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TMAP Typology of Coastal Vegetation in the Wadden Sea Area

WADDEN SEA ECOSYSTEM No. 32 - 2014

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TMAP-Typology of Coastal Vegetation in the Wadden Sea Area


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PublishersCommon Wadden Sea Secretariat (CWSS), Wilhelmshaven, Germany;Trilateral Salt Marsh and Dunes Expert Group

AuthorsJörg Petersen nature-consult, Hackelbrink 21, D-31139 Hildesheim, [email protected] Kers Rijkswaterstaat, Central Information Services (RWS-CIS). Ministry of Infrastructure & Environment, P.O. Box 5023, NL-2600 GA Delft, [email protected] Stock Landesbetrieb für Küstenschutz, Nationalpark und Meeresschutz Schleswig-

Holstein, Schloßgarten 1, D-25832 Tönning, [email protected]

Cover photosMartin Stock

Lay-outGerold Lüerßen

Source:TMAP-Typology of Coastal Vegetation in the Wadden Sea Area. Version 1.0, 2014.www.waddensea-secretariat.org, Wilhelmshaven Germany.Possible updates can be downloaded at www. waddensea-secretariat.org/saltmarsh

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WADDEN SEA ECOSYSTEM No. 32


Jörg PetersenBas Kers

Martin Stock

2014Common Wadden Sea Secretariat

Trilateral Salt Marsh and Dunes Expert Group

WITH COMMENTS OF THE TMAP EXPERT GROUP SALT MARSH & DUNES: Jürn Bunje, Kees Dijkema, Olaf von Drachenfels, Willem van Duin, Marinus van

der Ende, Peter Esselink, John Frikke, Norbert Hecker, Ulrich Hellwig, Kai Jensen, Peter Körber, Evert-Jan Lammerts, Piet Schipper, Madelein Vreeken

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Contents

1 Introduction 5

2 Basic Information 7

3 Standard TMAP Types of Dunes 9

4 Standard TMAP Types of Dune Slacks 27

5 Standard TMAP Types of Salt Marshes 44

6 Overview of the complete TMAP Typology 71

7 References 86

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1 Introduction

Coastal landscapes and their typical ecosystems like salt marshes and sand dunes are highly dynamic, characterized by steep environmental gradients and controlled by geomorphological, physical and biological processes. In addition, both direct and indirect human impacts are im-portant factors that give shape to the spatial and temporal variation of coastal ecosystems. The conservation of these coastal ecosystems is a major aim of the Wadden Sea Plan (management plan of the Trilateral Cooperation). As a basis for conservation management, sound knowledge about the spatial distribution of coastal ecosys-tems and their temporal change is required. In the past, multiple regional vegetation or biotope typologies have been used to describe and map plant communities of coastal ecosystems in the Wadden Sea region (e.g. Westhoff & den Held 1969; Dijkema & Wolff 1983; Dierssen 1988; Preising et al. 1990; Schaminée et al. 1995, 1996, 1998; Petersen 2000, 2011; Petersen & Pott 2005; Drachenfels v. 2011). In addition, several regional or national classifications for mapping purposes have been developed (SALT-Code, Rijkswater-staat, s. Jong de et al. 1998; Staatsbosbeheer-Code, s. Schipper 2002; Ringot-Code, s. Ringot 1992; SH-Code, LKN, s. Stock et al. 2005). For a trilateral assessment, however, these different regional approaches have been shown to be dif-ficult to compare at first sight.

The Trilateral Monitoring and Assessment Pro-gramme (TMAP; 1997) aimed to harmonize these different approaches for the Wadden Sea region as a basis for improving conservation manage-ment and to reach the conservation targets listed in the Wadden Sea Plan (see CWSS 2010). Since 2000, the TMAP salt marsh expert group developed and discussed a common commu-nity typology for the Wadden Sea saltmarshes. This TMAP salt marsh typology, which the ex-pert group endorsed in 2003, encompassed 29 different types or plant communities from the pioneer, low and high salt marshes and from brackish marshes, including reed beds (Bakker et al. 2005, Esselink et al. 2009). It is a simplified aggregation of vegetation types which is based on the presence of characteristic or dominant plant species. The TMAP salt-marsh typology has been widely used for monitoring the extent and composition of this ecosystem in both the Neth-erlands and in Germany for more than 10 years. In addition, a complementary TMAP dune typol-ogy was developed, in which 68 types of dunes and dune slacks are differentiated (Petersen & Lammerts 2005, Lammerts et al. 2009). However, a common TMAP synopsis of coastal ecosystems

that covers both salt marshes and dunes of the Wadden Sea region was lacking so far.

The synopsis presented here aims to give a comprehensive overview about the existing salt marsh and dune vegetation types in the Wad-den Sea region. Further, a ‘translation’ of these TMAP types to the typologies used in the differ-ent regions of the Wadden Sea is given. Last but not least, an assignment of the presented TMAP types to the habitat types protected by the Eu-ropean Fauna-Flora-Habitat directive (European Commission 2007, Drachenfels v. 2012) is also included in the synopsis.

The synopsis consists of two parts: In the first part, the most important TMAP types are pre-sented on separate and similarly arranged pages. Here, information on typical plant species and on plant communities, which belong to this type, is included. Further, the use of the type in map-ping programs in the Wadden Sea regions and the assignment to regional typologies are listed. Finally, a picture gives a visual impression of the described TMAP type. In the second part of the synopsis, a complete overview about all TMAP salt marsh and dune types of the Wadden Sea region is presented in a table. We, nevertheless, would like to emphasize that this synopsis is not thought to be used as a field key, allthough it has been successfully practised. Regionally field keys are available for this purpose.

We hope that the synopsis will be widely dis-tributed and repeatedly used by nature conserva-tion in all countries and regions of the Wadden Sea. We are sure that this synopsis will contrib-ute to the harmonization of different typologies of coastal habitats, and that it assists us in the process towards a better understanding of the spatial and temporal variation of coastal habi-tats in the Wadden Sea region.

Finally, the synopsis has been developed to improve conservation management of coastal habitats in the Wadden Sea region and thus to contribute to the long-term preservation of the Wadden Sea Natural Heritage.

Kai Jensen, Hamburg University, chair of the TMAP expert group

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2 Basic Information

Column header in tableTMAP Code and TypeCode and description of the TMAP types for dunes, dune slacks and salt marshes.(Bakker et al. 2005 and Petersen & Lammerts 2005).

Standard TMAP TypeDescription of the minimum standard/level of TMAP typology applied in each region. Since Denmark only maps habitat types the assign-ment to the standard TMAP typology constitutes an exception to the rule.

Typical / characteristic speciesDescriptiom of the TMAP type; species written in bold are the important representative taxa; +/- means: not always all, but some of the follow-ing species indicate the TMAP type (Garve 2004, Hauck & de Bruyn 2010, Koperski 2011).

Typical / characteristic vegetationRepresentative plant communities of the TMAP types (Westhoff & Held den 1969, Dierssen 1988, Preising et al. 1990, Pott 1995, Schaminée et al. 1995, 1996, 1998, BFN 2000, Petersen 2000, Petersen & Pott 2005).

Applied in country / federal stateStandard TMAP (in chapter 4-6 only Standard TMAP types are displayed) Netherlands (NL),Lower Saxony (LS), Hamburg (HH), Schleswig-Holstein (SH),Denmark (DK).

Values in the columns“X” = mapped/applied in this Wadden Sea region, “-“ = not mapped in this Wadden Sea region, “N” = no occurrence in this Wadden Sea region.

Reference to other typologiesNatura 2000 Code: European Commission (2007).Types of Annex 1 of Habitat Directive; in addition to the following tables the TMAP-types of salt marshes may also be part of 1130 (in estuaries).

Mapping Code, NL: Bas Kers (Rijkswaterstaat, NL),

RWS SALT 2008 Code, NL: Bas Kers (Rijkswa-terstaat, NL),

Staatsbosbeheer Code, NL: Piet Schipper, Evert Jan Lammerts (Staatsbosbeheer, NL),

Biotope Code, LS: Olaf von Drachenfels (Nds. Landesbetrieb für Wasserwirtschaft, Küsten- und Naturschutz, GER),

Biotope Code, HH: Peter Körber (Nationalpark Hamburgisches Wattenmeer, GER),

Mapping Code SH: Martin Stock (Landes-betrieb für Küstenschutz, Nationalpark und Meeresschutz Schleswig-Holstein, GER),

Biotope Code, SH: Martin Stock (Landesbetrieb für Küstenschutz; Marinus van der Ende (Landes-amt für Landwirtschaft, Umwelt und ländliche Räume des Landes Schleswig-Holstein, GER),

Mapping Code, DK identical with Natura 2000 Code: John Frikke (Miljøministeriet Naturstyrel-sen, DK).

PhotographsJörg Petersen (X.2.2, X.5, X.6.2, X.7.1, X.7.2, X.7.3, X.7.4, X.8.0, X.8.1, X.8.2, X.8.3, X.10, H.0, H.1.1, H.1.2, H.1.3, H.1.4, H.2.1, H.2.2, H.3.1, H.4.1, H.5.1, H.5.2, H.6.1, H.6.2, H.8, H.9, S.1.2, S.2.1, S.2.3, S.3.2, S.3.6, S.3.8, S.3.10, S.3.12, S.3.14, S.5.1, S.5.3),

Martin Stock (X.2.1, X.3.1, X.4.1, X.5, X.6.1, X.12, S.0.0, S.0.1, S.0.2, S.0.3, S.1.1, S.2.2, S.2.4, S.3.1, S.3.7, S.3.12, S.5.1, S.5.3),

Bas Kers (S.2.3, S.3.3, S.3.4, S.3.5, S.3.9, S.3.10, S.3.11, S.3.13, S.5.2, S.6.0, S.6.1),

Uwe de Bruyn (X.8.0, X.8.3).

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3 Standard TMAP Types of Dunes

TMAPCode:

TMAP Type:Description: Typical / characteristic -species -vegetation

X.2.1 Cakile maritima type

Cakile maritimaand +/- Salsola kali, Honckenya pep-loides, Atriplex species, Elymus farctus ssp. boreoatlanticus

Cakiletum maritimae

Applied in the country/federal state:

Standard TMAP

Netherlands Lower Saxony Hamburg Schleswig-Holstein

Denmark

X X X X X -Reference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL

RWS SALT 2008 Code NL

Staats-bosbe-heer Code NL

Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1210(in LS 2110)

XEd Dxc, Dxh, Deu, Dxs

22-a, 22B1

(KDV) KSV KSa CM 2110

Notes: Annual drift lines are short-lived and thus unstable stages during dune development (xeroserie). The vegetation is often of narrow-linear shape, developing on alluvial organic matter deposited during high water levels. A characteristic species in this environment is the Sea Rocket Cakile maritima (see picture).

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TMAPCode:


X.2.2 Crambe maritima type

Crambe maritimaand +/- Honckenya peploides, Silene vulgaris ssp. maritima, Glaucium flavum, Eryngium maritimum, Rumex crispus, Leymus arenarius

Crithmo-Crambetum, Honkenya peploides and Silene vulgaris ssp. maritima veg.


Standard TMAP


Denmark

X N N N X -Reference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1220 - - - - KSV KSv CR -

Notes: Perennial drift-line vegetation at beaches with gravel and boulders, which usually consists of perennial, salt-tolerant and nitrophilous species. Typical species are e.g. Sea Sandwort (Honckenya peploides), Sea-Holly (Eryngium maritimum; see picture) and Yellow Horned-Poppy (Glaucium flavum; see small picture).

Standard TMAP Types of Dunes

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TMAPCode:


X.3.1 Elymus farctus type Elymus farctus ssp. boreoatlanticus (dominant) and +/- Leymus arenarius, Ammo-phila arenaria, Honckenya peploides, Salsola kali, Cakila maritima

Elymo-Agropyretum, Honkenyo-Agropyretum juncei


Standard TMAP


Denmark

X X X X X XReference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2110 Xef Def, Deg, Dd

22A1, 22B1, 22B-a, 22-a, 23A1, 23B-a

KDV KDV KDv EG 2110

Notes: Embryonic dunes are usually only a few decimetres high and their vegetation is often formed solely by Sand Coach (Elymus farctus ssp. boreoatlanticus; see picture). Embryonic dunes represent the earliest stage of dune development (xeroserie).


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TMAPCode:


X.4.1 Ammophila arenaria type

Ammophila arenaria, x Calammophi-la baltica, Leymus arenarius (at least one species dominant)and +/- Sonchus arvense, Oenothera parviflora var. ammophila, Hieracium umbellatum, Calystegia soldanella (rare), Eryngium maritimum (rare), Lathyrus maritimus (rare), Festuca rubra ssp. arenaria, Carex arenaria, Honckenya peploides, Elymus farctus ssp. boreoatlanticus, Cakile maritima

Elymo-Ammophiletum, x Calammophila baltica-veg., Potentillo-Leymetum arenariae (Potentillo-Elymetum)


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2120 - Dd 23B1, 23B-a, 23-a

KDW KDW KDw EM, E 2120

Notes: White dunes are more stable than drift lines and embryonic dunes. They can form the high-est dunes in coastal areas, for which a crucial factor is constant sand supply. If sand supply is reduced or stopped, dune vegetation will develop into later successional stages. Vegeta-tion of white dunes is often patchy without a closed canopy. Typical species are European Marram Grass (Ammophila arenaria; see picture) and Lyme Grass (Leymus arenarius). Spe-cies richness of the vegetation is increasing in later successional stages.


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TMAPCode:


X.5 Dune grassland species of X.5: Carex arenaria, Agrostis capillaris, Festuca rubra ssp. arenaria, Aira praecox, Anthox-anthum odoratum, Luzula campes-tris, Viola canina, Viola tricolor (ssp. curtisii), Jasione montana, Hiera-cium umbellatum, Rumex aceto-sella, Hypochaeris radicata, Galium mollugo, Cerastium semidecan-drum, Lotus corniculatus, Hypnum cupressiforme, Dicranum scoparium, Polytrichum juniperinum, Campylo-pus introflexus, Ceratodon pu-pureus, Cetraria aculeata, Cladonia ciliata, C. portentosa, C. furcata, C. subulata, C. grayi, C. cervicornis, C. ramulosa and see: X.5.1 – X.5.5

Koelerio-Corynepho-retea-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2130 - Dd 14-c, 14-d, 14-g, 14-h, 14-l,m

KDG KDG KDg TR 2130


X.5.1

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Notes: Dune grasslands are characterized by mostly small-growing species. The canopy of the vegetation is often almost closed, organic material is slowly accumulating in the soil and sand supply occurs only occasionally. In this stage of dune development, cryptogams be-come established for the first time. The various sub-types clearly differ (see pictures: X.5.1, X.5.2 and X.5.4) and commonly co-occur within small areas.


