—197—

This is part of a systematic study of East Asian Saussurea (Asteraceae) (Kadota 1987, 2004, 2007, 2008, 2009, 2010).

In 1974 Goda (1981) found unfamiliar plants of Saussurea in Hamanaka Moor, Hamanaka-cho, Akkeshi-gun, Kushiro Subprefecture, Hokkaido, northern Japan. He supposed the plants to belong an undescribed species and sent specimens to Dr. Hiroshige Koyama, enrolled in Kyoto University at that time, for determination. Koyama identified the plants as Saussurea acuminata Turcz. subsp. duiensis (F. Schmidt) Kitam. [= S. duiensis F. Schmidt] and told to him that Goda’s finding was the first report for this species in Japan. Goda (1981) published the record with a photograph and a line drawing in his work based on Koyama’s determination. One

of the specimens collected by Goda in 1974 is also kept in TNS (Y. Goda 120, TNS 334831). However, any plants belonging to ‘S. duiensis’ had never been collected from eastern Hokkaido since the time of Goda’s discovery. While S. duiensis has not been recognized in Japan (Kitamura 1937, 1980, 1981, Masamune 1974, Ito 1981, Takita 1987, Koyama 1995) except for Ito and Hinoma (1987).

In July 2010 the plants in question were obtained from Nemuro Subprefecture, eastern Hokkaido, by Ms. Yachiyo Takashima and her colleagues. However, they collected the plants from a single locality. Hence I asked Mr. Shun Umezawa to do a thorough survey in eastern Hokkaido in order to understand the occurrence of the plants. Consequently he found other

J. Jpn. Bot. 86: 197–204 (2011)

Systematic Studies of Asian Saussurea (Asteraceae) V.Two New Species from Eastern Hokkaido, Japan and

Sakhalin, Russia

Yuichi kadota

Department of Botany, National Museum of Nature and Science,4–1–1, Amakubo, Tsukuba, 305-0005 JAPAN

E-mail: kadota@kahaku.go.jp

(Accepted on March 3, 2011)

Two new species, Saussurea hamanakaensis Kadota and S. shiretokoensis Sugaw. ex Kadota, are described from eastern Hokkaido (northern Japan) and Sakhalin, respectively. Saussurea hamanakaensis is different from S. duiensis F. Schmidt by having 5-seriate involucral phyllaries, narrowly but distinctly winged stem, more shallowly incised corolla-lobes and cauline leaves devoid of cobwebby hairs on the abaxial side. Saussurea hamanakaensis occurs in moors of eastern Hokkaido, Japan. Saussurea shiretokoensis is distinguished from both S. duiensis and S. hamanakaensis by having outer and middle involucral phyllaries with caudate apices and is known from maritime herbal stands of Sakhalin.

Key words: Japan, Hokkaido, new species, Sakhalin, Saussurea duiensis, Saussurea hamanakaensis, Saussurea shiretokoensis.

198 植物研究雑誌　第 86 巻　第 4 号 2011 年 8 月

Fig. 1. Type of Saussurea hamanakaensis Kadota (JAPAN: Hokkaido. Kushiro Subpref., Akkeshi-gun, Hamanaka-cho, 3 Aug. 2010, S. Umezawa s.n., TNS 1114462, holotype).

August 2011 Journal of Japanese Botany Vol. 86 No.4 199

localities and collected an enough amount of specimens of the plants.

Taxonomic comparison was subsequently done between the plants from eastern Hokkaido and S. duiensis from Sakhalin. As a result it is concluded that the plants from eastern Hokkaido represent a distinct species from S. duiensis F. Schmidt. The plants from eastern Hokkaido are described as a new species, Saussurea hamanakaensis, after the locality name of the discovery.

During the comparative studies of S. hamanakaensis and the Saussurea species from Sakhalin “Saussurea shiretokoensis” (Sugawara 1937, 1940) was revised. “Saussurea shiretokoensis Sugaw.” was ascribed to a nomen nudum because the name was not accompanied by Latin descriptions nor type designation.

A Shigezo Sugawara collection is preserved at the Herbarium of Field Science Center for Northern Biosphere, Hokkaido University (SAPT). The specimens “Sakhalin, Kita-Shiretoko Peninsula, Funakoshi, 11 Aug. 1927, S. Sugawara 13993” quoted by Sugawara (1940, p. 1881) as S. shiretokoensis were not found even in this collection. However, the specimens agree with the description in Japanese and the line drawing of S. shiretokoensis (Sugawara 1940; tab. 864) have been located in SAPT. These specimens are provided with outer and middle involucral phyllaries with caudate apices and consequently are significantly different from S. duiensis. Then S. shiretokoensis is here described as a distinct species with accompanying typification and Latin description.

