scale insects and mealy bugs (insecta: homoptera: coccoidea)

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SCALE INSECTS AND MEALY BUGS (INSECTA: HOMOPTERA: COCCOIDEA)

R.K.VARSHNEY*, M.J. JADHAV AND R.M.SHARMA

Zoological Survey of India, Western Regional Centre, Pune-411044 *Raj Bhawan, Manik Chowk, Aligarh-202001

The Scale insects and mealy bugs fall under the Superfamily Coccoidea, Order

Homoptera. The name refers to their secretion of waxy scales or mealy that serves as a protective

covering. They range in size from 1.5 mm to 25 mm in length. They are economically important

insects, since many species attack different parts of plant and feed by sucking the plant sap. On

the other hand a few species of scale insects are beneficial to mankind, e.g. the cochineal insect

(Dactylopius indicus) yields a carmine dye and the lac insect (Kerria lacca) is the source of

shellac. They are found on nearly all parts of the host plant including the leaves, branches,

trunks, fruits, and roots. They sometimes occur under bark and may cause various kinds of plant

deformities including chlorotic spots, pits, and galls. They cause severe damage to economically

important plants like sugarcane, tea, coffee, cotton, sorghum, vegetables, greenhouse plants and

almost all kinds of fruit trees.

About 7,500 species of Scale insects and Mealy bugs are known at present from the

world which inhabits almost all parts of the globe, in all floral habitats, from the tundra to the

tropics. The Indian diversity accounts to 409 species under 166 genera spread over 14 families

and 14 subfamilies (Varshney; 1992, 2002) and other scattered literature.

SYSTEMATIC ACCOUNT Superfamily: Coccoidea Family: Margarodidae

Subfamily: Margarodinae Tribe: Kuwanini

1. Genus: Kuwania Cockerell in Fernald, 1903

1. Kuwania zeylanica (Green) Fernald, 1903

Distribution: Kerala (Calicut).

Tribe: Margarodini

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2. Genus: Margarodes Guilding, 1829

2. Margarodes formicarum Guilding, 1829

Distribution: South India.

3. Margarodes indicus Green, 1912

Distribution: Mumbai and Bengaluru.

4. Margarodes papillosus Green, 1912

Distribution: Karnataka (Bengaluru, Bellary).

Subfamily: Neomargarodinae

3. Genus: Neomargarodes Green, 1914

5. Neomargarodes niger Morrison, 1928

Distribution: Karnataka (Mysore, Bellary), Tamil Nadu (Coimbatore).

Family: Monophlebidae Subfamily: Monophlebinae

Tribe: Iceryini 4. Genus: Crypticerya Cockerell, 1895

Crypticerya sp.

Distribution: Odisha (Bhubaneshwar, Taptapani, Talcher and Angul).

6. Crypticerya jacobsoni Morrison, 1928

Distribution: Assam (Dibrugarh, Jokai Forest), Karnataka (Bengaluru), Kerala (Cochin).

7. Crypticerya jaihind Rao, 1951

Distribution: Karnataka (Kanara Dist., Mangalore), Kerala (Cochin, Nelliampathy hills, Travancore), Tamil Nadu (Nilambur).

8. Crypticerya kumari Rao, 1951

Distribution: Karnataka (Gopalswami hills).

5. Genus: Icerya Signoret, 1875

Icerya sp.

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Distribution: Odisha (Bhubaneshwar, Taptapani, Dhauli, Konark, Rambha, Gopalpur-on-sea, Kuliana and Baripada).

9. Icerya aegyptiaca (Douglas, 1890)

Distribution: Bihar, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu, Tripura, West Bengal.

10. Icerya formicarum Newstead, 1897

Distribution: Andhra Pradesh, Bihar, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu, Uttar Pradesh, West Bengal and North-East India.

11. Icerya menoni Rao, 1951

Distribution: Kerala (Maradu).

12. Icerya minor Green, 1908

Distribution: Assam (Haflong), Bihar (Pusa), Uttar Pradesh (Varanasi).