X.5.2

X.5.4

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TMAPCode:


X.6.1 Empetrumnigrum type

Empetrum nigrum (dominant)and +/- Polypodium vulgare, Calluna vulgaris, Hieracium umbellatum, Salix repens agg., Lonicera periclymenum, Carex arenaria, Agrostis capillaris, Anthoxanthum odoratum, Luzula campestris, Festuca ovina, Dantho-nia decumbens, Dryopteris dila-tata, Ammophila arenaria, Hypnum jutlandicum / cupressiforme, Pleuro-zium schreberi, Dicranum scoparium, Pseudoscleropodium purum, Hylocom-nium splendens, Cladonia portentosa, C. furcata

Hieracio-Empetretum, Polypodio-Empetretum


Standard TMAP


Denmark

X X X N X XReference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2140 - Dd 20A3, 20-a, 11A3, 11-b, 11/a

KDE - KDe HE 2140

Notes: Black Crowberry (Empetrum nigrum, see picture) is the dominant species of this type, in which Common Polypody (Polypodium vulgare, see picture) does regularly occur too. Or-ganic material is accumulating in the soil at higher rates than in dune grasslands. Low soil pH values due to decalcification are characteristic.


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TMAPCode:


X.6.2 Calluna vulgaris type

Calluna vulgaris (dominant)and + (nearly) no Empetrum nigrumand +/- Salix repens agg., Carex arenaria, Agrostis capillaris, Anthox-anthum odoratum, Luzula campestris, Festuca ovina, Danthonia decumbens, Hieracium umbellatum, Hypnum jutlandicum / cupressiforme, Pleuro-zium schreberi, Dicranum scoparium, Pseudoscleropodium purum

Hieracio-Empetretum - dom. Calluna vulgaris


Standard TMAP


Denmark

X X X N X -Reference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2150 - Dd 20A1 KDC - KDc CV -

Notes: This dune type is characterized by the dominantly occurring Common Heather (Calluna vul-garis, see picture), whereas Crowberry (Empetrum nigrum) is mostly lacking. Accumulation of organic material in the soil and low soil pH values are characteristic for this dune type.


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TMAPCode:


X.7.1 Hippophaë rhamnoides type

Hippophaë rhamnoides (dominant) and +/- Salix repens (ssp. dunensis), Sambucus nigra, Carex arenaria, Ammophila arenaria, Galium mollugo, Festuca rubra agg., Poa pratensis, Li-naria vulgaris, Rubus caesius, Senecio jacobaea

Hippophao-Sambuce-tum nigrae, Salici are-nariae-Hippophaetum


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2160 - Dd 37B1, 37B2, 37B3, 37B-c, 37B-d, 37B-e

KGS - KHh HR 2160

Notes: This shrub vegetation usually occurs on the still calcareous transition zone between white and gray dunes. A characteristic species is the Common Sea-Buckthorn (Hippophaë rham-noides, see picture). Elderberry (Sambucus nigra) is characteristic for later stages of this type.


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TMAPCode:


X.7.2 Salix repens agg. type

Salix repens (ssp. dunensis) (dominant) and +/- Rosa spinosissima, Pyrola rotundifoliaand +/- Carex arenaria, Ammophila arenaria, Galium mollugo, Festuca rubra agg., Poa pratensis

dry Salix repens agg.-veg., Pyrolo-Salicetum, Rosa spinosissima-Salix arenaria-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2170 - Dd 19-f, 20-a, 36A2

KGK - KHs X 2170

Notes: Creeping Willow (Salix repens) is the dominant species of this dune type with shrubby vege-tation at rather dry sites. Burnet Rose (Rosa spinosissima; syn. R. pimpinellifolia) and Round-Leaved Wintergreen (Pyrola rotundifolia, see small picture) are further typical species.


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TMAPCode:


X.7.3 Rosa canina type Rosa canina, Rosa rubiginosa, Rubus fruticosus s.l., Crataegus spp., less Betula spp., less Populus tremula, Sambucus nigra, Sorbus aucuparia, etc. (at least 1 species dominant)and +/- Salix repens (ssp. dunensis), less Hippophaë rhamnoides, Lonicera periclymenum, Rubus caesius, Carex arenaria, Ammophila arenaria, Galium mollugo, Festuca rubra agg., Poa pratensis

Rhamno-Prunetea-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

as sur-rounding Natura 2000 type

- Dd 37-f,37B-b,37-c,37-e,14-m,15-d,17A-b

KGH - - GD -

Notes: At dry to mesic sites, tall-growing shrubby vegetation dominated by Roses (Rosa sp., e.g. Rosa canina, see small picture), Common Hawthorn (Crataegus monogyna) or other woody species is developing.


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TMAPCode:


X.7.4 Rosa rugosa type Rosa rugosa (dominant)

and +/- species of the X.5/X.6/X.7

Rosa rugosa-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

- - Dd - KGX - - GR -

Notes: This type is dominated by the non-native Japanese Rose (Rosa rugosa, see picture). Stands often occur close to human settlements and/or were originally planted.


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TMAPCode:


X.8.0 Dune woodland, unspecific

Populus alba, Prunus serotina(at least one species dominant)and +/- species of X.8


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

NL, DK 2180

- Dd 37B-b,41A/c,43-a,43-h,43-i

KGY - KHx UW -

Notes: Shrubby vegetation dominated by non-native species like White Poplar (Populus alba, see picture), Rum Cherry (Prunus serotina) or other woody species.


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TMAPCode:


X.8.1 Populus tremula type

Populus tremula, Betula pendula / Betula pubescens (at least one spe-cies dominant)and +/- Sorbus aucuparia, Carex arenaria, Agrostis capillaris, Anthox-anthum odoratum, Dryopteris dilatata, Empetrum nigrum, Holcus lanatus, Lonicera periclymenum

Populus tremula / Betula pendula Quercus robur-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2180 - Dd 43-j KGP - KHt PT 2180

Notes: Shrubby vegetation at dry to mesic sites dominated by Eurasian Aspen (Populus tremula, see picture) or Birch trees (Betula pendula).


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TMAPCode:


X.8.2 Pinus spp. type Pinus sylvestris, Pinus mugo, Pinus nigra (at least one species domi-nant) and +/- species of X.8

Pinus spp.-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

- - Dd 41-a, 41-b,41A-b,41A/a,41A2,A3

KGY,WZK

- KHp KI -

Notes: Coniferous forests at dry sites, usually with Scots Pine (Pinus sylvestris, see picture), Moun-tain Pine (Pinus mugo) or European Black Pine (Pinus nigra).


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TMAPCode:


X.8.3 Quercus type Quercus robur (dominant)and +/- Betula pendula, Betula pubescens, Populus tremula, Sorbus aucuparia, Agrostis capillaris, Anth-oxanthum odoratum, Carex arenaria, Dryopteris dilatata, Empetrum nigrum, Holcus lanatus, Lonicera pericly-menum


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2180 - Dd 42A1,2, 42A-a,b, 42A-c, 43A1, 43A-b, 43B-c,d, 43B-e, 43B2, 43B-b,c, 43B-d, 43-d

KGQ - KHq QR 2180

Notes: Deciduous forest at dry to mesic sites dominated by Pedunculate Oak (Quercus robur, see picture).


25


TMAPCode:


X.10 Ruderal dune areas Epilobium angustifolium, Cirsium arvense, Cirsium vulgare, Urtica dioica, Urtica urens, Rubus caesius, Anchusa officinalis, Rubus frutico-sus, Linaria vulgaris (at least one species dominant)and +/- species of X.5

dry. veg. of Urtica spp., Epilobium angustifolium


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK


- Dd,Rrl

14/a, 14E-a, 14-h, 15-d,e, 17A2, 17A-a, 31-e, 33-a, 33A1,4, 33A5, 33/a, 34A1, 37-g

KDR KDW, KDG

KDr EA -

Notes: Dune areas dominated by species indicating nutrient enrichment such as Fireweed (Epilobium angustifolium, see picture), Creeping Thistle (Cirsium arvense) or Stinging Nettle (Urtica dioica).


26


TMAPCode:


X.12 Drifting sand duneApplied in the country/federal state:

Standard TMAP


Denmark

X N N N X X

Reference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

as sur-rounding Natura 2000 type (mostly 2120)

- Dd KDO - - KDm WD 2120

Notes: Drifting dune mostly without vegetation (see picture). Marram Grass (Ammophila arenaria) might occur occasionally (X.4.1).


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TMAPCode:


H.0 Dune slacks (humid)Applied in the country/federal state:

Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv - KN,KB

KDT - - 2190

Notes: Wet dune slacks arise either after the accretion of a new dune line (primary dune slack; mostly linear in shape and parallel to the coastline) or if sand from older dunes is blown out by wind (secondary dune slack; mostly of roundish appearance).

4 Standard TMAP Types of Dune Slacks

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TMAPCode:


H.1.1 Centaurium littorale type

Centaurium littorale, Sagina nodosa (at least one species)and +/- species of hygroserie: Juncus anceps, Agrostis stolonifera, Potentilla anserina, Juncus articulatus, Holcus lanatus, Mentha aquatica, Linum catharticum, Carex flacca, Bryum algovicum and +/- species of xeroserie: Leon-todon saxatilis, Carex arenaria, Lotus corniculatus, Honckenya peploides, Festuca rubra, Ammophila arenaria, Sonchus arvensis, Sedum acre, Hip-pophaë rhamnoidesand +/- species of haloserie: Plan-tago coronopus, Plantago maritima, Armeria maritima, Juncus gerardii, Odontites vernus, Glaux maritima, Carex distans, Carex extensa, Trifolium fragiferum

Centaurio-Saginetum (samoletosum, parnas-siotosum)


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 HSs Crs 27A2, 27A1

KNH KDT KPc CS 2190

Notes: Plant community representing the earliest stage of vegetation development in wet primary dune slacks (hygroserie). Seaside Centaury (Centaurium littorale, see picture) and Knotted Pearlwort (Sagina nodosa, see picture) are characteristic species. This plant community is related to those of the xero- and haloserie, because dune and salt marsh species do also occur here.

Standard TMAP Types of Dune Slacks

29


TMAPCode:


H.1.2 Radiola linoides type Radiola linoides, Anagallis minima, Juncus pygmaeus, Cicendia fili-formis (at least one species)and +/- Juncus bufonius, Gnapha-lium uliginosum, Isolepis setacea, Eleocharis quinqueflora, Centaurium pulchellum, Sagina procumbens, Carex viridula, Plantago major, Poa annua, Calliergonella cuspidata, Drepano-cladus aduncus, Ceratodon purpureus, Bryum spec, Pohlia nutans, Scapania irrigua, Fossombronia foveolata, Jungermannia gracillima, Cephalozia bicuspidata and +/- species of H.1.4/H.3.1or +/- species of H.2/H.2.1or +/- species of S.0 and H.1.1

Cicendietum filiformis, Isoeto-Nanojuncetea-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv 28A1 KNP (KNH)

KDT KPr RL 2190

Notes: The early successional Dwarf Rush community occurs at sites with different site conditions. The characteristic species All-Seed (Radiola linoides, see picture) and Chaffweed (Anagallis minima, see small picture) are commonly occurring, while Pygmy Rush (Juncus pygmaeus) and Yellow Centaury (Cicendia filiformis) occur very rarely.


30


TMAPCode:


H.1.3 Littorella uniflora type

Littorella uniflora, Potamogeton polygonifolius, Eleocharis multicaulis, Deschampsia setacea, Apium inunda-tum, Pilularia globulifera, Scirpus flui-tans, Baldellia ranunculoides, Samolus valerandi, Potamogeton gramineus, Juncus bulbosus, Lythrum portula, Spar-ganium minimum (at least one species)and +/- Hydrocotyle vulgaris, Ranun-culus flammula, Carex viridula, Juncus articulatus, Juncus anceps, Potentilla anserina, Eleocharis palustris ssp. palustris, uniglumis, Agrostis stolonifera, Carex nigra, Phragmites australis, Drepanocladus spec., Calliergonella cuspidata.and +/- (wet situation) Utricularia spp. (often U. australis), Chara spp. (often C. globularis), Myriophyllum spp. (often M. spicatum), Potamogeton spp. (often P. natans), Ranunculus peltatus ssp. baudotii, Hippuris vulgaris, Polygonum amphibium. (basic situation) Mentha aquatica, Carex flacca, Campylium polygamum, and rare: Scorpidium scorpioides, Drepanocladus sendtneri, Scorpidium revolvens

Littorelletea uniflorae-veg.


Standard TMAP


Denmark

X X X N X -


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv 06-a, 06C4

KNP (KNKn, KNT)

KDT KPl LU 2190


Littorella uniflora (1)

31


Notes: The early-successional vegetation type is named after the Shoreweed (Littorella uniflora, see picture 1). It is occurring in wet dune slacks, which regularly dry out during summer. Vegetation is characterized by a variety of typical species and can be divided into different sub-types. At sites with a rather high soil pH (pictures 2-3), Lesser Water-Plantain (Baldel-lia ranunculoides), Various-Leaved Pondweed (Potamogeton gramineus) and Brookweed (Samolus valerandi) regularly occur. At sites with a low soil pH (picture 4), Bog Hair-Grass (Deschampsia setacea) is a typical species.


Baldellia ranunculoides and Potamogeton gramineus (2)

Samolus valerandi (3) leftDeschampsia setacea (4)

right

32


TMAPCode:


H.1.4 Lycopodiellainundata type

Lycopodiella inundata, Rhynchospora alba, Rhynchospora fusca, Drosera intermedia (at least one species, and rare Hammarbya paludosa)and +/- species of H.3.1: Erica tetralix, Drosera rotundifolia, Vac-cinium uliginosum, Empetrum nigrum, Narthecium ossifragum, Drosera rotundifolia, Oxycoccus palustris, Gen-tiana pneumonanthe, Molinia caeru-lea, Hydrocotyle vulgaris, Carex nigra, Carex panicea, Eriophorum angusti-folium, Juncus bulbosus, Salix repens, Polytrichum commune, Dicranum scoparium, Sphagnum denticulatum, Drepanocladus exannulatus and other Drepanocladus, Sphagnum spec. and +/- Liverworts: Gymnocolea in-flata, Scapania irrigua, Fossombronia foveolata, Jungermannia gracillima, Cephalozia bicuspidata

Lycopodiella inundata-veg., Sphagno-Rhyn-chosporetum, Lycopo-dio-Rhynchosporetum


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv 11A1 KNE KDT KPi LI 2190

Notes: A characteristic species of this pioneer community is the Marsh Clubmoss (Lycopodiella inundata, see picture). This vegetation type usually occurs in small patches within wet heathlands (H.3.1), often in areas with bare soil resulting from anthropogenic disturbances.


33


TMAPCode:


H.2.1 Carex trinervis type Carex nigra, Carex trinervis, Poten-tilla palustris (at least one species dominant)and +/- Salix repens agg. (often dominant), Calamagrostis canescens, Eriophorum angustifolium, Epilobium palustre, Stellaria palustre, Galium palustre, Viola palustris, Menyanthes trifoliata, Lycopus europaeus, Drepan-ocladus aduncus, Calliergon cordifo-lium, Sphagnum fimbriatumand +/- species of H.2

Caricetum trinervi-nigrae, Caricion nigrae veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv 09A1, 09A3, 09A-a, 09-a,b,c, 09-e,f,h, 09B-b, 09/c

KNA KDT KMm CN 2190

Notes: This vegetation type is characterized by the Three-Nerved Sedge (Carex trinervis, see small picture). It occurs in older, acidic dune slacks with generally high water level fluctuations. The vegetation is often a homogeneous mixture of sedges and creeping willows.


