sinuous stemstoward the surface, each topped by a single oval leaf that soaks up

life-giving sunlight.In water tintedbronze by planttannins, the lightslants through thefragile grove, illu-minating schools ofgolden shiners andminnows and alone painted turtleswimming throughthe delicate under-story. In deeperwater, a large snap-ping turtle stirs asmy shadow crossesits prehistoric eye,then nonchalantly

swims away over the muddy bottom. Leeches, undulating ribbonsof black and red flesh, ply the shallows in search of a host. Dragonfliesdab the water above me, creating tiny concentric ripples as theypluck prey from the surface. A dark shape moves through the waterin the distance, perhaps a pied-billed grebe diving for minnows.

In a wetland there really is no boundary separating the sunlitworld of trees, birds and sky from the dim world of pond lilies, fishand mud. All organisms here are members of a food chain thatconnects a minnow about to be snapped up by a hungry perch toan osprey 100 feet overhead, ready to dive and capture that sameperch, thus transforming predator into prey.

Ultimately everything in a wetland, including birds, is depend-ent for survival upon the primary producers. Microscopic algae,

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Frogs are a key component of the freshwater wet-land ecosystem, providing food for wading birds.

Unfortunately, their populations are plummeting forvarious human-caused environmental reasons

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duck weed, pond lilies, reeds, cattails and countless other species ofplants combine sunlight with the incredibly rich supply of nutrientsfound in the water and soil to create the most productive habitat inNorth America. Birds may be some of the most “charismatic” (touse a term popular with biologists to describe highly visible species)residents of wetlands, but they are also parts of the ecosystem,playing a role no more or no less vital than every other organismthat lives there.

Evolution has crafted a jigsaw puzzle of enormous complexityin this environment, where birds fit perfectly in virtually everyniche that is available to them. Each niche of a given bird is madeup of layer upon layer of trophic (feeding) relationships populatedby lower organisms that go all the way down to the most basic levelof the algaes. To illustrate thispoint, consider the particularchain of events that must occurbefore a tree swallow can get ameal for its hungry chicks. Thelate spring sun warms the shallow,nutrient-rich water. Microscopicaquatic algae drink up the sunlightand bloom into profusion, tinge-ing the water a light green andproviding food for newly hatchedinsects such as mosquito larvae.Some time later, millions of theselittle wrigglers eventually developinto clouds of mosquitoes, whichin turn become prey for elegantcivil-bluet damselflies that pluckthem from the air. Gracefullyswooping over the pond, with itsmouth gaping wide, a tree swallowscoops hapless damselflies on thewing, then returns to feed them tohungry chicks in the nest. Andthis extremely simplified example

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Muskrats, the most common andwidespread mammals inhabiting

North America’s wetlands, areimportant to the health of the

ecosystem as they collectively con-sume enormous amounts of plantmatter, which are ultimately recy-

cled back into the environment

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represents just a fraction of the actual complexity of trophic rela-tionships involved in sustaining just one bird in a wetland!

In reality, there are probably as many kinds of places to findfood in a wetland as there are species that inhabit it. Roughlyspeaking, a wetland can be divided into six zones within whichfood resources are found: the water column where fish, leeches andaquatic insects live; the bottom where mollusks, insects, larvae andcrustaceans live; the submerged plants that produce stems and onwhich amphibians, insects and mollusks cling; the surface whereduckweed and algae grow; exposed plants that bear fruits, leavesand seeds; and the air where insects and birds fly. Each species ofbird in a wetland will exploit one or more of these places accord-ing to how it has evolved—a bittern hunting frogs in the shallows,a teal gleaning duckweed from the surface, a grebe fishing forleeches in deeper water, for example. The interconnections betweeneverything that lives, even in a small wetland pond, are incompre-hensibly vast. But thanks to millions of years of adaptation throughnatural selection, the system works!

In addition to all of the biological interactions between birdsand other species, there are complex relationships and balances

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Most wetlands are immensely productive ecosystems, and contain as muchbiomass below the water as above

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between birds and thephysical environmentof the wetland itself.The influence of waterlevels, water tempera-ture and pH, weather,the amount of coverand shelter availableand the presence ofpredators all affect thelives of wetland birds.

For birds the mostimportant physical feature of any wetland is the availability ofwater. Water cover might be permanent or seasonal (ponds,marshes, fens, bogs), during high tide (tidal marshes) or after aheavy rainfall (vernal pools, playa lakes). Even when the surfacewater disappears for a time, the soil may be saturated enough tosupport the aquatic plants, such as cattails, reeds and rushes, thatmany birds rely on for shelter and as cover.