Taxonomic treatmentSaussurea DC. in Ann. Mus. Natl. Hist.

Nat. Paris 16: 156, 198 (1810) & Prodr. 6: 531 (1837).

Subgen. Saussurea: Lipsch., Fl. URSS 27: 392 (1962) & Genus Saussurea 95 (1979) – H. Koyama in K. Iwats. & al., Fl. Jap. 3a: 153 (1885) – C. Shih & S.-Y. Jing in Y. L. Chen & C. Shih, Fl. Reipub. Popul. Sin. 78(2): 66 (1999).

Sect. Saussurea: Lipsch., Gen. Saussurea 178 (1979) – H. Koyama in K. Iwats. & al., Fl. Jap. 3a: 153 (1995) – C. Shih & S.-Y. Jing in Y. L. Chen & C. Shih, Fl. Reipub. Popul. Sin. 78(2): 158 (1999).

Ser. Amurensia Lipsch., Fl. URSS 27: 559 (1962) & Genus Saussurea 215 (1979).

Ser. Acuminata Kitam. in Acta Phytotax. Geobot. 4: 7 (1935), p. p. & in Mem. Coll. Sci., Kyoto Imp. Univ., ser. B, 13: (Compos. Jap. I) 159 (1937), p. p.

Saussurea hamanakaensis Kadota, sp. nov. [Figs. 1, 2]

Saussurea duiensis auct. non F. Schmidt: Goda, Kushiro-chiho no Shokubutsu Sôsetsu: cover &124 (1981).

Differt a Saussurea duiensi phyllariis involucrorum 5-seriatis, caule anguste alato, corollis non profunde lobatis, foliis ventraliter non araneosis.

Type: JAPAN. Hokkaido, Kushiro Subpref., Akkeshi-gun, Hamanaka-cho [detailed locality data concealed for plant protection], 3 August 2010, S. Umezawa s.n. (TNS 1114462–holotype, Fig. 1; TNS 114463–isotype).

A slender, herbaceous perennial, 60–140 cm tall. Rhizome oblique, ca. 1.5 cm in diameter, with string-like roots. Stem erect, striate, narrowly but clearly winged, glandular-dotted in the upper part, 2–5 times branched; wings up to 2 mm wide. Basal leaves withering at anthesis. Lower cauline leaves subcoriaceous, narrowly lanceolate, 10–15 cm long, 1–2 cm wide, serrate, glabrous on the adaxial side, densely glandular-dotted on the abaxial side, cuneate at base, acuminate at apex: petioles 2–4 cm long, amplexicaul, glabrous. Upper cauline leaves linear to narrowly lanceolate, 5–10 cm long, 3–8 mm wide, serrate to entire, acuminate at apex, amplexicaul, similarly glandular-dotted to the lower cauline leaves, sessile and amplexicaul. Flowers in July to August, with 10–20 capitula, arranged in a compact corymb; peduncles 2–7 mm long in the terminal corymb, ascending,

200 植物研究雑誌　第 86 巻　第 4 号 2011 年 8 月

sparingly glandular-dotted. Involucres narrowly cylindrical, 4–5 mm in diameter, 8–10 mm long, sericeous; phyllaries 5-seriate; outer phyllaries broadly ovate to ovate, 2–3 mm long, cuspidate; inner phyllaries lanceolate, 8–10 mm long, acuminate; setae 2 mm long. Corollae pale purplish violet, 10 mm long; lobes 3 mm long; throats 1 mm long; tubes 6 mm long; anthers 4 mm long, purplish black. Pappi 2-whorled, inner 8 mm long, outer 3 mm long, grayish white. Achenes 3.5–4 mm long, glabrous, slightly reddish brown.

Japanese name: Konsen-tôhiren (nom. nov.).新和名：コンセントウヒレンDistribution: Nemuro and Kushiro Subprefs.,

Hokkaido, Japan. Endemic.Other specimens examined: JAPAN. Hokkaido,

Kushiro Subpref., Akkeshi-gun, Hamanaka-cho, 8 Aug. 1974, Y. Goda 120 (TNS 334831); Hamanaka-cho, 3 Aug. 2010, S. Umezawa s.n. (TNS 1114464–1114466).