34


TMAPCode:


H.2.2 Schoenus nigricans type

Schoenus nigricans, Liparis loeselii, Parnassia palustris, Epipactis palus-tris, Dactylorhiza incarnata, Pellia endiviifolia, Campylium polygamum, C. stellatum, Bryum pseudotri-quetrum, and rare: Gentianella amarella, Carex pulicaris, Juncus subnodulosus, Taraxacum palus-tre, Fissidens adianthoides, Aneura pinguis, Preissia quadrata, Moerckia hibernica (minimum three species)and +/- Mentha aquatica, Eleocharis quinqueflora, Carex flacca, Linum catharticum, and rare: Equisetum var-iegatum, Pedicularis palustris, Juncus arcticus ssp. balticus, Gymnadenia conopsea, Anagallis tenellaand +/- species of H.2and +/- species of H.1.1/H.1.2

Junco baltici-Schoene-tum nigricantis, Juncus subnodulosus-veg., Parnassio-Juncetum atricapilli, Caricion davallianae veg.


Standard TMAP


Denmark

X X X N N -Reference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 HSc Dv, Dvs, Dvp

09C3 KNK KDT - - 2190


Schoenus nigricans (1)

35


Notes: This calcareous fen, which is characterized by Schoenus nigricans (see picture 1), usually harbors several plant species with colorful flowers, e.g. Marsh Grass-of-Parnassus (Par-nassia palustris; see picture 2), Fen Orchid (Liparis loeselii) and Early Marsh Orchid (Dac-tylorhiza incarnata, see picture 3). It colonizes pioneer sites in weakly brackish, young primary dune slacks, in secondary dune slacks and in enclosed or reclaimed coastal areas. Under favorable conditions, the vegetation type is persistent for long periods. This com-munity contains higher numbers of “Red list species” than all other plant communities of the Wadden Sea islands.


Parnassia palustris (2)

Dactylorhiza incarnata (3)

36


TMAPCode:


H.3.1 Erica tetralix type Erica tetralix (dominant) and +/- Empetrum nigrum, Narthe-cium ossifragum, Vaccinium uligino-sum, Vaccinium oxycoccos, Vaccinium macrocarpos, Drosera rotundifolia, Juncus squarrosus, Gentiana pneu-monanthe, Scirpus cespitosus ssp. germanicus, Aulacomnium palustre, Polytrichum commune, Sphagnum fimbriatum, S. subnitens, S. palustre, S. fallax, S. squarrosum, S. molleand +/- Salix repens agg., Cal-luna vulgaris, Potentilla erecta, Viola palustris, Carex trinervis, Carex panicea, Agrostis canina, Eriophorum angustifo-lium, Pedicularis sylvatica, Danthonia decumbens, Molinia caerulea, Nardus stricta, Hypnum jutlandicum / cupres-siforme, Pleurozium schreberi, Dicranum scoparium, Pseudoscleropodium purum

Empetro-Ericetum, Narthecium ossifragum-veg.


Standard TMAP


Denmark

X X X N X -


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK2190 - Dv 11A3 KNE KDT KMt ET 2190

Notes: The coastal wet heathlands are characterized by the dominance of Cross-leaved Heath (Erica tetralix, see pictures). The community occurs in old, acidic dune slacks.


37


TMAPCode:


H.4.1 Phragmites australis type

Phragmites australis (dominant) and +/- Iris pseudacorus, Typha angustifolia, Typha latifolia, Schoeno-plectus tabernaemontaniand +/- species of H.2

Scirpo-Phragmitetum, Schoenoplecto-Phrag-mitetum, Typho-Phrag-mitetum


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv 08B2, 08B3, 26-b

KNR KDT KMr R 2190

Notes: This community occurs in wet dune slacks and consists of reed beds dominated by Common Reed (Phragmites australis, see picture).


38


TMAPCode:


H.5.1 Salix cinerea type Salix cinerea (dominant), Salix repens agg.

and +/- species of H.2/H.4/H.6

Salix cinerea-Salix arenaria-veg., Salice-tum cinereae – saliceto-sum repentis


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 (NL, DK 2170)

- Dv 36A2 KBH KDT KMw SA 2170

Notes: This tall-growing willow shrubbery occurs in moist to wet dune slacks. It is usually domi-nated by Grey Willow (Salix cinerea, dominant, see picture). Creeping Willow (Salix repens) is another, commonly occurring species.


39


TMAPCode:


H.5.2 Myrica gale type Myrica gale (dominant), Salixrepens agg.

and +/- species of H.2/H.4/H.6

Myricetum galis


Standard TMAP


Denmark

X X X N N NReference to other typologies:

Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 (NL, DK 2170)

- Dv 09-h, 36A-b

KBH KD - - 2170

Notes: This tall-growing shrubbery dominated by Bog Myrtle (Myrica gale, see picture) does mainly occur on the West Frisian Islands in moist to wet dune slacks with a low soil pH.


40


TMAPCode:


H.6.1 Betula pubescens type

Betula pubescens (dominant)and +/- Carex nigra, Calamagrostis canescens, Hydrocotyle vulgaris, Men-tha aquatica, Phragmites australis, Erica tetralix, Salix repens agg., Salix cinerea, Calliergonella cuspidata, Sphagnum spp. and +/- Calamagrostis epigejos, Carex arenaria, Lonicera periclymenum, Agrostis capillaris, Empetrum nigrum, Dicranum scoparium, Rubus spec.

Betula pubescens-veg., Empetro-Betuletum carpaticae


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2180 - Dv 40A2, 40A-c, 40A-d

KBA, KBR

KDT KMb BP 2180

Notes: This forest community in moist to wet dune slacks is mostly dominated by Downy Birch (Betula pubescens, see picture). In the herb layer, plant species indicating high soil moisture like Common Sedge (Carex nigra), Purple Small-Reed (Calamagrostis canescens) or Com-mon Reed (Phragmites australis) do typically occurr.


41


TMAPCode:


H.6.2 Alnus glutinosa type Alnus glutinosa (dominant)and +/- Carex nigra, Calamagrostis canescens, Hydrocotyle vulgaris, Men-tha aquatica, Phragmites australis, Iris pseudacorus, Erica tetralix, Salix repens agg., Salix cinerea, Calliergo-nella cuspidata, Sphagnum spec.and +/- Calamagrostis epigejos, Carex arenaria, Lonicera periclymenum, Agrostis capillaris, Empetrum nigrum, Ribes nigrum, Dicranum scoparium, Rubus spec., Solanum dulcamara, Glyceria maxima, Thypha spec., Carex riparia

Alnus glutinosa-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2180 - Dv 39A1,2, 39A-a,b, 39A-d,e, 39A-f

KBE KDT KMe AG 2180

Notes: This forest community in moist to wet dune slacks is dominated by Common Alder (Alnus glutinosa, see picture). As in type H.6.1, plant species indicating high soil moisture like Common Sedge (Carex nigra), Purple Small-Reed (Calamagrostis canescens) or Common Reed (Phragmites australis) do typically occurr in the herb layer.


42


TMAPCode:


H.8 Aquatic vegetation in dune slacks - Hydroserie

Utricularia spp. (often U. austra-lis), Chara spp. (often C. globula-ris), Myriophyllum spp. (often M. spicatum), Potamogeton spp. (often P. natans), Ruppia spp., Ranunculus peltatus ssp. baudotii, etc. (at least one species dominant)

Charetea fragilis, Pota-mogetoneta, Utriculari-etea-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv 04-a, 04B2,3, 04C1,3, 04D1a, 04D1b, 05A1,2, 05A-a, 05E3, 05-f,h

KNT KDT KMf HY 2190

Notes: Aquatic dune slack vegetation with Bladderworts (Utricularia), Watermilfoil (Myriophyl-lum), Pondweed (Potamogeton) or Chara (see picture).


43


TMAPCode:


H.9 Ruderal Dune slack areas with litter brushwood

Epilobium hirsutum, Cirsium vul-gare, arvense. (at least one species dominant)and +/- species of H.2/H.4

Moist vegetation of Epilobium hirsutum,Cirsium vulgare a. arvense.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

2190 - Dv 32B2, 32B3, 32A1, 32-a, 32-b, 32-c, 32-d, 32-e, 32-f, 32-g

KNS KDT KMh EH 2190

Notes: Areas in dune slacks with vegetation dominanted by species indicating high nutrient avail-ability like Great Willowherb (Epilobium hirsutum, see picture), Spear Thistle (Cirsium vul-gare) or Creeping Thistle (Cirsium arvense).


44


TMAPCode:


S.0.0 Vegetation not pre-sent, disturbed


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK


Su* - 50A, 50B, 50C

- KWO KNx 90 -

Notes: Highly disturbed sites with bare soil without vegetation (see picture) within salt marshes are mapped as type S.0.0 (vegetation not present, unspecific).

5 Standard TMAP Types of Salt Marshes

Standard TMAP Types of Salt Marshes

45


Notes: Old creeks or inundated areas are mapped as open water without vegetation.

TMAPCode:


S.0.1 Open water


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK


Sw water - KLA, KLZ

- KNt 55 -


46


Notes: Bare sand or shell is mapped as type S.0.2.

TMAPCode:


S.0.2 Bare sand or shells in salt marsh sys-tems


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK


Ss zand; schelp

- (KS) KWO KNo 88 -


47


Notes: The type ‘bare mud or clay’ is found at inundated sites in the salt marsh area or along the salt marsh shore below mean high tide.

TMAPCode:


S.0.3 Bare mud or clay in salt marsh systems


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK


Sm slik, Qq0p, Qq0e, QuO, Ss0

- (KWK, KWB)

KWO KWv VO -


48


Notes: Common Cordgrass (Spartina anglica, see picture) is the first perennial species establishing around mean high tide. This vegetation can accelerate sedimentation and thus facilitate salt marsh development.

TMAPCode:


S.1.1 Spartina anglica type

Spartina anglica Spartinetum anglicae


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1320 SPs Ss5, Ss3, Bs3, Bs5

24A1 KWG KWG KNs S 1320


49


Notes: This pioneer vegetation is characterized by a few annual plants as Long-Spiked Glasswort (Salicornia stricta) and Common Glasswort (Salicornia europaea, see picture). At nutrient-rich sites, the Annual Sea-blite (Suaeda maritima) and the Hairy Sea-blite (Bassia hirsuta, see small picture) can be dominant.

TMAPCode:


S.1.2 Salicornia type Salicornia spp., Suaeda maritima, Bassia hirsuta (at least one species dominant)

Salicornietum strictae, Salicornietum rammo-sissimae, Salicornietum decumbentis, Suaede-tum maritimae, Thero-Salicornion strictae-veg., Suaedo-Bassietum hirsutae


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1310 SPq Qqp, Qqe, Qu

25A2, 25A-a, 26A-c

KWQ, KWQW, KWQV, KSF

KWQ KQb, KQd, KQr, KQs,KQu

SS, SR, SD, U, BH

1310


50


Notes: Common Saltmarsh-Grass (Puccinellia maritima, see picture) is the characteristic species of the low salt marsh. The plant tissue of this species contains a high content of proteins and is therefore favoured by many herbivores.

TMAPCode:


S.2.1 Puccinellia maritima type

Puccinellia maritima (dominant)and +/- Spergularia spp., Salicornia spp., Puccinellia distans, Spartina anglica, Suaeda maritima, Limonium vulgare, Aster tripolium

Puccinellietum mariti-mae


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SLp P, Pp, Ppq, P-d, P-q, P-u, Pps, Ppu, Pe, Pt, Pex, Pj, Pg

26A1, 26A-a

KHUP KHU, KHW

KNd, KNp, KNv, KNw

P, Ps 1330


51


Notes: Common Sea-Lavender (Limonium vulgare, see picture), a species with its optimum in the low salt marsh and with beautiful flowers, can create whole areas of purple colors. The flowers are also important for many nectarivore insects like butterflies (see small picture).

TMAPCode:


S.2.2 Limonium vulgare / Puccinellia maritima type

Limonium vulgare and +/- Plantago maritima, Puccinel-lia maritima, Spergularia maritima, Aster tripolium

(Plantagini-Limonietum)


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SLl Ppl, Pl 26A2 KHUL KHU, KHW

- P 1330


52


Notes: This vegetation is characterized by the dominance of Sea Aster (Aster tripolium, see pic-ture). The species with purple flowers is attractive for many nectarivore insects (see small picture) and is also favoured by herbivores. In moderately grazed brackish low salt marshes the species can reach a height of up to 1.5 meters.

TMAPCode:


S.2.3 Aster tripolium / Puccinellia maritima type

Aster tripolium (dominant)and +/- Puccinellia maritima, Sper-gularia maritima, Limonium vulgare, Plantago maritima

Aster tripolium-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 Sla Ppa,Pa,Jfa,Jja, Ppab, Ba

26A-b KHUA KHU, KHW

KNa A 1330


53


Notes: This low shrubby vegetation is characterized by the dominance of Sea Purslane (Atriplex portulacoides, see picture). This type mainly occurrs in the low salt marsh on clayey soils. During cold winters, this vegetation severely suffers from ice formation.

TMAPCode:


S.2.4 Atriplex portula-coides type

Atriplex portulacoides (dominant)and +/- Puccinellia maritima, Spergu-laria maritima, Salicornia spp., Suaeda maritima

Halimionetum portula-coidis


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SLh Ph 26A3 KHUH KHU, KHW

KNh H 1330


54


Notes: This vegetation type with the Common Sea-Lavender (Limonium vulgare) occurs in in-undated areas at higher, sandy salt marshes. Other characteristic species are Saltmarsh Rush (Juncus gerardii), Sea Milkwort (Glaux maritima), Red Fescue (Festuca rubra) and Sea Wormwood (Artemisia maritima).

TMAPCode:


S.3.1 Limonium vulgare / Juncus gerardi type

Limonium vulgare (dominant), Jun-cus gerardii, Glaux maritimaand +/- species of S.3

(Plantagini-Limonietum)


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHl Jjl, Jfl 26A2, 26C-a, 26-a

KHOL KHO - - 1330


55


Notes: This usually grazed vegetation type with a dominance of Saltmarsh Rush (Juncus gerardii, see picture) occurs at regularly inundated areas at high sandy salt marshes. Sea Milkwort (Glaux maritima) is often co-dominant.

TMAPCode:


S.3.2 Juncus gerardi type Juncus gerardi Glaux maritima and +/- species of S.3

Juncetum gerardii


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHj Jj, Jex 26C1, 26C-c

KHOJ KHO, KHI

KOj J 1330


56


Notes: This vegetation of the high salt marsh is dominated by Red Fescue (Festuca rubra). Other often occurring species are Sea Thrift (Armeria maritima), Creeping Bentgrass (Agrostis stolonifera) and Sea Couch Grass (Elytrigia atherica).

TMAPCode:


S.3.3 Festuca rubra type Festuca rubra (dominant) and +/- species of S.3

Festuca rubra-veg., Armerio-Festucetum littoralis


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHf Jf 26C-b KHOR KHO, KHI

KOf F 1330


57


Notes: This vegetation occurs at high marsh sites and is dominated by Sea Wormwood (Artemisia maritima, see picture). Other regularly occurring species are Red Fescue (Festuca rubra) and Sea Couch Grass (Elytrigia atherica).