13. Icerya pilosa Green, 1896

Distribution: Andhra Pradesh (Sarola), Bihar (Shahabad, Juneshwargarh farm), Delhi, Haryana (Karnal), Madhya Pradesh (Gwalior), Kerala (Perungaloor, Travancore), Tamil Nadu (Chennai, Hasanoor, Coimbatore, Kanyakumari, Satyamangalam, Coromandel coast), Uttar Pradesh (Aligarh, Meerut, Shahjahanpur) and West Bengal.

14. Icerya pulcher Morrison, 1928

Distribution: Bihar (Pusa).

15. Icerya purchase Maskell, 19879

Distribution: Karnataka, Kerala, Maharashtra, Odisha, and Tamil Nadu.

16. Icerya seychellarum (Westwood, 1855)

Distribution: Andaman Is., Karnataka (Bellary, Kottur), Kerala, Madhya Pradesh, Maharashtra, Odisha, and Tamil Nadu.

17. Icerya travancorensis Rao, 1951

Distribution: Kerala, Tamil Nadu.

18. Icerya zimmermani Green, 1932

Distribution: Bihar (Bhagalpur).

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6. Genus: Steatococcus Ferris, 1921

19. Steatococcus assamensis Rao, 1951

Distribution: Meghalaya (Dawki).

Tribe: Drosichini

7. Genus: Drosicha Walker, 1858

Drosicha sp.

Distribution: Tamil Nadu (Chennai), Bihar (Pusa).

20. Drosicha burmeisteri Cockerell, 1902

Distribution: North India.

21. Drosicha contrahens Walker, 1858

Distribution: North-East India.

22. Drosicha dalbergiae (Green, 1902)

Distribution: Bihar, Punjab (Sutlej Valley) and Uttarakhand (Dehra Dun).

23. Drosicha leachii (Westwood, 1845)

Distribution: Kerala, Uttarakhand (Dehra Dun).

24. Drosicha mangiferae (Green, 1903)

Distribution: Bihar (Bhagalpur, Pusa, Sitamarhi, Dalsingsarai, Santhal Paraganas, Darbhanga, Muzaffarpur), Delhi, Uttar Pradesh (Fatehpur), Uttarakhand (Dehra Dun).

25. Drosicha quadricaudata (Green, 1922)

Distribution: Uttar Pradesh (Varanasi).

26. Drosicha saundersii (Westwood) Foldi, 2001


27. Drosicha stebbingii (Green); Morrison, 1928

Distribution: Bihar (Darbhanga, Saran, Muzaffarpur), Haryana (Karnal), Himachal Pradesh (Shimla), Madhya Pradesh, Rajasthan (Jodhpur), Uttarakhand (Dehra Dun), Uttar Pradesh (Saharanpur, Kanpur, Varanasi), West Bengal (Howrah & Darjeeling Dist.)

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28. Drosicha tectonae (Green); Morrison, 1928

Distribution: Madhya Pradesh (Damoh), Punjab, Uttar Pradesh.

Tribe: Monophlebini

8. Genus: Aspidoproctus Newstead, 1901

29. Aspidoproctus pertinax Newstead, 1901

Distribution: Karnataka (Bengaluru).

9. Genus: Hemaspidoproctus Morrison, 1927

30. Hemaspidoproctus cinereus Morrison, 1927

Distribution: Bihar (Pusa), Karnataka, Kerala, Gujarat, Rajasthan and Tamil Nadu.

10. Genus: Labioproctus Green, 1922

31. Labioproctus poleii Green, 1896

Distribution: Karnataka (Mysore, Coorg), West Bengal (Darjeeling Dist.).

11. Genus: Misracoccus Rao, 1950

32. Misracoccus assamensis Rao, 1950

Distribution: Meghalaya (Shilong).

33. Misracoccus xyliae Rao, 1950

Distribution: Karnataka (Molakalmurn), Kerala (Malabar) and Tamil Nadu (Coimbatore).

12. Genus: Monophlebus Guerin-Meneville, 1827

Monophlebus sp.

Distribution: Tamil Nadu.