Nemuro Subpref., Nemuro-shi, Hattaushi, 18 July 2010, Y. Takashima & Y. Yoshiazawa s.n. (TNS 1114472–1114473); Nemuro-shi, Hattaushi, 30 July 2010, Y. Takashima, Y. Yoshiazawa & Takeda s.n. (TNS 1114471); Nemuro-shi, Attoko, 28 July 2010, S. Umezawa s.n. (TNS 1114459–1114461); Nemuro-shi, Attoko, alt. ca. 40 m, 43°14′N 145°19′E, 3 Aug. 2010, S. Umezawa s.n. (TNS 1114467–1114469, 1114474).

Note: Saussurea hamanakaensis resembles S. duiensis in having narrowly cylindrical involucres and numerous heads arranged in a compact corymb. However, S. hamanakaensis is clearly discriminated from S. duiensis by having 1) 5-seriate involucral phyllaries (Fig. 2D), 2) narrowly winged stem, 3) shallowly lobed corollae, 4) leaves without cobwebby hairs. The

Fig. 2. Saussurea hamanakaensis Kadota. A. Habit. B. Habitat. C. Undersurface of leaf. D. Capitula. A–C. Hamanaka-cho, Kushiro Subpref., Hokkaido. Photos by Mr. S. Umezawa. D. Nemuro-shi, Nemuro Subpref., Hokkaido. Photo by Y. Kadota. Scale = 1cm.

August 2011 Journal of Japanese Botany Vol. 86 No.4 201

cauline leaves and the stem are densely covered with golden glandular dots (Fig. 2C) and they are very glutinous in the living state. Saussurea duiensis is similar to S. hamanakaensis in having glandular-dotted leaves and the stem. In Hokkaido, Japan, the other species of ser. Amurensia, S. riederi Herder, S. sachalinensis F. Schmidt and S. yesoensis Franch. are distinguished from S. hamanakaensis by the absence of glandular dots on leaves and stems.

Saussurea hamanakaensis grows along the edge of moors (Takashima pers. comm., Umezawa pers. comm.; Fig. 2B). While S. duiensis occurs in herb-stands and/or along the edge of woodlands (Sugawara 1940, Lipschitz 1962, 1979, Barkalov 1993). In this way the two species are different in habitat preference.

Saussurea shiretokoensis Sugaw. ex Kadota, sp. nov. [Fig. 3]

Saussurea shiretokoensis Sugaw., Fl. Saghal. 312 (1937), nomen & Fl. Saghal. 4: 1881, tab. 804 (1940), Japonice, nom. nud. – Honda, Nom. Pl. Jap. 369 (1939), nomen – Lipsch., Fl. URSS 27: 563 (1962), nom. nud. & Genus Saussurea: 247 (1979), nom. nud.

Di ffe r t a Saussurea duiens i e t S . hamanakaensi phyllariis involucrorum exterioribus et mediis apice caudatis; a S. sachalinensi foliis anguste ovato-lanceolatis infra glandulo-punctatis.

Type: RUSSIA. Sakhalin [Karafuto]. Shikka [Shisuka] Subpref., Chirie-gun, Chirie-mura, Funakoshi (currently no villages, near Peskovskiy), 10 Aug. 1927, S. Sugawara 22976 (SAPT–holotype; Fig. 3). Funakoshi, 10 August 1927, S. Sugawara 22975, 22977 (SAPT–isotypes).

A perennial herb, 12–30 cm high. Rootstock suberect, 4–6 mm in diameter. Stem erect, simple, pubescent with multicellular hairs in the upper half, glabrous in the lower half. Basal leaves withered at anthesis. Lower cauline leaves subscabrid, narrowly ovato-lanceolate, 7–9.5 cm long, 1.5–2 cm wide, serrate with awn-shaped