TMAPCode:


S.3.5 Artemisia maritima / Festuca rubra type

Artemisia maritima (dominant) and +/- Festuca rubra, Elymus atheri-cus

Artemisietum maritimae


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHz Pz, Jf-z, Jz

26-c KHOB KHO, KHI

KOt T 1330


58


Notes: This vegetation of the high salt marsh is dominated by Sea Rush (Juncus maritimus, see picture). Due to its stiff and stinging leaves it is hardly grazed by herbivores.

TMAPCode:


S.3.6 Juncus maritimus type

Juncus maritimus (dominant) and +/- Festuca rubra, Juncusgerardii, Elymus athericus

Juncus maritimus-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHm Pm, Jm

26-e KHBN KHO, KHB

KOm MG 1330


59


Notes: The vegetation dominated by Sea Couch Grass (Elytrigia atherica) is late successional stage at high marshes, often occurring on ungrazed sites. Vegetation is mostly species-poor be-cause of the tall grass with a dense cover and a thick litter layer.

TMAPCode:


S.3.7 Elymus athericus type

Elymus athericus (dominant)and +/- Festuca rubra, Atriplex prostrata

species of S.3

Agropyretum litoralis, Agropyretum pungentis, Atriplici-Agropyretum pungentis, Atriplici-Elymetum atherici, Elymetum pycnanthi, Astero-Agropyretum repentis, Elymus repens ssp. littoralis-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 Shy Py,Jy3,Jy5, Ry3, Ry5, By3,By5

26-d KHQA KHQ KOq Q 1330


60


Notes: Vegetation dominated by Long-Bracted Sedge (Carex extensa, see picture) is typical for high sandy salt marshes and upper beach plains. Other typical species are Lesser Centaury (Centaurium pulchellum), Red Bartsia (Odontites vernus), Sea Milkwort (Glaux maritima) and Sea Hard-Grass (Parapholis strigosa).

TMAPCode:


S.3.8 Carex extensa type Carex extensa and +/- Odontites vernus ssp. seroti-nus, Centaurium pulchellum, Glaux maritima, Juncus maritimus, Agrostis stolonifera, Parapholis strigosa and +/- species of S.3

Junco Caricetumextensae


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHe Je 26C2 KHS, KHBN

KHB KOc CE 1330


61


Notes: At high tide marks and guano-rich sites, a nutrient-rich vegetation with Orache (Atriplex) occurs (see picture). Typical species are Spear-Leaved Orache (Atriplex prostrata), Grass-Leaved Orache (Atriplex littoralis) and Sea Mayweed (Tripleurospermum maritimum).

TMAPCode:


S.3.9 Atriplex prostrata / A. littoralis type

Atriplex prostrata, A. littoralis(at least one species dominant) and +/- Tripleurospermum maritimum

Atriplex prostrata / lit-toralis veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHx Xx, Xk 22A1, 29A3

KHM KHO KOl, KOp

AP, AX 1330


62


Notes: This dense grassland vegetation is characterized by Creeping Bent (Agrostis stolonifera) and different Clovers such as Strawberry Clover (Trifolium fragiferum, see small picture) and White Clover (Trifolium repens). It occurs in high salt marshes and in low dunes. Other typical species are Common Silverweed (Potentilla anserina), Lesser Centaury (Centaurium pulchellum), Red Bartsia (Odontites vernus), Red Fescue (Festuca rubra) and Bird’s-Foot Trefoil (Lotus corniculatus).

TMAPCode:


S.3.10 Agrostis stolonifera / Trifolium fragifer-um type

Agrostis stoloniferaand +/- Potentilla anserina, Festuca rubra, Trifolium fragiferum, Trifolium repens, Lotus corniculatus and +/- species of S.3

Trifolio fragiferi-Agrostietum stoloniferae, Agrostis stolonifera-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHg Jg, Rgf, Rpf, Rgc, Rgl, Rgt

12B2, 12B3, 12B-m

KHB oder KHF

KHI, KHO

KOa W 1330


63


Notes: This type occurs on low brakish dunes. Distant Sedge (Carex distans) and Spiny Restharrow (Ononis spinosa, see small picture) are typical species.

TMAPCode:


S.3.12 Ononis spinosa / Carex distans type

Carex distans, Ononis spinosa (at least one species) and +/- species of S.3and +/- species of X.3/X.5

Ononido-Caricetum distantis


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHo Ro 12B4 KHBT KHB KOo OC 1330


64


Notes: On high salt marshes and low dunes vegetation with Couch Grass (Elytrigia repens) as the dominant species can occur (see picture). If Sea Couch Grass (Elytrigia atherica) is also present, it is likely to find the bastard (E. x oliveri), too. On the small picture: left E. repens, middle: E. x oliveri and right: E. atherica.

TMAPCode:


S.3.13 Elymus repens type Elymus repens (dominant)and +/- species of S.3/X.5

Agropyretum litoralis, Agropyretum pungentis, Atriplici-Agropyretum pungentis, Atriplici-Elymetum atherici, Elymetum pycnanthi, Astero-Agropyretum repentis, Elymus repens ssp. littoralis-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHr Re 26-d, 16/c

KHQR KHQ KOd DQ 1330


65


Notes: At ruderal high salt marshes and low dunes vegetation with Creeping Thistle (Cirsium ar-vense) can occur. At more nutrient-rich sites, Stinging Nettle (Urtica dioica) can be domi-nant. Formerly classified as S.6.0 in SH

TMAPCode:


S.3.14 Ruderal salt marsh areas

Cirsium arvense, Urtica dioica, Sonchus spec. (at least one species dominant)and +/- species of S.0


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SHu Rru,Rrc

- KHQS - KOh,KOr

RU, CA 1330


66


Notes: In brackish salt marshes at permanently wet sites, vegetation with Sea Club-Rush (Bol-boschoenus maritimus, see picture) or Softstem Bulrush (Schoenoplectus tabernaemontani) can occur. Other typical species are Common Reed (Phragmites australis) and Creeping Bentgrass (Agrostis stolonifera).

TMAPCode:


S.5.1 Bolboschoenus / Schoenoplectus type

Bolboschoenus maritimus and / or Schoenoplectus tabernaemontani (at least one species dominant) and +/- Phragmites australis, Aster tripolium, Agrostis stoloniferaand +/- species of S.5

Bolboschoenetum maritimae


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 (LS partly)

SBb Bis, Bi3, Bi5

12B-d, 26-b, 08B2

KRS, KWRT, KWRS

KBS KRb B 1330


67


Notes: If vegetation of the brackish marsh reaches its climax stage, it will be dominated by Com-mon Reed (Phragmites australis, see picture). It is very common in ungrazed areas, such as brackish dune slacks and at brackish marshes along estuaries.

TMAPCode:


S.5.2 Phragmites australis type

Phragmites australis (dominant) and +/- Atriplex prostrataand +/- species of S.5

Phragmites australis-veg. (brackish marsh)


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 (LS partly)

SBp Bb, Bc 08-f, 26-a

KRP, KWRP

KBP KRs RH 1330


68


Notes: This community is an assemblage of multiple brackish pioneer and grassland communi-ties. Most common species are Creeping Bentgrass (Agrostis stolonifera) and Common Sil-verweed (Potentilla anserina). Common Saltmarsh-Grass (Puccinellia maritima), Saltmarsh Rush (Juncus gerardii), Slender Spike-Rush (Eleocharis uniglumis) and at some places Red Goosefoot (Chenopodium rubrum), Saltmarsh Flat-Sedge (Blysmus rufus) or Golden Buttons (Cotula coronopifolia, see pictures) can be dominant.

TMAPCode:


S.5.3 Brackish flooded grassland type

Agrostis stolonifera, Potentilla anserina, Blysmus rufus, Cotula coronopifolia, Eleocharis uniglumis, Triglochin palustre and +/- species of S.5

Blysmetum rufi, Eleo-charis uniglumis-veg., Cotula coronopifolia-veg., Alopecurus bulbosus-veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

1330 SBg P—b, Pp-b, Pe-b, Bcs, Bg, Bt, Bj, Br, Bgt, Bgn, Bpj, Bo, Bpg, Bp

26C3 KHBN, KHF

KHB KRc, KRe, KRr

BR, EU, C

1330


69


Notes: This vegetation is mostly intensively grazed and occurs in high salt marshes, summer pol-ders. Due to antropogenic influences in such areas, it’s not a real salt or brackish vegetation by definition. These areas might be of less interest for nature managers, and are therefore mapped provisionally as ‘grassland unspecific’.

TMAPCode:


S.6.0 Fresh grasslands, unspecific


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK

- SF* - 12B-m G-types G KOr,KOh

CA, SP -


70


Notes: At high salt marshes, summer dikes (see picture) and summer polders in intensively man-aged (grazed) areas, a vegetation with glycophytes can occur. Typical species are Perennial Ryegrass (Lolium perenne, see small picture), Crested Dogstail Grass (Cynosurus cristatus), White Clover (Trifolium repens), Meadow Barley (Hordeum secalinum), Broadleaf Plantain (Plantago major) and Common Silverweed (Potentilla anserina).

TMAPCode:


S.6.1 Lolium perenne, Cynosurus cristatus and other fresh spe-cies types

Lolium perenne, Cynosurus crista-tus, etc.and +/- species of S.6

Lolio-Cynosuretum, Plantagini-Lolietum perennis, Polygonion veg.


Standard TMAP


Denmark


Natura 2000 CodeEU

Mapping Code

NL



Biotope Code

LS

Biotope Code

HH

Biotope Code

SH

Mapping Code

SH

Mapping Code

DK


SFl Dd, Dv, Rgv, Rp, Rgh

16C4 GM, Gl GM KGf, KGg, KGy

AR, MO, Y

-


71


6 Overview of the complete TMAP Typology

72


Overview of the complete TMAP Typology

TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

X.0

Dune

s

XEDd

KD

KDKD

ySW

X.1

Beac

h pl

ains

No

vege

tatio

n

Ss,

X.1

zand

KS

, KS

N,

KSB,

KS

P,

KSI,

(KSA

)

KSKS

s99

X.2

Beac

h dr

iftl

ine

2110

, 12

10,

1220

XE,

SU*

veek

KSV

CM

, CR

X.2.

0

Be

ach

drif

tlin

e,

unsp

ecifi

c21

10,

1210

, 12

20

XE*

KS

V

CM,

CR

X.2.

1X

XX

XX

Ca

kile

mar

itim

a ty

peCa

kile

mar

itim

a an

d +/

Sa

lsol

a ka

li, H

onck

enya

pep

loid

es, A

trip

lex

spec

ies,

Elym

us

farc

tus

ssp.

bor

eoat

lant

icus

Caki

letu

m m

ariti

mae

1210

, (in

LS

2110

)

XEd

Dxc,

Dx

h,

Deu,

Dx

s

22a

, 22

B1(K

DV)

KSV

KSa

CM21

10

X.2.

2X

NN

NX

Cr

ambe

mar

itim

a ty

peCr

ambe

mar

itim

a an

d +/

H

onck

enya

pep

loid

es, S

ilene

vul

garis

ssp.

mar

itim

a, G

lauc

ium

fla

vum

, Ery

ngiu

m m

ariti

mum

, Rum

ex c

rispu

s, Le

ymus

are

nariu

s

Crith

mo-

Cram

betu

m, H

onke

-ny

a pe

ploi

des a

nd

Sile

ne v

ulga

ris s

sp.

mar

itim

a ve

g.

1220

KSV

KSv

CR

X.3

Embr

yoni

c du

nes

2110

XEDd

KD

VKD

V

EG21

10X.

3.0

Embr

yoni

c du

nes,

unsp

ecifi

c 21

10XE

*Dd

KD

VKD

V

EG21

10

X.3.

1X

XX

XX

XEl

ymus

far

ctus

ty

peEl

ymus

far

ctus

spp

. bor

eoat

lant

icus

(do

min

ant)

an

d +/

Le

ymus

are

nariu

s, Am

mop

hila

are

naria

, Hon

cken

ya p

eplo

ides

, Sa

lsol

a ka

li, C

akila

mar

itim

e

Elym

o-Ag

ropy

retu

m,

Hon

keny

o-Ag

ropy

re-

tum

junc

ei

2110

Xef

Def,

Deg,

Dd

22A1

, 22

B1,

22B

a,

22a

, 23

A1,

23B

a

KDV

KDV

KDv

EG21

10

X.4

Whi

te d

unes

21

20

Dd

KDW

KDW

EM

, E21

20

X.4.

0

W

hite

dun

es,

unsp

ecifi

c21

20

Dd50

A,

50B,

50

C

KDW

KDW

EM

, E21

20

X.4.

1X

XX

XX

XAm

mop

hila

ar

enar

ia t

ype

Amm

ophi

la a

rena

ria, A

mm

ocal

amag

rost

is ba

ltica

, Ley

mus

are

na

rius

(at

leas

t on

e sp

ecie

s do

min

ant)

an

d +/

So

nchu

s arv

ense

, Oen

othe

ra p

arvi

flora

var

. am

mop

hila

, Hie

-ra

cium

um

bella

tum

, Cal

yste

gia

sold

anel

la (r

are)

, Ery

ngiu

m m

ariti

mum

(r

are)

, Lat

hyru

s mar

itim

us (r

are)

, Fes

tuca

rubr

a ss

p. a

rena

ria, C

arex

ar

enar

ia, H

onck

enya

pep

loid

es, E

lym

us fa

rctu

s ss

p. b

oreo

atla

ntic

us,

Caki

le m

ariti

ma

Elym

o-Am

mop

hi-

letu

m, x

Cal

amm

o-ph

ila b

altic

a-ve

g.,

Pote

ntill

o-Le

ymet

um

aren

aria

e (P

oten

til-

lo-E

lym

etum

)

2120

Dd

23B1

, 23

Ba,

23

a

KDW

KDW

KDw

EM, E

2120

73



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

X.5

XX

XX

XX

Dune

gra

ssla

ndsp

ecie

s of

X.5

: Car

ex a

rena

ria, A

gros

tis c

apill

aris

, Fes

tuca

rubr

a ss

p.

aren

aria

, Aira

pra

ecox

, Ant

hoxa

nthu

m o

dora

tum

, Luz

ula

cam

pest

ris,

Viol

a ca

nina

, Vio

la tr

icol

or (s

sp. c

urtis

ii), J

asio

ne m

onta

na, H

iera

cium

um

bella

tum

, Rum

ex a

ceto

sella

, Hyp

ocho

eris

radi

cata

, Gal

ium

mol

lugo

, Ce

rast

ium

sem

idec

andr

um, L

otus

cor

nicu

latu

s, H

ypnu

m c

upre

ssifo

rme,

Di

cran

um sc

opar

ium

, Pol

ytric

hum

juni

perin

um, C

ampy

lopu

s int

ro-

flexu

s, Ce

rato

don

pupu

reus

, Cet

raria

acu

leat

a Cl

adon

ia c

iliat

a,

C. p

orte

ntos

a, C

. fur

cata

, C. s

ubul

ata,

C. g

rayi

, C. c

ervi

corn

is, C

. ram

u-lo

sa

Koel

erio

-Cor

ynep

ho-

rete

a-ve

g.21

30

Dd14

c,

14d

, 14

g,

14h

, 14

l,

14m

KDG

KDG

KDg

TR21

30

X.5.