Water conditions that may be sufficient for some birds mightnot be for others. For example, sora rails are able to survive in verysmall patches of extremely shallow or dried-out habitats (as well asother wetter and larger wetlands) that are just damp enough tosupport cattails and other moisture-loving plants. In such an envi-ronment they have a source of food (seeds), nest-building materialsand enough cover from predators. In contrast, the pied-billedgrebe, another small bird, can’t survive in such places because itneeds a relatively large flooded area to land and take off from anddeeper water to dive in for its prey. An osprey requires deep waterthat it can safely dive into from a great height, while a dowitcherneeds water shallow enough to wade through while it feeds byprobing the mud with its long bill. Each species has a particular setof conditions (in this case water depth) that must be met before itcan successfully inhabit a wetland.

Fortunately, most wetlands aren’t exclusively moist or flooded,deep or shallow, but are a combination of conditions that are suit-able for a variety of birds. For example, in salt marshes the water

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The great majority of turtles worldwide dependon freshwater wetlands as habitat

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level changes daily, flooding on the incoming tide and draining on theebb tide. Shorebirds will feed on mudflats for crustaceans and mol-lusks during low tide, while other species like cormorants or greatblue herons will often wait for high tide before they begin to fish.

Other factors also determine how many bird species will be foundin a wetland. Larger wetlands tend to be more heterogeneous witha wider range of water depths and a more varied distribution ofvegetation. This can result in a “patchiness” that provides a greaterchoice of potential microhabitats resulting in a higher diversity ofspecies. The species of birds found in a wetland will also depend onthe kinds of plants present.

Water temperature has an impact on how abundant food willbe for birds. Lower temperatures usually mean a slower growthrate for the primary producers in the wetland—the algae, duck-weeds and other plants. If the water temperature is too low, algaewon’t grow as quickly, and as one of the foundations upon whicheverything else depends, the entire food chain right up to the topwill be affected. In other words, it might be a little harder for thattree swallow to feed her young if the water is just a little colder!Warmer water temperatures, of course, will have the oppositeeffect, generally increasing the mass of vegetation available and thenumber of birds present.

It should come as no surprise that wetlands, with such a highconcentration of birds, attract large numbers of predators. Like amoat around a castle, water is a buffer against incursions by manypredators, preventing their access to eggs or young in the nest. Butsome are undeterred by water and prey heavily on birds, eggs andyoung. Raccoons, skunks, minks, snapping turtles, pickerel, large-mouth bass, snakes and alligators all take their toll, especially onthe eggs and chicks. During particularly dry breeding seasons,water levels decrease and greater access to previously unreachableareas may open up, allowing predators to take more birds.Breeding success may also be lower for birds that nest near outeredges of a wetland, where they are more easily reached.Fortunately, the heavy vegetation that is a feature of the breedingareas of most wetlands provides a way for most nesting birds toconceal their nests effectively, enabling breeding success.

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In a wetland, or any complex ecosystem for that matter, every-thing is more or less dependent on everything else. Tugging at justone strand in this web of biological and physical interactions cancause one species or a group of them to be less successful andanother to become more so. For example, in the case of globalwarming, in which hotter, drier weather results in lower waterlevels in wetlands, pied-billed grebepopulations may decline significantlywhile soras may survive with rela-tively few difficulties, at least for atime. Eventually global warmingwill affect everything in our wet-lands in some way, possibly resultingin catastrophic changes.

Different species of birds utilizewetlands in different ways through-out the year. Wetland-dependentbirds, including many ducks,grebes, rails, herons, loons, cranes,cormorants, shorebirds, some song-birds and a few birds of prey, relyalmost exclusively on this environ-ment for nesting, food and shelterduring the breeding season. They arealso highly dependent on them duringmigration and for overwintering.

Many other birds rely on wet-lands for at least part of the year forvital activities such as feeding,drinking water or shelter from badweather. For example, yellow war-blers, song sparrows, bobolinks andcommon grackles, among others,while not completely dependent onthem, nevertheless are commonly found about wetlands, tappingthem for food and nesting materials. To varying degrees certainraptors also depend on wetland areas as a source of prey or, in

31

These pond lilies, seen under-water, are part of an aquatic

ecosystem that is integral to thehealth of the whole habitat.

There’s often more than meetsthe eye in a wetland environment

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the case of some, such as red-shouldered hawks, as a place tonest near.