teeth, acuminate at apex, roundish at base, sparingly arachnoid on the adaxial side, densely glandular-dotted on the abaxial side, devoid of cobwebby hairs, long petiolate; petioles 5–9 cm long, winged, amplexicaul at base. Middle cauline leaves oblanceolate to narrowly ovate, 5–9 cm long, 0.8–2 cm wide, similarly serrate to the lower ones, short-petiolate; petioles 0.5–1.5 cm long or subsessile, winged. Upper cauline leaves narrowly oblanceolate, 3–5 cm long, ca. 5 mm wide, subentire, sessile. Inflorescence terminal, corymbose, 10–15-flowered; peduncles 3–5 mm long, thick, densely pubescent with brownish multicellular hairs and glandular-dotted; bracts foliaceous, narrowly lanceolate, acuminate, entire. Involucres narrowly cylindrical, 5–6 mm in diameter, with linear bracteoles. Involucral phyllaries 5-seriate, tinged dark purplish black along the margin and at base, sericeous and ciliate-margined, herbaceous; outermost phyllaries narrowly ovate, 2–3 mm long, caudate at apex; middle ones narrowly ovato-lanceolate, 6–7 mm long, shortly acuminate; innermost ones narrowly ovato-lanceolate, ca. 1 cm long, acute at apex. Florets purplish pink. Corollae purplish violet, 12 mm long; lobes 6 mm long; throats 1 mm long; tubes 6 mm long; anthers 4 mm long, purplish black. Pappi 2-whorled, inner 10 mm long, outer 2 mm long, grayish white.

Japanese name: Shiretoko-azami (Sugawara 1937).

和名：シレトコアザミ（菅原 1937）Distribution: Central Sakhalin (Karafuto).

Endemic.Habitat: Maritime herb-stands (Sugawara

1940).Other specimens examined: RUSSIA. Sakhalin

[Karafuto]. Esutori Subpref., Nayoshi-gun, Nayoshi-mura, Yokunai (currently near Lesogorsk), 10 Aug. 1933, S. Sugawara 22978 (SAPT). Motodomari Subpref., Motodomari-gun, Hoyori-mura, Maguntan (currently Pugachevo), 19 Aug. 1927, S. Sugawara 22979 (SAPT).

Note: Saussurea shiretokoensis is different from S. duiensis and S. hamanakaensis by having outer and middle involucral phyllaries

202 植物研究雑誌　第 86 巻　第 4 号 2011 年 8 月

Fig. 3. Type of Saussurea shiretokoensis Sugaw. ex Kadota (RUSSIA: Sakhalin, Sikka Distr., Funakoshi, 10 Aug. 1927, S. Sugawara 22976, SAPT, holotype). Right bottom corner inset shows the inflorescence of the right indi-vidual.

August 2011 Journal of Japanese Botany Vol. 86 No.4 203

with caudate apices. Saussurea sachalinensis F. Schmidt is similar to S. shiretokoensis in having caudate involucral phyllaries but the former is distinguished from the latter by having leaves glandular-dotted on the abaxial side.

Saussurea shiretokoensis has not been recognized to the flora of Sakhalin (Miyabe and Miyake 1915, Kudo 1927, Voroschilov 1982). Barkalov (1993) treated S. shiretokoensis as a synonym of S. duiensis. Lipschitz (1962, 1979) regarded S. shiretokoensis as a closer species to S. duiensis based on Sugawara’s line drawing because he could not understand the description written in Japanese. Lipschitz (1962) also stated that S. shiretokoensis might be of hybrid origin between S. duiensis and S. sachalinensis due to the shapes of involucral phyllaries and cauline leaves. The hybridity of S. shiretokoensis remains unsolved and it must be qualified in the field.

I would like to express my sincere thanks to Mr. Shun Umezawa for his presents of living material, specimens and photographs of Saussurea hamanakaensis; to Mses. Yachiyo Takashima, Yoshiko Yoshizawa, Messrs. Norihisa Kondo, for their present and valuable information on S. hamanakaensis; to Mr. Michio Sato, Hakodate City Museum, for his information on the specimens collected by Mr. Shigezo Sugawara; to Professor Emeritus Dr. Hiroyoshi Ohashi for his nomenclatural advice on S. shiretokoensis; to Dr. S. V. Smirnov, South-Siberian Botanical Garden, Altai State University, Barnaul, for his present of type images and information on East Asian Saussurea; to Dr. Hideki Takahashi and Dr. Takayuki Azuma for their kind permission of examination of Saussurea specimens deposited at SAPT; to Mr. Shungo Kariyama for his kind access to Saussurea specimens from Sakhalin housed at KURA.

Literatures citedBarkalov V. Yu. 1993. Saussurea. In: Charkevicz S. S.