0

Du

ne g

rass

land

, un

spec

ific

2130

Dd

KD

GKD

G

TR21

30

X.5.

1

XX

X

Co

ryne

phor

us

cane

scen

s ty

pe

Cory

neph

orus

can

esce

ns

and

+/

Poly

tric

hum

pili

feru

m, C

lado

nia

arbu

scul

a, C

. coc

cife

ra, C

. flo

erke

ana,

C. g

raci

lis, C

. unc

ialis

an

d +/

sp

ecie

s of

X.5

an

d no

or l

ess

spec

ies

of X

.5.2

Viol

o-Co

ryne

phor

e-tu

m, C

oryn

epho

rion-

veg.

2130

Dd

14A1

, 14

A2KD

GAKD

G

TR21

30

X.5.

2

XX

X

Ko

eler

ia a

rena

ria

type

Ph

leum

are

nariu

m, T

ortu

la r

ural

is va

r. ru

ralif

orm

is, K

oele

ria a

re

naria

an

d +/

sp

ecie

s of

X.5

.2: S

ilene

otit

es, T

halic

trum

min

us, A

nthy

llis

vuln

erar

ia, G

aliu

m v

erum

, Trif

oliu

m a

rven

se, E

uphr

asia

str

icta

, Le

onto

don

saxa

tilis,

Rhi

nant

us m

inor

, Sed

um a

cre,

Poa

pra

tens

is ss

p. ir

rigat

a, M

yoso

tis r

amos

issim

a, S

cler

anth

us p

eren

nis,

Trifo

lium

ca

mpe

stre

, Sen

ecio

jaco

baea

, Bra

chyt

heci

um a

lbic

ans,

Hypn

um

lacu

nosu

m, R

acom

itriu

m c

anes

cens

, Cla

doni

a ra

ngifo

rmis,

C. f

olia

cea,

C. s

cabr

iusc

ula,

C. h

umili

s, C.

poc

illum

, Pel

tiger

a sp

ec. (

min

. 3

5 sp

ecie

s)

and

+/

spec

ies

of X

.5.3

an

d +/

sp

ecie

s of

X.5

Tort

ulo-

Phle

etum

, Ph

leo-

Tort

ulet

um,

Fest

uco-

Galie

tum

, Ai

retu

m p

raec

ocis

, Ko

eler

ion-

veg.

2130

Dd

14B1

, 14

D4,

14D

d,

14E1

, 14

Ea

KDGK

KDG

TR

2130

X.5.

3

XX

N

Bo

tryc

hium

lu

naria

typ

e N

ardu

s st

ricta

, Bot

rych

ium

luna

ria (

rare

), An

tenn

aria

dio

ica

(rar

e),

spec

ies

of t

he X

.5.3

: Pol

ygal

a vu

lgar

is, H

iera

cium

pilo

sella

, Fes

tuca

ov

ina,

Ver

onic

a of

ficin

alis,

Dan

thon

ia d

ecum

bens

, Pot

entil

la e

rect

a (m

inim

um 3

spe

cies

) an

d +/

sp

ecie

s of

X.5

an

d +/

sp

ecie

s of

X.5

.2

Botr

ychi

o-Po

lyga

le-

tum

, Nar

do-G

alio

n-ve

g.

2130

Dd

19A3

, 19

Aa,

19

Ac,

19

b,

19a

KDGB

KDG

TR

2130

X.5.

4

XX

X

Ca

rex

aren

aria

ty

pe

Care

x ar

enar

ia (

dom

inan

t)

and

+/

spec

ies

of X

.5

Care

x ar

enar

ia-v

eg.

2130

Dd

14i

, 14

jKD

GSKD

GKD

gTR

2130

X.5.

5

XX

De

scha

mps

ia

flexu

osa

type

De

scha

mps

ia fl

exuo

sa (

dom

inan

t)

and

+/

spec

ies

of X

.5

Desc

ham

psia

flexu

osa-

veg.

2130

Dd

19d

, 19

Ad

KDGS

KDG

KDg

TR21

30

X.6

N

Dune

hea

th

2140

Dd

KD

E,

KDC

HE,

CV

2140

74



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

X.6.

0

N

Du

ne h

eath

, un

spec

ific

Dd

KDE,

KD

C

H

E,CV

2140

X.6.

1X

XX

NX

XEm

petr

um

nigr

um t

ype

Empe

trum

nig

rum

(do

min

ant)

an

d +/

Po

lypo

dium

vul

gare

, Cal

luna

vul

garis

, Hie

raci

um u

mbe

llatu

m,

Salix

repe

ns a

gg.,

Loni

cera

per

icly

men

um, C

arex

are

naria

, Agr

ostis

ca

pilla

ris, A

ntho

xant

hum

odo

ratu

m, L

uzul

a ca

mpe

stris

, Fes

tuca

ovi

na,

Dant

honi

a de

cum

bens

, Dry

opte

ris d

ilata

ta. A

mm

ophi

la a

rena

ria, H

yp-

num

jutla

ndic

um /

cupr

essi

form

e, P

leur

oziu

m sc

hreb

eri,

Dicr

anum

sco-

pariu

m, P

seud

oscl

erop

odiu

m p

urum

, Hyl

ocom

ium

sple

nden

s, Cl

adon

ia

port

ento

sa, C

. fur

cata

Hie

raci

o-Em

pe-

tret

um, P

olyp

odio

-Em

petr

etum

2140

Dd

20A3

, 20

a,

11A3

, 11

b,

11/a

KDE

KD

eH

E21

40

X.6.

2X

XX

NX

Ca

lluna

vul

garis

ty

peCa

lluna

vul

garis

(do

min

ant)

an

d +

(nea

rly)

no E

mpe

trum

nig

rum

and

+/

Salix

repe

ns a

gg.,

Care

x ar

enar

ia, A

gros

tis c

apill

aris

, An-

thox

anth

um o

dora

tum

, Luz

ula

cam

pest

ris, F

estu

ca o

vina

, Dan

thon

ia

decu

mbe

ns, H

iera

cium

um

bella

tum

, Hyp

num

jutla

ndic

um /

cupr

essi

-fo

rme,

Ple

uroz

ium

schr

eber

i, Di

cran

um sc

opar

ium

, Pse

udos

cler

opod

ium

pu

rum

Hie

raci

o-Em

petr

e-tu

m -

dom

. Cal

luna

vu

lgar

is

2150

Dd

20A1

KDC

KD

cCV

X.7

Dune

scr

ub

Dd

KG

KHg

GD,

HR,

X

X.7.

0

Du

ne s

crub

, un

spec

ific

Dd

KG

GD

, H

R, X

X.7.

1X

XX

NX

XHi

ppop

haë

rham

noid

es t

ype

Hipp

opha

ë rh

amno

ides

(do

min

ant)

an

d +/

Sa

lix re

pens

(ssp

. dun

ensi

s), S

ambu

cus n

igra

, Car

ex a

rena

ria,

Amm

ophi

la a

rena

ria, G

aliu

m m

ollu

go, F

estu

ca ru

bra

agg.

, Poa

pra

ten-

sis,

Lina

ria v

ulga

ris, R

ubus

cae

sius

, Sen

ecio

jaco

baea

Hip

poph

ao-S

am-

buce

tum

nig

rae,

Sa

lici a

rena

riae-

Hip

poph

aetu

m

2160

Dd

37B1

, 37

B2,

37B3

, 37

Bc,

37

Bd,

37

Be

KGS

KH

hH

R21

60

X.7.

2X

XX

XX

XSa

lix r

epen

s ag

g.

type

Salix

rep

ens

(ssp

. dun

ensis

) (d

omin

ant)

an

d +/

Ro

sa sp

inos

issi

ma,

Pyr

ola

rotu

ndifo

lia

and

+/

Care

x ar

enar

ia, A

mm

ophi

la a

rena

ria, G

aliu

m m

ollu

go, F

estu

ca

rubr

a ag

g., P

oa p

rate

nsis

dry

Salix

repe

ns

agg.

-veg

., Py

rolo

-Sa

licet

um, R

osa

spin

osis

sim

a-Sa

lix

aren

aria

-veg

.

2170

Dd

19f

, 20

a,

36A2

KGK

KH

sX

2170

X.7.

3X

XX

NX

Ro

sa c

anin

a ty

pe

Rosa

can

ina,

Ros

a ru

bigi

nosa

, Rub

us f

rutic

osus

s.l.

, Cra

taeg

us s

pp.,

less

Bet

ula

spp.

, les

s Po

pulu

s tr

emul

a, S

ambu

cus

nigr

a, S

orbu

s au

cu

paria

(at

leas

t on

e sp

ecie

s do

min

ant)

an

d +/

Sa

lix re

pens

(ssp

. dun

ensi

s), l

ess

Hip

poph

aë rh

amno

ides

, Lo

nice

ra p

eric

lym

enum

, Rub

us c

aesi

us, C

arex

are

naria

, Am

mop

hila

are

-na

ria, G

aliu

m m

ollu

go, F

estu

ca ru

bra

agg.

, Poa

pra

tens

is

Rham

no-P

rune

tea-

veg.

unti

l 10

0 m

²

as s

ur

roun

din

g N

atur

a 20

00

type

Dd

37f

, 37

Bb,

37

c,

37e

, 14

m,

15d

, 17

Ab

KGH

KD

???

GD

X.7.

4X

XX

XX

Ro

sa r

ugos

a ty

pe

Rosa

rug

osa

(dom

inan

t)

and

+/

spec

ies

of t

he X

.5/X

.6/X

.7

Rosa

rugo

sa-v

eg.

Dd

KGX

KH

rGR

X.8

Dune

woo

dlan

d

Dd

36A2

KG

UW

,PT,

KI

, QR

75



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

X.8.

0X

XX

NX

Du

ne w

oodl

and,

un

spec

ific

Popu

lus

alba

, Pru

nus

sero

tina

(at

leas

t on

e sp

ecie

s do

min

ant)

an

d +/

sp

ecie

s of

X.8

(NL,

DK 2180

)

Dd

37B

b,

41A/

c,

43a

, 43

h,

43i

KGY

KH

xUW

X.8.

1X

XX

NX

XPo

pulu

s tr

emul

a ty

pe

Popu

lus

trem

ula,

Bet

ula

pend

ula

/ Bet

ula

pube

scen

s (a

t le

ast

1 sp

ecie

s do

min

ant)

an

d +/

So

rbus

auc

upar

ia, C

arex

are

naria

, Agr

ostis

cap

illar

is, A

nth-

oxan

thum

odo

ratu

m, D

ryop

teris

dila

tata

, Em

petr

um n

igru

m, H

olcu

s la

natu

s, Lo

nice

ra p

eric

lym

enum

Popu

lus t

rem

ula

/ Bet

ula

pend

ula

Que

rcus

robu

r-ve

g.

2180

Dd

43j

KGP

KH

tPT

2180

X.8.

2X

XX

NX

Pi

nus

spp.

typ

e

Pinu

s sy

lves

tris,

Pin

us m

ugo,

Pin

us n

igra

(at

leas

t 1

spec

ies

dom

ina

nt)

and

+/

spec

ies

of X

.8

Pinu

s sp

p.-v

eg.

Dd41

a,

41b

, 41

Ab,

41

A/a,

41

A2,

41A3

KGY,

W

ZK

KHp

KI

X.8.

3X

XX

NX

Q

uerc

us t

ype

Que

rcus

rob

ur (

dom

inan

t)

and

+/

Betu

la p

endu

la, B

etul

a pu

besc

ens,

Popu

lus t

rem

ula,

Sor

bus

aucu

paria

, Agr

ostis

cap

illar

is, A

ntho

xant

hum

odo

ratu

m, C

arex

are

-na

ria, D

ryop

teris

dila

tata

, Em

petr

um n

igru

m, H

olcu

s lan

atus

, Lon

icer

a pe

ricly

men

um

2180

Dd

42A1

, 42

A2,

42A

a,

42A

b,

42A

c,

43A1

, 43

Ab,

43

Bc,

43

Bd,

43

Be,

43

B2,

43B

b,

43B

c,

43B

d,

43d

KGQ

KH

qQ

R21

80

X.9

XX

X

Ope

n du

ne

area

s no

veg

etat

ion

as s

ur

roun

din

g N

atur

a 20

00

type

Dd

KD

OKD

OKD

oKD

76



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

X.10

XX

XX

X

Rude

ral d

une

area

s Ep

ilobi

um a

ngus

tifol

ium

, Cirs

ium

arv

ense

, Cirs

ium

vul

gare

, Urt

ica

dioi

ca, U

rtic

a ur

ens,

Rubu

s ca

esiu

s, An

chus

a of

ficin

alis,

Rub

us f

ru

ticos

us, L

inar

ia v

ulga

ris (

at le

ast

one

spec

ies

dom

inan

t)

and

+/

spec

ies

of X

.5

dry.

veg.

of U

rtic

a sp

p., E

pilo

bium

an-

gust

ifoliu

m

as s

ur

roun

din

g N

atur

a 20

00

type

Dd

,Rr

l14

/a,

14E

a,

14h

, 15

d,

15e

, 17

A2,

17A

a,

31e

, 33

a,

33A1

, 33

A4,

33A5

, 33

/a,

34A1

, 37

g

KDR

KDW

, KD

GKD

rEA

X.11

X

X

Salt

y du

ne a

reas

sp

ecie

s of

X.5

and

Sal

t m

arsh

spe

cies

an

d +/

Cl

adon

ia ra

ngifo

rmis

, Cla

doni

a fu

rcat

a ss

p. s

ubra

ngifo

rmis

Salty

dun

e ve

g.

(xer

oser

ie-h

alos

erie

)as

sur

ro

und

ing

Nat

ura

2000

ty

pe

R.

., C.

., X.