All told, the number of bird species that require wetlands fortheir survival to some degree is estimated between one third andhalf of the roughly 650 species that regularly breed in NorthAmerica, north of Mexico. The importance of protecting these vitalhabitats becomes clear when you consider that nearly half of ourbreeding birds, as well as countless other species, depend on a pre-cious 700,000 square miles or 10 percent of the land area that isoccupied by wetlands (not including Alaska). In fact, wetlands con-tain the greatest number of endangered, threatened and vulnerablespecies of any habitat in North America. In the United Statesapproximately one half of all officially endangered species aredependent on wetlands, including seventeen species of birds. Asimilar situation exists in Canada.

The high proportion of endangered, threatened or vulnerablewildlife found in wetlands can be traced to two related facts. Theyare home to an inordinately large number of species in relation totheir land area when compared with other North American habi-tats, and a greater percentage of wetlands have already been lost

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A tree swallow completes the food cycle in a wetland by feeding its youngsome damselflies

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(about half) compared to many other habitat types. Indeed, of theeight species of birds that have become extinct in North America inhistorical times (as recently as 1987 for the dusky seaside sparrow),five of them were birds that relied on wetlands. While variouscauses may have contributed to the extinction of the Eskimocurlew, Carolina parakeet, Bachman’s warbler, dusky seaside sparrowand the Labrador duck (little is known of this bird’s life history, butthat it utilized tidal wetlands to some degree isn’t far-fetched), theloss of a wetland habitat likely played at least some part in all of them.

Today, a real threat of extinction hangs over the heads of themajestic whooping crane, the Mississippi sandhill crane, the woodstork, the Everglades snail kite and the piping plover, to name justa few. All are officially endangered in North America and the lossof their wetland habitat is one of the primary reasons that they maydisappear from this world forever. As well, geographical popula-tions of many other species are in peril of extinction within limitedranges (a phenomenon called extirpation) in North America.

Despite being the most biologically productive ecosystems inNorth America, and providing a home to perhaps hundreds ofmillions of its avian residents, wetlands are being destroyed at afrightening rate. It is estimated that between 1780 and 1980 theconterminous U.S. lost over half or nearly 100 million acres of itsoriginal wetland habitats (most of Alaska’s wetlands are still intact)and continues to lose about 110,000 acres annually. In Canada thenumbers are no more encouraging. Estimates put the historical lossdue to agricultural expansion alone at over 49 million acres. Thelosses continue. Over the past twenty years the world’s focus hasbeen on the destruction of tropical rain forests. North America’swetland ecosystems are like our tropical rain forests: They are thebiological engines that drive avian (and much other) life on thiscontinent. It is time we regard them in proportion to their importance.

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CANADA GOOSE Branta Canadensis

When most North Americans think of geese, they think of Canadageese. As the most common species of goose on the continent, it isfound practically everywhere.

APPEARANCELength 25–45 inches. Wingspan 43–68 inches. A large, heavygoose, gray-brown overall, with lighter undersides, a long blackneck, black head, tail and bill and a striking white cheek patch.Great size differences exist among the various races of the species.The smallest, an Arctic species, is the size of a large duck, while thelargest race may have nearly twice the wingspan and weigh almostthree times as much.

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Canada goose lands on a marsh early in spring

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HABITATIts habitats are ecologically diverse from vast, treeless areas in theArctic to wetlands surrounded by heavy forests to prairie sloughs

and coastal areas. Lakes, ponds,bogs, fens, large streams, urbanponds, wet meadows, swamps,freshwater and saltwater marshesare all utilized by the Canadagoose. Grain and cornfields arealso favored as a convenientsource of food.

BEHAVIORThe most famous behavior of the

Canada goose is their habit of migrating in graceful V-formationsaccompanied by melancholic honking. It is very gregarious, and isfound in flocks. It forages by reaching into the water to gatherfood, and also picks food directly from the surface or from theground. When a flock is feeding, at least one bird will act as a sen-try to warn the others of danger. It is well adapted for walking andswimming, and runs a short distance to gather speed before takingoff. Flight is powerful.

CALLSCall is a resonant double honk that sounds quite pleasant in achorus and at a distance. The smallest race has a higher-pitchedcackling sound.

FOODDiet consists mostly of vegetable matter such as the seeds of grassesand sedges, shoots of aquatic plants, berries, grains and corn. Alsoeats crustaceans, mollusks and other invertebrates.