34. Monophlebus atripennis Burmeister, 1835


13. Genus: Monophlebidus Morrison, 1927

35. Monophlebidus indicus Morrison, 1927

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14. Genus: Perissopneumon Newstead, 1900

36. Perissopneumon ferox Newstead, 1900

Distribution: Kerala, Maharashtra, Tamil Nadu, Uttar Pradesh.

37. Perissopneumon phyllanthi (Green, 1923)

Distribution: Andhra Pradesh (Godavari Dist.), Karnataka (Bengaluru), Kerala (Malabar, Walayar Forest) and West Bengal (Kolkata).

38. Perissopneumon tamarinda Rao, 1950

Distribution: Andhra Pradesh (Godavari Dist.) and West Bengal (Kolkata).

15. Genus: Walkeriana Signoret, 1876

39. Walkeriana senex Green, 1896

Distribution: Kerala (Wynaad) and Tamil Nadu.

Family: Ortheziidae

16. Genus: Orthezia Bosc d’ Antic, 1784

40. Orthezia insignis Browne, 1887

Distribution: Karnataka (Bengaluru), Kerala (Wynad), Tamil Nadu (Salem Dist.), Sikkim (Rangpo) and West Bengal.

Family: Tachardiidae Subfamily: Tachardiinae

Tribe: Tachardini

17. Genus: Kerria Targioni-Tozzetti, 1884

Subgenus: (i) Kerria

Kerria sp. Varshney, 2000

Distribution: Odisha (Bhubaneshwar) and Tamil Nadu (Jawalgiri).

41. Kerria (K.) albizziae (Green, 1911)

Distribution: Bihar, Uttarakhand (Dehra Dun), West Bengal.

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42. Kerria (K.) brancheata Varshney, 1966

Distribution: Chhatisgarh, Jharkhand.

43. Kerria (K.) chamberlini Varshney, 1966

Distribution: Rajasthan (Jodhpur).

44. Kerria (K.) chinensis chinensis (Mahdihassan, 1923)

Distribution: North- East India.

45. Kerria (K.) chinensis kydia (Misra, 1930)

Distribution: Assam (Cachar Dist.).

46. Kerria (K.) communis (Mahdihassan, 1923)

Distribution: Andhra Pradesh, Goa, Karnataka, Kerala, Maharashtra, Tamil Nadu.

47. Kerria (K.) ebrachiata (Chamberlin, 1923)

Distribution: Chhatisgarh, Jharkhand, Karnataka (Bengaluru).

48. Kerria (K.) fici fici (Green, 1903)

Distribution: Bihar (Patna), Delhi, Madhya Pradesh, Uttar Pradesh, Jammu & Kashmir, Rajasthan, Tamil Nadu (Coimbatore), West Bengal.

49. Kerria (K.) fici jhasiensis (Misra, 1930)

Distribution: Uttar Pradesh (Jhansi).

50. Kerria (K.) indicola (Kapur, 1958)

Distribution: Bihar.

51. Kerria (K.) lacca lacca (Kerr, 1782)

Distribution: Andhra Pradesh, Bihar, Karnataka, Odisha, Tamil Nadu, Uttar Pradesh and West Bengal.

52. Kerria (K.) lacca ambigua (Misra, 1930)

Distribution: Uttar Pradesh (Guna, Jhansi).

53. Kerria (K.) lacca mysorensis (Mahadihassan, 1923)

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Distribution: Karnataka (Bengaluru, Mysore).

54. Kerria (K.) nagoliensis (Mahadihassan, 1923)

Distribution: Madhya Pradesh (Raipur, Sohagpur).

55. Kerria (K.) nepalensis Varshney, 1977


56. Kerria (K.) pusana (Misra, 1930)


57. Kerria (K.) laca mysorensis (Mahadihassan, 1923)

Distribution: Karnataka.

58. Kerria (K.) sharda Misra & Sushil, 2000 rangoonensis (Chamberlin, 1925)

Distribution: Odisha.