KD

F

KDn

DS

X.12

XN

NN

XX

Drif

ting

san

d du

ne

as s

ur

roun

din

g N

atur

a 20

00

type

(m

ost

ly

2120

)

KD

O

KDm

WD

2120

H.0

XX

XX

XX

Dune

sla

cks

(hum

id)

2190

Dv

KN

,KB

KDT

2190

H.1

X

Pion

eer

dune

sl

acks

2190

HS

Dv

KNP

KDT

KPy

PI21

90

H.1

.0

Pi

onee

r du

ne

slac

ks, u

nspe

cific

2190

HS*

Dv

KNP

KDT

PI

2190

H.1

.1X

XX

XX

Ce

ntau

rium

lit

tora

le t

ype

Cent

auriu

m li

ttor

ale,

Sag

ina

nodo

sa (a

t le

ast

one

spec

ies)

an

d +/

sp

ecie

s of

hyg

rose

rie: J

uncu

s anc

eps,

Agro

stis

stol

onife

ra, P

o-te

ntill

a an

serin

a, Ju

ncus

art

icul

atus

, Hol

cus l

anat

us, M

enth

a aq

uatic

a,

Linu

m c

atha

rtic

um, C

arex

flac

ca, B

ryum

alg

ovic

um

and

+/

spec

ies

of x

eros

erie

: Leo

ntod

on sa

xatil

is, C

arex

are

naria

, Lo

tus c

orni

cula

tus,

Hon

cken

ya p

eplo

ides

, Fes

tuca

rubr

a, A

mm

ophi

la

aren

aria

, Son

chus

arv

ensi

s, Se

dum

acr

e, H

ippo

phaë

rham

noid

es

and

+/

spec

ies

of h

alos

erie

: Pla

ntag

o co

rono

pus,

Plan

togo

mar

itim

a,

Arm

eria

mar

itim

a, Ju

ncus

ger

ardi

i, Od

ontit

es v

ernu

s, Gl

aux

mar

itim

a,

Care

x di

stan

s, Ca

rex

exte

nsa,

Trifo

lium

frag

iferu

m

Cent

aurio

-Sa

gine

tum

(sam

olet

osum

,pa

rnas

siot

osum

)

2190

HSs

Crs

27A2

, 27

A1KN

HKD

TKP

cCS

2190

77



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

H.1

.2X

XX

NX

Ra

diol

a lin

oide

s ty

pe

Radi

ola

linoi

des,

Anag

allis

min

ima,

Jun

cus

pygm

aeus

, Cic

endi

a fil

ifo

rmis

(at

leas

t on

e sp

ecie

s)

Junc

us b

ufon

ius,

Gnap

haliu

m u

ligin

osum

, Iso

lepi

s set

acea

, Ele

ocha

ris

quin

quefl

ora,

Cen

taur

ium

pul

chel

lum

, Sag

ina

proc

umbe

ns, C

arex

vi

ridul

a, P

lant

ago

maj

or, P

oa a

nnua

, Cal

lierg

onel

la c

uspi

data

, Dre

pano

-cl

adus

adu

ncus

, Cer

atod

on p

urpu

reus

, Bry

um sp

ec, P

ohlia

nut

ans,

Scap

ania

irrig

ua, F

osso

mbr

onia

fove

olat

a, Ju

nger

man

nia

grac

illim

a,

Ceph

aloz

ia b

icus

pida

ta

and

+/

spec

ies

of H

.1.4

/H.3

.1 or

+/

spe

cies

of H

.2/H

.2.1

or

+/

spe

cies

of S

.0/H

.1.1

Cice

ndie

tum

filif

or-

mis

, Iso

eto-

Nan

ojun

cete

a-ve

g.

2190

Dv

28A1

KNP

(KN

H)

KP

rRL

2190

H.1

.3X

XX

NX

Li

ttor

ella

uni

flora

ty

pe

Litt

orel

la u

niflo

ra, P

otam

oget

on p

olyg

onifo

lius,

Eleo

char

is m

ul

ticau

lis, D

esch

amps

ia s

etac

ea, A

pium

inun

datu

m, P

ilula

ria g

lobu

lif

era,

Sci

rpus

flui

tans

, Bal

delli

a ra

nunc

uloi

des,

Sam

olus

val

eran

di,

Pota

mog

eton

gra

min

eus,

Junc

us b

ulbo

sus,

Lyth

rum

por

tula

, Spa

rga

nium

min

imum

(at

leas

t on

e sp

ecie

s)

and

+/

Hyd

roco

tyle

vul

garis

, Ran

uncu

lus fl

amm

ula,

Car

ex v

iridu

la,

Junc

us a

rtic

ulat

us, J

uncu

s anc

eps,

Pote

ntill

a an

serin

a, E

leoc

haris

pa

lust

ris s

sp. p

alus

tris

+ u

nigl

umis

, Agr

ostis

stol

onife

ra, C

arex

nig

ra,

Phra

gmite

s aus

tral

is, D

repa

nocl

adus

spe

c., C

allie

rgon

ella

cus

pida

te

and

+/

(wet

situ

atio

n) U

tric

ular

ia s

pp. (

ofte

n U.

aus

tral

is),

Char

a sp

p. (o

ften

C. g

lobu

laris

), M

yrio

phyl

lum

spp

. (of

ten

M. s

pica

tum

), Po

tam

oget

on s

pp. (

ofte

n P.

nat

ans)

, Ran

uncu

lus p

elta

tus

ssp.

bau

dotii

, H

ippu

ris v

ulga

ris, P

olyg

onum

am

phib

ium

an

d +/

(b

asic

situ

atio

n) M

enth

a aq

uatic

a, C

arex

flac

ca, C

ampy

lium

po

lyga

mum

, and

rare

: Sco

rpid

ium

scor

pioi

des,

Drep

anoc

ladu

s sen

dt-

neri,

Sco

rpid

ium

revo

lven

s

Litt

orel

lete

a un

iflo

rae-

veg.

2190

Dv

06a

, 06

C4KN

P (K

NKn

, KN

T)

KDT

KPl

LU21

90

H.1

.4X

XX

NX

Ly

copo

diel

la

inun

data

typ

e Ly

copo

diel

la in

unda

ta, R

hync

hosp

ora

alba

, Rhy

ncho

spor

a fu

sca,

Dr

oser

a in

term

edia

(at

leas

t on

e sp

ecie

s, an

d ra

re H

amm

arby

apa

ludo

sa)

and

+/

spec

ies

of H

.3.1

: Eric

a te

tral

ix, D

rose

ra ro

tund

ifolia

, Vac

ci-

nium

ulig

inos

um, E

mpe

trum

nig

rum

, Nar

thec

ium

oss

ifrag

um, D

rose

ra

rotu

ndifo

lia, O

xyco

ccus

pal

ustr

is, G

entia

na p

neum

onan

the,

Mol

inia

ca

erul

ea, H

ydro

coty

le v

ulga

ris, C

arex

nig

ra, C

arex

pan

icea

, Erio

phor

um

angu

stifo

lium

, Jun

cus b

ulbo

sus,

Salix

repe

ns, P

olyt

richu

m c

omm

une,

Di

cran

um sc

opar

ium

, Sph

agnu

m d

entic

ulat

um, D

repa

nocl

adus

exa

n-nu

latu

s an

d ot

her D

repa

nocl

adus

, Sph

agnu

m s

pec.

an

d +/

Li

verw

orts

: Gym

noco

lea

infla

ta, S

capa

nia

irrig

ua, F

osso

mbr

o-ni

a fo

veol

ata,

Jung

erm

anni

a gr

acill

ima,

Cep

halo

zia

bicu

spid

ata

Lyco

podi

ella

inun

-da

ta-v

eg.,

Spha

gno-

Rhyn

chos

pore

tum

, Ly

copo

dio-

Rhyn

cho-

spor

etum

2190

Dv

11A1

KNE

KDT

KPi

LI21

90

78



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

H.2

Dune

sla

ck f

ens

spec

ies

of H

.2: S

alix

repe

ns a

gg.,

Hyd

roco

tyle

vul

garis

, Ran

uncu

lus

flam

mul

a, A

gros

tis st

olon

ifera

, Jun

cus a

rtic

ulat

us, J

uncu

s anc

eps,

Eleo

char

is p

alus

tris

ssp

. pal

ustr

is +

uni

glum

is, P

hrag

mite

s aus

tral

is,

Pote

ntill

a an

serin

a, C

allie

rgon

ella

cus

pida

ta

Sche

uchz

erio

-Car

i-ce

tea

fusc

ae-v

eg.

2190

HS

DvKN

KDT

CN

2190

H.2

.0

Du

ne s

lack

fen

s, un

spec

ific

2190

Dv

09/a

KNS

KDT

CN

2190

H.2

.1X

XX

NX

Ca

rex

trin

ervi

s ty

pe

Care

x ni

gra,

Car

ex t

riner

vis,

Pote

ntill

a pa

lust

ris (

at le

ast

one

spec

ies

dom

inan

t)

and

+/

Salix

repe

ns a

gg. (

ofte

n do

min

ant)

, Cal

amag

rost

is c

anes

cens

, Er

ioph

orum

ang

ustif

oliu

m, E

pilo

bium

pal

ustr

e, S

tella

ria p

alus

tre,

Gal

i-um

pal

ustr

e, V

iola

pal

ustr

is, M

enya

nthe

s trif

olia

ta, L

ycop

us e

urop

aeus

, Dr

epan

ocla

dus a

dunc

us, C

allie

rgon

cor

difo

lium

, Sph

agnu

m fi

mbr

iatu

m

and

+/

spec

ies

of H

.2

Caric

etum

trin

ervi

-ni

grae

, Car

icio

n ni

grae

veg

.

2190

Dv

09A1

, 09

A3,

09A

a,

09a

, 09

b,

09c

, 09

e,

09f

, 09

h,

09B

b,

09/c

KNA

KDT

KMm

CN21

90

H.2

.2X

XX

NN

Sc

hoen

us

nigr

ican

s ty

pe

Scho

enus

nig

rican

s, Li

paris

loes

elii,

Par

nass

ia p

alus

tris,

Epi

pact

is pa

lust

ris, D

acty

lorh

iza

inca

rnat

a, P

ellia

end

iviif

olia

, Cam

pyliu

m p

oly

gam

um, C

. ste

llatu

m, B

ryum

pse

udot

rique

trum

, and

rar

e:

Gent

iane

lla a

mar

ella

, Car

ex p

ulic

aris,

Jun

cus

subn

odul

osus

, Tar

axa

cum

pal

ustr

e, F

issid

ens

adia

ntho

ides

, Ane

ura

ping

uis,

Prei

ssia

qu

adra

ta, M

oerc

kia

hibe

rnic

a (m

inim

um 3

spe

cies

) an

d +/

M

enth

a aq

uatic

a, E

leoc

haris

qui

nque

flora

, Car

ex fl

acca

, Lin

um

cath

artic

um, a

nd ra

re: E

quis

etum

var

iega

tum

, Ped

icul

aris

pal

ustr

is,

Junc

us a

rctic

us ss

p. b

altic

us, G

ymna

deni

a co

nops

ea, A

naga

llis t

enel

la

and

+/

spec

ies

of H

.2

and

+/

spec

ies

of H

.1.1

/H.1

.2

Junc

o ba

ltici

-Sc

hoen

etum

ni

gric

antis

, Jun

cus

subn

odul

osus

-veg

., Pa

rnas

sio-

Junc

etum

at

ricap

illi,

Caric

ion

dava

llian

ae v

eg.

2190

HSc

Dv,

Dvs,

Dvp

09C3

KNK

KDT

???

21

90

H.2

.3

XX

N

Ca

lam

agro

stis

epig

ejos

typ

e Ca

lam

agro

stis

epig

ejos

(do

min

ant)

, Sal

ix r

epen

s ag

g., (

and

rare

O

phio

glos

sum

vul

gatu

m),

and

+/

Care

x ni

gra,

Phr

agm

ites a

ustr

alis

, Lyc

opus

eur

opae

us, H

olcu

s la

natu

s, Ci

rsiu

m p

alus

tre,

Lat

hyru

s pra

tens

is, P

oa p

rate

nsis

, Gal

ium

pa-

lust

re, H

ydro

coty

le v

ulga

ris, A

gros

tis st

olon

ifera

, Ran

uncu

lus fl

amm

ula,

Ca

llier

gone

lla c

uspi

data

, Bra

chyt

heci

um ru

tabu

lum

Cala

mag

rost

is

epig

ejos

-veg

.21

90

Dv09

d,

09/c

KNS

KDT

2190

H.3

N

Dune

sla

ck h

eath

Oxyc

occo

-Sph

ag-

nete

a-ve

g.21

90

DvKN

EKD

T

ET21

90

H.3

.0

N

Du

ne s

lack

hea

th,

unsp

ecifi

c21

90

Dv

KNE

KDT

ET

2190

79



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

H.3

.1X

XX

NX

Er

ica

tetr

alix

ty

pe

Eric

a te

tral

ix (

dom

inan

t)

and

+/

Empe

trum

nig

rum

, Nar

thec

ium

oss

ifrag

um, V

acci

nium

ulig

i-no

sum

, Vac

cini

um o

xyco

ccos

, Vac

cini

um m

acro

carp

os, D

rose

ra ro

tun-

difo

lia, J

uncu

s squ

arro

sus,

Gent

iana

pne

umon

anth

e, S

cirp

us c

espi

tosu

s ss

p. g

erm

anic

us, A

ulac

omni

um p

alus

tre,

Pol

ytric

hum

com

mun

e, S

phag

-nu

m fi

mbr

iatu

m, S

. sub

nite

ns, S

. pal

ustr

e, S

. fal

lax,

S. s

quar

rosu

m, S

. m

olle

an

d +/

Sa

lix re

pens

agg

., Ca

lluna

vul

garis

, Pot

entil

la e

rect

a, V

iola

pa

lust

ris, C

arex

trin

ervi

s, Ca

rex

pani

cea,

Agr

ostis

can

ina,

Erio

phor

um

angu

stifo

lium

, Ped

icul

aris

sylv

atic

a, D

anth

onia

dec

umbe

ns, M

olin

ia

caer

ulea

, Nar

dus s

tric

ta, H

ypnu

m ju

tland

icum

/ cu

pres

sifo

rme,

Ple

uro-

zium

schr

eber

i, Di

cran

um sc

opar

ium

, Pse

udos

cler

opod

ium

pur

um

Empe

tro-

Eric

etum

, N

arth

eciu

m o

ssifr

a-gu

m-v

eg.

2190

Dv

11A3

KNE

KDT

KMt

ET21

90

H.3

.2

X

N

O

xyco

ccus

m

acro

carp

os t

ype

Oxy

cocc

us m

acro

carp

os (

dom

inan

t)

and

+/

spec

ies

of H

.3.1

/H.2

.1

Oxyc

occu

s mac

ro-

carp

os-v

eg.

2190

Dv

11b

KNE

KDT

ET

2190

H.3

.3

X

N

M

olin

ia c

aeru

lea

type

M

olin

ia c

aeru

lea

(dom

inan

t)

and

+/

spec

ies

of H

.3.1

/H.2

.1

Mol

inia

cae

rule

a-ve

g.21

90

Dv10

e,

11c

KNE,

KN

AKD

T

ET21

90

H.4

Dune

sla

ck

reed

bed

Phra

gmite

tea-

veg.