FAMILY LIFEFemale builds a down-lined ground nest made of grasses, sticks andother vegetation, often in a high spot near water or on a small isletin a marsh or pond. Rarely an abandoned tree nest of an osprey

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The range of theCanada goose

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may be used. Four to ten eggs are incubated for 25 to 30 days bythe female. First flight occurs between 40 to 73 days, depending onthe race. The young are tended by both parents and stay with themthrough their first winter until they return to the breeding groundsin the spring. One brood per year.

MIGRATIONIts perennial V-flight pilgrimage to the North in early spring andback again in late fall has made the Canada goose the archetype ofmigrating birds in North America. Spring birds arrive as the icebreaks up, generally from February to April. Fall migration gener-ally occurs from September to December. In the southern part of itsrange, it is a year-round resident.

CONSERVATION CONCERNSSpecies status overall in North America is secure and stable. TheCanada goose is abundant and widespread across the continentdespite being heavily hunted as a game bird. The Atlantic Canadianpopulation appears to be declining. Although the Aleutian Canadagoose of Alaska, subspecies Leucopareia, is listed as a federallythreatened species in the United States, its population has increased

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The familiar Canada goose is the quintessential wetland bird of North America

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from only 800 in the 1960s to approximately 60,000 birds now.Another subspecies Occidentalis, the dusky Canada goose, is con-sidered in peril of extinction in British Columbia, Oregon andAlaska.

RELATED SPECIESOnly the Brant, a somewhat similar and smaller goose that breedsin the high Arctic, shares the genus Branta with the Canada gooseas a breeding bird in North America.

Did You Know?

The familiar V-formation of Canada geese and other species suchas cormorants has evolved to help them save energy while inflight. In geese, the “V” puts them in a configuration where the aircurrents produced by the wings of a leading bird reduces theamount of drag behind and off to the sides, making flight easierfor the flanking birds that follow. This may reduce energyexpended by up to 50 percent. Obviously, the leading bird doesn’tbenefit and that’s why the position is rotated regularly so all birdscan take advantage.

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WOOD DUCKAix sponsa

In full breeding plumage, the male wood duck is the most beauti-ful of all North American ducks.

APPEARANCELength 18.5 inches. Wingspan 30 inches. The male is unlike anyother North American duck. Its most prominent feature is the long,swept-back crest, which looks like an oversized helmet, and adefined U-shaped spur of white on the side of the head. The rest ofthe bird is an assortment of bright iridescent greens, blues and pur-ples. The similarly shaped female is much duller than the male withan overall variegated brown accented by white stripes and iridescentblue patches. The head is gray with a prominent white eye-ring.Both sexes have long tails and short bills.

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A female wood duck and her ducklings

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HABITATA tree-loving duck, it preferswooded wetlands such asswamps, ponds, lakes, streamsand flooded forest.

BEHAVIORIt is usually seen in pairs, lessfrequently in small flocks. It for-ages by gleaning food from the

surface of the water. Very adept at walking on land, the wood duckoften comes right out of the water to pick the seeds and acorns ofshrubs, trees and other vegetation, and perches in trees. It takes offalmost vertically from water with no running and flight is very rapid.

CALLS Quite vocal. The male utters an upward pitching whistle, while thefemale gives a shrill oo-weet, oo-weet call when alarmed.

FOODDiet includes seeds, acorns, berries, beechnuts and wild rice. Alsotakes aquatic insects, mollusks and amphibians.

FAMILY LIFENest is in a natural cavity or rotted-out branch of a tree.Occasionally nests in a cavity made by another species such as apileated woodpecker. Will also nest in artificial nesting boxes. Thefemale lines it with wood chips and down before laying ten to fifteeneggs, which she incubates for 28 to 37 days. Despite sometimesbeing placed 50 feet or higher, the newly hatched downy young willexit the nest by jumping to the ground unharmed. The mothertends the young for 56 to 70 days until their first flight. One or twobroods per year.

MIGRATIONMigrant birds generally arrive on their breeding grounds inMarch or April. Fall migration occurs from mid-September to

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The range of thewood duck

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November. In the southern part of its range the wood duck is ayear-round resident.

CONSERVATION CONCERNSSpecies status overall in North America is secure and its populationis increasing. Hunted almost to extinction by the late nineteenthand early twentieth century. Fortunately, since the inception of theCanada–United States Migratory Bird Convention of 1916, thepopulation has increased dramatically. Threats still face the species,especially the destruction of the wet forested habitats where it nests.

RELATED SPECIESAlthough considered a “dabbling duck” along with ducks of thegenus Anas, the wood duck belongs to its own separate genus Aix.

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The male wood duckis the most spectacularNorth American wet-

land bird

A female wood duckshows just a hint ofthe male’s brilliant

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