(ed.), Plantae Vasculares Orientis Extremi Sovietici 6:

254–296. Nauka, Saint Petersburg (in Russian).Goda Y. 1981. Kushiro-chihô no Shokubutsu Sôsetsu. 265

pp. Self-publishing (in Japanese).Ito K. 1981. Saussurea. Alpine Plants of Hokkaido. pp.

14–15. Seibundo Shinkosha, Tokyo (in Japanese).Ito K. and Hinoma A. 1987. Saussurea. Check List

of Higher Plants in Hokkaido IV (Sympetalae): 180–184. Takugin Research Institute, Sapporo (in Japanese).

Kadota Y. 1987. A new variety of Saussurea kudoana Tatew. & Kitam. (Asteraceae) from Hokkaido, Japan. Mem. Natn. Sci. Mus., Tokyo (20): 83–90.

Kadota Y. 2004. A new species of Saussurea (Asteraceae) from Honshu, Japan. J. Jpn. Bot. 79: 235–240.

Kadota Y. 2007. Systematic studies of Asian Saussurea (Asteraceae) I. Saussurea kubotae, a new species from western Japan. J. Jpn. Bot. 82: 259–265.

Kadota Y. 2008. Systematic studies of Asian Saussurea (Asteraceae) II. Two new species from northern Japan. J. Jpn. Bot. 83: 284–294.

Kadota Y. 2009. Systematic studies of Asian Saussurea (Asteraceae) III. Saussurea fuboensis, a new species from the southernmost part of Tohoku District, northern Japan. J. Jpn. Bot. 84: 177–183.

Kadota Y. 2011. On the entity of Saussurea triptera Maxim. var. mikurashimensis Kitam. (Asteraceae) from Mikurajima Island, Idzu Islands, Japan (Systematic studies of Asian Saussurea (Asteraceae) V). Mem. Natl. Mus. Nat. Sci., Tokyo (47): 411–415.

Kitamura S. 1935. Les Saussurées du Japon; leur classification et leur distribution. Acta Phytotax. Geobot. 4: 1–14.

Kitamura S. 1937. Compositae Japonicae. Pars prima. Mem. Coll. Sci., Kyoto Imp. Univ., ser. B, 13: 140–212.

Kitamura S. 1980. Saussurea. In: Kitamura S., Murata G. and Hori M., Coloured Illustrations of Herbaceous Plants of Japan (Sympetalae), revised edition. pp. 24–29, figs. 7–8, pl. 7. Hoikusha Publishing Co. Ltd., Osaka (in Japanese).

Kitamura S. 1981. Saussurea. In: Satake Y., Ohwi J., Kitamura S., Watari S. and Tominari T. (eds.), Wild Flowers of Japan. Herbaceous Plants (including Dwarf Subshrubs). 3: 220–224, pls. 201–207. Heibonsha Ltd., Tokyo (in Japanese).

Koyama H. 1995. Saussurea. In: Iwatsuki K., Yamazaki T., Boufford D. E. and Ohba H. (eds.), Flora of Japan 3a: 152–162. Kodansha Ltd., Tokyo.

Kudo Y. 1924. Saussurea. Kita-Karafuto Shokubutsu Chôsasho [Contribution to the Flora of Northern Sakhalin]. pp. 241–243. Saghalien Gunsei-bu, Toyohara (in Japanese).

Lipschitz S. 1962 (1997). Saussurea. In: Shishkin B. K. and Bobrov E. G. (eds.), Flora of the U.S.S.R. (Flora SSSR) 27: 447–658. Bishen Singh Mahendra Pal