2190

Dv

08B3

KNR

KDT

R

2190

H.4

.0

Du

ne s

lack

re

edbe

d,

unsp

ecifi

c

2190

Dv

08d

, 08

e,

08f

, 08

g,

08h

, 08

k,

32c

, 32

f

KNR

KDT

R

2190

H.4

.1X

XX

XX

Ph

ragm

ites

aus

tral

is ty

pe

Phra

gmite

s au

stra

lis (

dom

inan

t)

and

+/

Iris p

seud

acor

us, T

ypha

ang

ustif

olia

, Typ

ha la

tifol

ia, S

choe

no-

plec

tus t

aber

naem

onta

ni

and

+/

spec

ies

of H

.2

Scirp

o-Ph

ragm

ite-

tum

, Sch

oeno

plec

to-

Phra

gmite

tum

, Ty-

pho-

Phra

gmite

tum

2190

Dv

08B2

, 08

B3,

26b

KNR

KDT

KMr

R21

90

H.4

.2

XX

N

Ca

rex

spp.

typ

e Ca

rex

pseu

docy

peru

s, Ca

rex

ripar

ia, C

arex

acu

ta, C

arex

ova

lis, C

la

dium

mar

iscus

, Car

ex s

pp. (

at le

ast

1 sp

ecie

s do

min

ant)

an

d +/

sp

ecie

s of

H.4

.1/H

.2

Mag

noca

ricio

n-ve

g.21

90

Dv08

C1,

08C2

, 08

C6,

08C

a,

08C

b,

08C

c,

08C

d,

08C

e,

08C

f

KNA

KDT

2190

H.5

X

Dune

sla

ck

will

ow s

hrub

bery

2190

Dv

36A2

KD

T

SA21

70

80



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

H.5

.0

X

Du

ne s

lack

w

illow

shr

ubbe

ry,

unsp

ecifi

c

2190

(L

S 21

70)

Dv

09/a

KBK

KDT

SA

2170

H.5

.1X

XX

XX

Sa

lix c

iner

ea t

ype

Salix

cin

erea

(do

min

ant)

, Sal

ix r

epen

s ag

g.

and

+/

spec

ies

of H

.2/H

.4/H

.6

Salix

cin

erea

-Sal

ix

aren

aria

-veg

., Sa

li-ce

tum

cin

erea

e –

sa-

licet

osum

repe

ntis

2190

(N

L,DK

21

70)

Dv

36A2

KBH

KDT

KMw

SA21

70

H.5

.2X

XX

NN

NM

yric

a ga

le t

ype

Myr

ica

gale

(do

min

ant)

, Sal

ix r

epen

s ag

g.

and

+/

spec

ies

of H

.2/H

.4/H

.6

Myr

icet

um g

alis

2190

(N

L,DK 21

70)

Dv

09h

, 36

Ab

KBH

KDT

2170

H.6

X

Dune

sla

ck

woo

dlan

d21

80

Dv

KD

T

BP, A

G21

80

H.6

.0

Du

ne s

lack

w

oodl

and,

un

spec

ific

2190

Dv

KB

SKD

T

BP, A

G21

80

H.6

.1X

XX

NX

Be

tula

pub

esce

ns

type

Be

tula

pub

esce

ns (

dom

inan

t)

and

+/

Care

x ni

gra,

Cal

amag

rost

is c

anes

cens

, Hyd

roco

tyle

vul

garis

, M

enth

a aq

uatic

a, P

hrag

mite

s aus

tral

is, E

rica

tetr

alix

, Sal

ix re

pens

agg

., Sa

lix c

iner

ea, C

allie

rgon

ella

cus

pida

ta, S

phag

num

spp

. an

d +/

Ca

lam

agro

stis

epi

gejo

s, Ca

rex

aren

aria

, Lon

icer

a pe

ricly

me-

num

, Agr

ostis

cap

illar

is, E

mpe

trum

nig

rum

, Dic

ranu

m sc

opar

ium

, Rub

us

spec

.

Betu

la p

ubes

cens

-ve

g., E

mpe

tro-

Betu

-le

tum

car

patic

ae

2180

Dv

40A2

, 40

Ac,

40

Ad

KBA,

KB

RKD

TKM

bBP

2180

H.6

.2X

XX

NX

Al

nus

glut

inos

a ty

pe

Alnu

s gl

utin

osa

(dom

inan

t)

and

+/

Care

x ni

gra,

Cal

amag

rost

is c

anes

cens

, Hyd

roco

tyle

vul

garis

, M

enth

a aq

uatic

a, P

hrag

mite

s aus

tral

is, E

rica

tetr

alix

, Sal

ix re

pens

agg

., Sa

lix c

iner

ea, C

allie

rgon

ella

cus

pida

ta, S

phag

num

spe

c.

and

+/

Cala

mag

rost

is e

pige

jos,

Care

x ar

enar

ia, L

onic

era

peric

lym

e-nu

m, A

gros

tis c

apill

aris

, Em

petr

um n

igru

m, R

ibes

nig

rum

, Sol

anum

dul

-ca

mar

a, D

icra

num

scop

ariu

m, R

ubus

spec

., So

lanu

m d

ulca

mar

a, G

lyce

-ria

max

ima,

Iris

, pse

udac

orus

, Thy

pha

spec

., Ca

rex

ripar

ia, P

hrag

mite

s

Alnu

s glu

tinos

a-ve

g.21

80

Dv39

A1,

39A2

, 39

Aa,

39

Ab,

39

Ad,

39

Ae,

39

Af

KBE

KDT

KMe

AG21

80

H.7

X

Ope

n du

ne s

lack

ar

eas

no v

eget

atio

n21

90

Dv

KNP

KDT

KMy

OD

2190

H.8

XX

XN

X

Aqua

tic

vege

tati

on in

du

ne s

lack

s

Hyd

rose

rie

Utric

ular

ia s

pp. (

ofte

n U.

aus

tral

is), C

hara

spp

. (of

ten

C. g

lobu

la

ris),

Myr

ioph

yllu

m s

pp. (

ofte

n M

. spi

catu

m),

Pota

mog

eton

spp

. (o

ften

P. n

atan

s), R

uppi

a sp

p., R

anun

culu

s pe

ltatu

s ss

p. b

audo

tii

(at

leas

t on

e sp

ecie

s do

min

ant)

Char

etea

frag

ilis,

Pota

mog

eton

eta,

Ut

ricul

arie

tea-

veg.

2190

Dv

04a

, 04

B2,

04B3

, 04

C1,

04C3

, 04

D1a,

04

D1b,

05

A1,

05A2

, 05

Aa,

05

E3,

05f

, 05

h

KNT

KDT

KMf

HY

2190

81



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

H.9

XX

XX

X

Rude

ral d

une

slac

k ar

eas

wit

h lit

ter

brus

hwoo

d

Epilo

bium

hirs

utum

, Cirs

ium

vul

gare

+ a

rven

se (

at le

ast

one

spe

cies

dom

inan

t)

and

+/

spec

ies

of H

.2/H

.4

Moi

st v

eget

atio

n of

Ep

ilobi

um h

ir sut

um,

Cirs

ium

vul

gare

a.

arve

nse

2190

Dv

32B2

, 32

B3,

32A1

, 32

a,

32b

, 32

c,

32d

, 32

e,

32f

, 32

g

KNS

KDT

KMh

EH21

90

S

Sa

lt m

arsh

KH

S.0

No

vege

tati

on

Su

KN

o88

S.0.

0X

XX

XX

Ve

geta

tion

not

pr

esen

t, di

stur

bed

as s

ur

roun

din

g N

atur

a 20

00

type

Su*

50

A,

50B,

50

C

KW

OKN

x90

S.0.

1X

XX

XX

O

pen

wat

er

as s

ur

roun

din

g N

atur

a 20

00

type

Sww

ater

KL

A,KL

Z

KNt

55

S.0.

2X

XX

XX

Ba

re s

and

or

shel

ls in

sal

t m

arsh

sys

tem

s

as s

ur

roun

din

g N

atur

a 20

00

type

Ssza

nd;

sche

lp

(KS)

KWO

KNo

88

S.0.

3X

XX

XX

Ba

re m

ud o

r cl

ay

in s

alt

mar

sh

syst

ems

as s

ur

roun

din

g N

atur

a 20

00

type

Smsl

ik,

Qq0

p,

Qq0

e,

QuO

, Ss

0

(K

WK,

KW

B)KW

OKW

vVO

S.1

Pion

eer

zone

sp

ecie

s of

S.1

: Spa

rtin

a an

glic

a, S

alic

orni

a sp

ec.,

Suae

da m

ariti

ma

SP

KW

S.1.

0

Pion

eer

zone

, un

spec

ific

SP*

KW

S.1.

1X

XX

XX

XSp

artin

a an

glic

a ty

peSp

artin

a an

glic

aSp

artin

etum

angl

icae

1320

SPs

Ss5,

Ss

3,

Bs3,

Bs

5

24A1

KWG

KWG

KNs

S13

20

82



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

S.1.

2X

XX

XX

XSa

licor

nia

type

Sa

licor

nia

spp.

, Sua

eda

mar

itim

a, B

assia

hirs

uta

(at

leas

t on

e sp

eci

es d

omin

ant)

Salic

orni

etum

stric

-ta

e, S

alic

orni

etum

ra

mm

osis

sim

ae,

Salic

orni

etum

de

cum

bent

is, S

ua-

edet

um m

ariti

mae

, Th

ero-

Salic

orni

on

stric

tae-

veg.

, Su

aedo

-Bas

siet

um

hirs

utae

1310

SPq

Qqp

, Q

qe,

Qu

25A2

, 25

Aa,

26

Ac

KWQ,

KW

QW,

KWQV

, KS

F

KWQ

KQb,

KQ

d,

KQr,

KQs,

KQu

SS, S

R,

SD, U

, BH

1310

S.2

X

Low

mar

sh

spec

ies

of S

.2: P

ucci

nelli

a m

ariti

ma,

Atr

iple

x po

rtul

acoi

des,

Coch

lear

ia

angl

ica,

Ast

er tr

ipol

ium

, Spe

rgul

aria

mar

itim

a +

salin

a, T

riglo

chin

m

ariti

ma,

Lim

oniu

m v

ulga

re, P

lant

ago

mar

itim

a, P

arap

holis

strig

osa,

At

riple

x pe

dunc

olat

a

Pucc

inel

lion

mar

itim

ae-v

eg.

1330

SL

KH

U,

KHUS

, KH

UB,

KHU,

KH

W

13

30

S.2.

0

X

Lo

w m

arsh

, un

spec

ific

1330

SL*

Pw

KHUZ

KHU,

KH

W

KU13

30

S.2.

1X

XX

XX

Pu

ccin

ellia

mar

itim

a ty

pe

Pucc

inel

lia m

ariti

ma

(dom

inan

t)

and

+/

Sper

gula

ria s

pp.,

Salic

orni

a sp

p., P

ucci

nelli

a di

stan

s, Sp

artin

a an

glic

a, S

uaed

a m

ariti

ma,

Lim

oniu

m v

ulga

re, A

ster

trip

oliu

m

Pucc

inel

lietu

mm

ariti

mae

1330

SLp

P, P

p,

Ppq,

P

d,

Pq,

P

u,

Pps,

Ppu,

Pe

, Pt,

Pex,

Pj

, Pg

26A1

, 26

Aa

KHUP

KHU,

KH

WKN

d,KN

p,KN

v,KN

w

P, P

s13

30

S.2.

2X

XX

XX

Li

mon

ium

vu

lgar

e / P

ucci

ne

llia

mar

itim

a ty

pe

Lim

oniu

m v

ulga

re

and

+/

Plan

tago

mar

itim

a, P

ucci

nelli

a m

ariti

ma,

Spe

rgul

aria

mar

iti-

ma,

Ast

er tr

ipol

ium

(Pla

ntag

ini-

Lim

onie

tum

)13

30SL

lPp

l, Pl

26A2

KHUL

KHU,

KH

W

P13

30

S.2.

3X

XX

XX

As

ter

trip

oliu

m /

Pucc

inel

lia

mar

itim

a ty

pe

Aste

r tr

ipol

ium

(do

min

ant)

an

d +/

Pu

ccin

ellia

mar

itim

a, S

perg

ular

ia m

ariti

ma,

Lim

oniu

m v

ulga

-re

, Pla

ntag

o m

ariti

ma

Aste

r trip

oliu

m-v

eg.

1330

Sla

Ppa,

Pa

, Jf

a,

Jja,

Pp

ab,

Ba

26A

bKH

UAKH

U,

KHW

KNa

A13

30

S.2.

4X

XX

XX

At

riple

x po

rtul

acoi

des

type

Atrip

lex

port

ulac

oide

s (d

omin

ant)

an

d +/

Pu

ccin

ellia

mar

itim

a, S

perg

ular

ia m

ariti

ma,

Sal

icor

nia

spp.

, Su

aeda

mar

itim

a

Hal

imio

netu

m p

or-

tula

coid

is13

30SL

hPh

26A3

KHUH

KHU,

KH

WKN

hH

1330

S.3

X

Hig

h m

arsh

sp

ecie

s of

S.3

: Art

emis

ia m

ariti

ma,

Arm

eria

mar

itim

a, Ju

ncus

ger

ardi

i, Gl

aux

mar

itim

a, F

estu

ca ru

bra,

Ely

mus

ath

eric

us, J

uncu

s mar

itim

us,

Trifo

lium

frag

iferu

m, P

lant

ago

coro

nopu

s, Sa

gina

mar

itim

a, C

arex

ex

tens

a, C

arex

dis

tans

, Ono

nis s

pino

sa, H

orde

um se

calin

um, C

ochl

earia

da

nica

, Odo

ntite

s ver

nus s

sp se

rotin

us, E

lym

us re

pens

Arm

erio

n m

ariti

mae

-veg

.13

30SH

KHO

KHO

, KH

IKO

y

1330

S.3.

0

X

H

igh

mar

sh,

unsp

ecifi

c13

30SH

*Jw

KH

OZ

KHO

, KH

I

KO13

30

83



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

S.3.

1X

XX

XX

Li

mon

ium

vul

gare

/ J

uncu

s ge

rard

ii ty

pe

Lim

oniu

m v

ulga

re (

dom

inan

t), J

uncu

s ge

rard

i an

d +/

sp

ecie

s of

S.3

(Pla

ntag

ini-

Lim

oni-

etum

)13

30SH

lJj

l, Jfl

26A2

, 26

Ca,

26

a

KHO

LKH

O

13

30

S.3.

2X

XX

XX

Ju

ncus

ger

ardi

i ty

pe

Junc

us g

erar

dii

Glau

x m

ariti

ma

an

d +/

sp

ecie

s of

S.3

Junc

etum

ger

ardi

i13

30SH

jJj

, Jex

26C1

, 26

Cc

KHO

JKH

O,

KHI

KOj

J13

30

S.3.

3X

XX

XX

Fe

stuc

a ru

bra

type

Fe

stuc

a ru

bra

(dom

inan

t)

and

+/

spec

ies

of S

.3

Fest

uca

rubr

a-ve

g.,

Arm

erio

-Fes

tcet

um

litto

ralis

1330

SHf

Jf26

Cb

KHO

RKH

O,

KHI

KOf

F13

30

S.3.

4

X

At

riple

x po

rtul

acoi

des

/ Ar

tem

isia

mar

itim

a ty

pe

Atrip

lex

port

ulac

oide

s (d

omin

ant)

an

d +/

Ar

tem

isia

mar

itim

a, F

estu

ca ru

bra,

Junc

us g

erar

dii,

Elym

us

athe

ricus

Hal

imio

netu

mpo

rtul

acoi

dis

1330

SHh

Jfh

26A3

KHUH

KHO

13

30

S.3.

5X

XX

XX

Ar

tem

isia

mar

itim

a /

Fest

uca

rubr

a ty

pe

Arte

misi

a m

ariti

ma

(dom

inan

t)

and

+/

Fest

uca

rubr

a, E

lym

us a

ther

icus

Arte

mis

ietu

m

mar

itim

ae13

30SH

zPz

, Jf

z,

Jz

26c

KHO

BKH

O,

KHI

KOt

T13

30

S.3.

6X

XX

XX

Ju

ncus

mar

itim

us

type

Ju

ncus

mar

itim

us (

dom

inan

t)

and

+/

Fest

uca

rubr

a, Ju

ncus

ger

ardi

i, El

ymus

ath

eric

us

Junc

us m

ariti

mus

-ve

g.13

30SH

mPm

, Jm

26e

KHBN

KHO

, KH

BKO

mM

G13

30

S.3.

7X

XX

XX

El

ymus

ath

eric

us

type

El

ymus

ath

eric

us (

dom

inan

t)

and

+/

Fest

uca

rubr

a, A

trip

lex

pros

trat

a sp

ecie

s of

S.3

Agro

pyre

tum

lito

-ra

lis, A

grop

yret

um

pung

entis

, Atr

ipl-

ici-

Agro

pyre

tum

pu

ngen

tis, A

trip

lici-

Ely m

etum

ath

eric

i, El

ymet

um p

ycna

nthi

, As

tero

-Agr

opyr

etum

re

pent

is, E

lym

us

repe

ns s

sp. l

ittor

alis

-ve

g.

1330

SHy

Py,

Jy3,

Jy

5,

Ry3,

Ry

5,

By3,

By

5

26d

KHQ

AKH

QKO

qQ

1330

S.3.

8X

XX

XX

Ca

rex

exte

nsa

type

Ca

rex

exte

nsa

and

+/

Odon

tites

ver

nus

ssp

sero

tinus

, Cen

taur

ium

pul

chel

lum

, Gla

ux

mar

itim

a, Ju

ncus

mar

itim

us, A

gros

tis st

olon

ifera

, Par

apho

lis st

rigos

a an

d +/

sp

ecie

s of

S.3

Junc

o Ca

ricet

um

exte

nsae

1330

SHe

Je26

C2KH

S,

KHBN

KHB

KOc

CE13

30

S.3.

9X

XX

XX

At

riple

x pr

ostr

ata

/ A.

litto

ralis

typ

e

Atrip

lex

pros

trat

a, A

. litt

oral

is (a

t le

ast

one

spec

ies

dom

inan

t)

and

+/

Trip

leur

ospe

rmum

mar

itim

um

Atrip

lex

pros

trat

a /

litto

ralis

veg

.13

30SH

xXx

, Xk

22A1

, 29

A3KH

MKH

OKO

l,KO

pAP

, AX

1330

S.3.

10X

XX

XX

Ag

rost

is st

olon

ifera

/ Tr

ifoliu

m

frag

iferu

m t

ype

Agro

stis

stol

onife

ra

and

+/

Pote

ntill

a an

serin

a, F

estu

ca ru

bra,

Trifo

lium

frag

iferu

m, T

rifo-

lium

repe

ns, L

otus

cor

nicu

latu

s an

d +/

sp

ecie

s of

S.3

Trifo

lio fr

agife

ri-Ag

rost

ietu

m st

o-lo

nife

rae,

Agr

ostis

st

olon

ifera

-veg

.

1330

SHg

Jg,

Rgf,

Rpf,

Rg

c,

Rgl,

Rgt

12B2

, 12

B3,

12B

m

KHB

or

KHF

KHI,

KHO

KOa

W13

30

84



TMAP

Co

deSt

an

dard

TM

AP

Appl

ied

in c

ount

ry/

fede

ral s

tate

TMAP

Ty

pe

Typi

cal /

cha

ract

eris

tic

spe

cies

veg

etat

ion

Nat

ura

2000

Co

de

Map

pi

ng

Code

N

L

RWS

SALT

Co

de

NL

Staa

ts

bosb

. Co

de

NL

Bio

tope

Co

de

LS

Bio

tope

Co

de

HH

Bio

tope

Co

de

SH

Map

pi

ng

Code

SH

Map

pi

ng

Code

DK

N LL S

H HS H

D K

S.3.

11

X

XX

Pl

anta

go

coro

nopu

s /

Cent

auriu

m

litto

rale

typ

e

Plan

tago

cor

onop

us

and

+/

Sagi

na m

ariti

ma,

Cen

taur

ium

litt

oral

e, S

agin

a no

dosa

, Coc

h-le

aria

dan

ica,

Fes

tuca

rubr

a, B

uple

urum

tenu

issi

mum

, Sed

um a

cre,

Br

achy

heci

um a

lbic

ans,

Leon

todo

n sa

xatil

e, B

ryum

spp.

an

d +/

sp

ecie

s of

S.3

Sagi

no-C

ochl

ea-

rietu

m

and

Cent

aurio

-Sag

i-ne

tum

trifo

lieto

sum

1330

(in

N

L 13

10)

SHc

Cc,

Ccj,

Ccs,

Crt

27Aa

1,27

Aa2

KHBT

KHS

KOs

SC13

30

S.3.

12X

XX

XX

O

noni

s sp

inos

a / C

arex

dist

ans

type

Care

x di

stan

s, O

noni

s sp

inos

a (a

t le

ast

one

spec

ies)

an

d +/

sp

ecie

s of

S.3

an

d +/

sp

ecie

s of

X.3

/X.5

Onon

ido-

Caric

etum

di

stan

tis13

30SH

oRo

12B4

KHBT

KHB

KOo

OC

1330

S.3.

13X

XX

XX

El

ymus

rep

ens

type

El

ymus

rep

ens

(dom

inan

t)

and

+/

spec

ies

of S

.3/X

.5

Agro

pyre

tum

lito

-ra

lis, A

grop

yret

um

pung

entis

, Atr

ipl-

ici-

Agro

pyre

tum

pu

ngen

tis, A

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72


7 References

Bakker, J.P., J. Bunje, K.S. Dijkema, J. Frikke, B. Kers, P. Körber, J. Kohlus, & M. Stock. 2005. Salt marshes. In: K. Essink, C. Dettmann, H. Farke, K. Laursen, G. Lüerßen, H. Marencic & W. Wiersinga (eds). Wadden Sea Quality Sta-tus Report 2004, Wadden Sea Ecosystem No. 19, Trilater-al Monitoring and Assessment Group, Common Wadden Sea Secretariat. Wilhelmshaven. p. 163-179.

BFN (2000, Hrsg.): Verzeichnis und Rote Liste der Pflanz-engesellschaften Deutsch lands (Bearb. E. Rennwald). - Schriftenreihe für Vegetationskunde: 35, 800 S., Bonn – Bad Godesberg.

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Dierssen, K. (1988): Rote Liste der Pflanzengesellschaf-ten Schleswig-Holsteins. - Schriftenreihe Landesamt für Naturschutz und Landschaftspflege Schleswig-Holstein 6: 157 S., Kiel.

Dijkema, K. & Wolff, W. (1983): Flora and vegetation of the Wadden Sea islands and coastal areas. Report 9 of the Wadden Sea Working Group: 413 S. + maps, Rot-terdam.

Drachenfels, O. v. (2011): Kartierschlüssel für Biotoptypen in Niedersachsen. - Naturschutz und Landschaftspflege in Niedersachsen A/4, Hildesheim.

Drachenfels, O. v. (2012): Hinweise zur Definition und Kartierung der Lebensraumtypen von Anh. I der FFH-Richtlinie in Niedersachsen auf der Grundlage des Inter-pretation Manuals der Europäischen Kommission (Ver-sion EUR 27 vom April 2007) + Anhang: Hinweise und Tabellen zur Bewertung des Erhaltungszustands der FFH-Lebensraumtypen in Niedersachsen. - Hannover.

Esselink, P., Petersen, J., Arens, S., Bakker, J.P., Bunje, J., Dijkema, K.S., Hecker, N., Hellwig, U., Jensen, A.V., Kers, A.S., Körber, P., Lammerts, E.J., Stock, M., Veeneklaas, Vreeken M. & Wolters, M. (2009): Salt Marshes. Thematic Report No. 8. In: Marencic, H. & VLAS, J. de (Eds), 2009. Quality Status Report 2009. Wadden Sea Ecosystem No. 25. Common Wadden Sea Secretariat, Trilateral Monitor-ing and Assessment Group, Wilhelmshaven, Germany.

European Commission, DG Environment EUR 27 (2007): Interpretation Manual of European Union Habitats. – 144 S., Brüssel.

Garve, E. (2004): Rote Liste und Florenliste der Farn- und Blütenpflanzen in Nieder sachsen und Bremen. 5. Fassung, Stand 1.3.2004 – Informationsdienst Natur schutz Nied-ersachsen 24 (1/2004): 1-76 u. Anlage: 1-8. Hildesheim.

Hauck, M. & U. de Bruyn (2010): Rote Liste und Gesa-mtartenliste der Flechten in Niedersachsen und Bremen. - Informationsd. Naturschutz Niedersachsen 1/2010: 83 S., Hannover.

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Koperski, M. (2011): Rote Liste und Gesamtartenliste der Moose in Niedersach-sen und Bremen. - Informationsd. Naturschutz Niedersachsen 3/2011: 208 S., Hannover.

Lammerts, E.J., Petersen, J. & Hochkirch, A. (2009): Beaches and Dunes. Thematic Report No. 15. In: MAREN-CIC, H. & VLAS, J. de (Eds), 2009. Quality Status Report 2009. Wadden Sea Ecosystem No. 25. Common Wadden Sea Secretariat, Trilateral Monitoring and Assessment Group, Wilhelmshaven, Germany.

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Petersen, J. (2011): Orchideengrünland der Ostfriesischen Inseln. Arbeitskreis Heimische Orchideen Niedersachsen e. V., 30 Jahre AHO-Niedersachsen, 95-111, Hannover.

Petersen, J. & Lammerts, E.J. (2005): Dunes. Chapter 9.2, Wadden Sea Quality Status Report. Common Wadden Sea Secretariat, Trilateral Monitoring and Assessment Group, Wilhelmshaven. 249-266.

Petersen, J. & Pott, R. (2005): Ostfriesische Inseln. Land-schaft und Vegetation im Wandel. - 160 S. u. Kartenband (12 Vegetationskarten auf DIN A1), Hrsg: Nds. Heimat-bund, Hannover.

Pott, R. (1995): Die Pflanzengesellschaften Deutschlands. - 622 S., Stuttgart.

Preising, E., Vahle, H.-C., Brandes, D., Hofmeister, H., Tüxen, J. & Weber, H. (1990): Die Pflanzengesellschaf-ten Niedersachsens - Bestands-entwicklung, Gefährdung und Schutzprobleme - Wasser- und Sumpfpflanzenge-sellschaften des Süßwassers. - Naturschutz u. Land-schaftspfl. Niedersachsen 20 (8): 161 S., Hannover.

Ringot, J.-L. (1992): Luftbildinterpretation, Nationalpark Niedersächsisches Wattenmeer. Biotop- und Nutzung-stypenkartierung der ostfriesischen Inseln und des Fest-landsküstenvorlandes. Unveröff. Bericht im Auftrag der NLPV Wilhelmshaven.

Schaminée, J., Weeda, E. & Westhoff, V. (1995): De veg-etatie van Nederland 2. Plantengemeenschappen van wateren, moerassen en natte heiden - :358 S., Uppsala, Leiden.

Schaminée, J., Stortelder, A. & Weeda, E. (1996): De Veg-etatie van Nederland 3. Plantengemeenschappen van graslanden, zomen en droge heiden. - :356 S., Uppsala, Leiden.

Schaminée, J., Weeda, E. & Westhoff, V. (1998): De Veg-etatie van Nederland 4. Plantengemeenschappen van de kust en binnenlandse pionier-milieus. - :346 S., Uppsala, Leiden.

Schipper, P.C., (2002): Catalogus vegetatietypen. In: Staatsbosbeheer: Catalogi bedrijfssturing: natuur, bos, recreatie en landschap. Staatsbosbeheer, afdeling terre-inbeheer, Driebergen.

Stock, M., S. Gettner, H. Hagge, K. Heinzel, J. Kohlus & H. Stumpe. 2005. Salzwiesen an der Westküste von Schleswig-Holstein 1988 - 2001. Schriftenreihe des Na-tionalparks Schlleswig-Holsteinisches Wattenmeer Vol. 15: 1-239.

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86

73


Issues of the Publication Series „Wadden Sea Ecosystem“

No. 1: Breeding Birds in the Wadden Sea 1991. 1994.No. 2: Migratory Waterbirds in the Wadden Sea1992/93. 1994.No. 3: Guidelines for Monitoring of Breeding Birds in the Wadden Sea (in Dutch, German, Danish). 1995.No. 4: Breeding Birds on Census Arteas 1990 until 1994. Status of Shorelark, Twite and Snow Bunting

in the Wadden Sea. 1997.No. 5: Migratory Waterbirds in the Wadden Sea 1993/94. 1996.No. 6: Trilateral Monitoring and Assessment Program. TMAP Expert Workshops 1995/96. 1996.No. 7: Assessment of the Wadden Sea Ecosystem. 1997.No. 8: Monitoring Breeding Success of Coastal Birds. Monitoring Pollutants in Coastal Bird Eggs in the

Wadden Sea. 1998.No. 9: Wadden Sea Quality Status Report 1999. 1999.No. 10: Breeding Birds in the Wadden Sea in 1996. 2000.No. 11: Contaminants in Bird Eggs in the Wadden Sea. Spatial and Temporal Trends 1999 - 2000. 2001. No. 12: Lancewad. Landscape and Cultural Heritage in the Wadden Sea Region. 2001.No. 13: Final Report of the Trilateral Working Group on Coastal Protection and Sea Level Rise. 2001.No. 14: Wadden Sea Specific Eutrophication Criteria. 2001.No. 15: Common and Grey Seals in the Wadden Sea. TSEG-plus Report March/June 2001.2002.No. 16: High Tide Roosts in the Wadden Sea. A Review of Bird Distribution, Protection Regimes and

Potential Sources of Anthropogenic Discturbance. 2003.No. 17: Management of North Sea Harbour and Grey Seal Populations. Proceedings of the International

Symposium at EcoMare, Texel, The Netherlands November 29 - 30, 2002. 2003.No. 18: Contaminants in Bird Eggs in the Wadden Sea. Recent Spatial and Temporal Trends. Seabirds at

Risk? Effects of Environmental Chemicals on Reproductive Success and Mass Growth of Seabirds at the Wadden Sea in the Mid 1990s. 2004.

No. 19: Wadden Sea Quality Status Report 2004. 2005.No. 20: Migratory Waterbirds in the Wadden Sea 1980 - 2000. 2005.No. 21: Coastal Protection and Sea Level Rise - Solutions for Sustainable Coastal Protection. 2005 No. 22: Breeding Birds in the Wadden Sea in 2001. 2006.No. 23: Seriously Declining Trends in Migratory Waterbirds: Causes-Concerns-Consequences. Proceedings

of the International Workshop on 31 August 2005 in Wilhelmshaven, Germany. 2007.No. 24: Nomination of the Dutch-German Wadden Sea as World Heritage Site. 2008.No. 25: Wadden Sea Quality Status Report 2009. 2009.No. 26: Science for Nature Conservation and Managment: The Wadden Sea Ecosystem and EU Direc-

tives. Proceedings of the 12th International Scientific Wadden Sea Symposium in Wilhelmshaven, Germany, 30 March - 3 April 2009. 2010.

No. 27: Exploring contrasting trends of migratory waterbirds in the international Wadden Sea. 2010.No. 28: CPSL Third Report. The role of spatial planning and sediment in coastal risk management. 2010.No. 29: The Wadden Sea - A Universally Outstanding Tidal Wetland. The Wadden Sea Quality Status

Report. Synthesis Report 2010.No. 30: Migratory Waterbirds in the Wadden Sea 1987-2008. 2010.No. 31: Trends of Migratory and Wintering Waterbirds in the Wadden Sea 1987/1988-2010/2011. 2013.No. 32: TMAP-Typology of Coastal Vegetation in the Wadden Sea Area. 2014.

74


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