204 植物研究雑誌　第 86 巻　第 4 号 2011 年 8 月

門田裕一：アジア産トウヒレン属（キク科）の分類学的研究 V．北海道東部とサハリンからの 2 新種　 北 海 道 東 部とサハリン（ 樺 太 ） から， キク科トウヒレン属トウヒレン節の 2 新 種，コンセントウヒレンSaussurea hamanakaensis Kadota とシレトコアザミ S. shiretokoensis Sugaw. ex Kadota をそれぞれ記載した．　コンセントウヒレンはこれまでススヤアザミ S. duiensis F. Schmidt [= S. acuminata Turcz. subsp. duiensis (F. Schmidt) Kitam.] と考えられてきたものである．コンセントウヒレンはススヤアザミとは次の点で異なる．①総苞片は 5 列で，②茎には狭いが明瞭な翼があり，③花冠の切れ込みが浅く，④葉の下面には腺点のみでクモ毛がない．本種は北海道・道東地方の固有種で，ススヤアザミはサハリンの固有種である．　コンセントウヒレンは茎の上部や葉の下面に黄金色の腺点が密布し，生時は著しく粘着する．コンセントウヒレンの茎や葉の下面に腺点が密布するのはススヤアザミと共通しており，形態的に似た北海道産のエゾトウヒレンS. yesoensis Franch.，カラフトアザミ S. sachalinensis F. Schmidt，ナガバキタアザミ S. riederi Herder などとの明瞭な区別点となる．　学名の種形容語は最初に発見された厚岸郡浜中町に，和名は根室地方と釧路地方の双方に分布することによる．コンセントウヒレンは 1974 年に合田勇太郎によって浜中町で発見された（合田 1981）．これは小山博滋によりススヤアザミと同定された．浜中町で得られた後しばらく記録が途絶えていたが，2010 年になって浜中町と根室市で再発見された．道東地方での生育地は湿原とその周辺のやや水湿の地である（高嶋　私信，梅澤　私信）．ススヤア

Singh, Dehra Dun, and Koeltz Scientific Books, Koenigstein (translated from Russian).

Lipschitz S. 1979. Genus Saussurea DC. 282 pp. Nauka, Moskva (in Russian).

Masamune G. 1974. Saussurea. Color Illustrated Flora of Nippon 6-II: 331–341. Koyo Shoin, Tokyo (in Japanese).

Miyabe K. and Miyake T. 1915. Saussurea. Flora of Saghalin. pp. 276–281. The Government of Saghalin, Sapporo (in Japanese).

Shih C. and Jin S.-Y. 1999. Saussurea. In: Chen Y.-L. and Shih C. (eds.), Flora Reipublicae Popularis Sinicae 78(2): i–x, 1–232 (in Chinese).

Suagwara S. 1937. Plants of Saghalien. A List of the Vegetation – Totality in the Saghalien. With the

description of Ligneous Plants, Useful Plants and Naturalized Plants. 490 pp. Sugawara Shigezo Karafuto Shokubutsu Kenkyu Kôenkai, Toyohara.

Sugawara S. 1940. Saussurea. Flora of Saghalien with Descriptions and Figures of Phanerogams and Higher Cryptogams Indigenous to Saghalien. Diapensiaceae – Asteraceae. 4: 1868–1883, tabs 858–865 (in Japanese).

Takita K. 1987. Saussurea. Higashi-Hokkaido no Shokubustu [Plants of Eastern Hokkaido]. p. 736. Kato Shokan, Kushiro (in Japanese).

Voroschilov V. N. 1982. Saussurea. Opredelitely Rastenij Sovietskogo Dalynego Vostoka. pp. 572–580. Nauka, Leningrad (in Russian).

ザミは樹陰草原地（菅原 1940）や森林の林縁，潅木のある茂み，草原 (Lipschitz 1962, 1989) に生育するので，この 2 種には生育環境にも違いがあると考えられる．　サハリン（樺太）から記載されたシレトコアザミはコンセントウヒレンやススヤアザミと同じように葉の下面に腺点があるが，総苞外片と中片の先端が長く尾状に尖る点でこれら 2 種とは明瞭に区別される．シレトコアザミは菅原 (1937, 1940) によって発表されたものであるが，タイプ標本が指定されておらず，またラテン語の記載も与えられていない裸名であった．サハリンの植物誌的研究ではシレトコアザミは言及されることはないか，あるいはススヤアザミの異名として扱われることが多かった．Lipschitz (1962, 1979) は 菅 原 の 図（ 菅 原 1940；p. 1880, tab. 864）に基づいてシレトコアザミの存在を認めたが，正式に記載することはなかった．　菅原 (1940) に引用された標本（菅原 13993）の所在は現段階では不明のままであるが，北海道大学北方生物圏フィールド科学センター植物園に所蔵されている標本が記載に良く一致するので，その中から「樺太，北知床半島，舟越，1927 年 8 月 10 日，菅原繁蔵 22976」をタイプとして選定してラテン語の記載を与え，学名を Saussurea shiretokoensis Sugaw. ex Kadotaとした．シレトコアザミはサハリンの固有種で，総苞片の先端が尾状に尖る点ではナガバキタアザミに似るが，葉の下面に腺点を密布する点でナガバキタアザミとは異なる別の一群である．

（国立科学博物館植物研究部）