Subgenus: (ii) Chamberliniella Varshney, 1987

59. Kerria (C.) rangoonensis (Chamberlin, 1925)

Distribution: Assam.

Subfamily: Tachardininae

18. Genus: Paratachardina Balachowsky, 1950

60. Paratachardina mahdihassani (Kondo & Gullan, 2007)


61. Paratachardina mithila Varshney, 1968

Distribution: Meghalaya (Shillong).

62. Paratachardina silvestri (Mahdihassan, 1923)

Distribution: Andhra Pradesh, Karnataka (Bengaluru) and Tamil Nadu.

63. Paratachardina ternate (Chamberlin, 1923)

Distribution: Kerala (Travancore).

64. Paratachardina theae (Green) in Green & Mann, 1907

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Distribution: Assam, Sikkim and West Bengal.

Family: Eriococcidae

19. Genus: Eriococcus Targioni-Tozzetti, 1869

65. Eriococcus araucariae Maskell, 1879

Distribution: Karnataka (Bengaluru) and Tamil Nadu.

66. Eriococcus lagerstroemiae Kuwana, 1907


67. Eriococcus phyllanthi Ferris, 1957


68. Eriococcus rhodomyrti Green, 1922

Distribution: Maharashtra.

69. Eriococcus transwersus Green, 1922

Distribution: India

70. Eriococcus beesoni Green, 1926

Distribution: Odisha (Ganjam Dist.) and Tamil Nadu

Family: Kermesidae

20. Genus: Kermes Boitard, 1828

71. Kermes himalayensis Greee, 1909

Distribution: North-West India.

21. Genus: Pseudopulvinaria Atkinson, 1889

72. Pseudopulvinaria sikkimensis Atkinson, 1889

Distribution: Assam, Meghalaya, Sikkim, Shillong.

Family: Dactylopiidae

22. Genus: Dactylopius Costa, 1835

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Dactylopius sp.

Distribution: India: Odisha (Bhubaneswar).

73. Dactylopius coccus Costa, 1835

Distribution: India.

74. Dactylopius indicus Green, 1922

Distribution: Andhra Pradesh (Godavari Dist.), Bihar (Patna, Gaya), Himachal Pradesh, Odisha, Tamil Nadu and West Bengal.

75. Dactylopius tomentosus (Lamarck, 1801)

Distribution: Odisha, Karnataka, Maharashtra, Tamil Nadu and West Bengal.

Family: Pseudococidae Sub family: Pseudococcinae

Tribe: Phenacoccinini

23. Genus: Birendracoccus Ali, 1975

76. Birendracoccus saccharifolii (Green, 1908)

Distribution: Andhra Pradesh, Bihar, Delhi, Uttar Pradesh (Aligarh, Meerut) and West Bengal.

24. Genus Brevennia Goux, 1940

77. Brevennia rehi (Lindinger, 1943)

Distribution: Andhra Pradesh, Bihar, Delhi, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu and West Bengal.

25. Genus: Coccidohystrix Lindinger, 1943

Coccidohystrix sp.

Distribution: Punjab (Hoshiarpur).

78. Coccidohystrix insolita (Green, 1908)

Distribution: Andhra Pradesh, Bihar, Gujarat, Goa, Kerala, Haryana, Tamil Nadu and West Bengal.

26. Genus: Helicoccus Sulc, 1912

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79. Helicoccus singularis Avasthi & Shafee, 1982

Distribution: Andhra Pradesh.

27. Genus: Naiacoccus Green, 1919

80. Naiacoccus serpentines Green, 1919

Distribution: Punjab and Uttar Pradesh.

28. Genus: Nipacoccus Sulc, 1945

Nipacoccus sp.

Distribution: Punjab, Karnataka and Odisha.

81. Nipacoccus viridis (Newstead, 1894)

Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Himachal Pradesh, Maharashtra, Kerala, Karnataka, Uttar Pradesh and West Bengal.

29. Genus: Novonilacoccus Ghosh & Ghose, 1987

82. Novonilacoccus oryzae Ghosh & Ghose, 1987

Distribution: West Bengal (Kalyani).

30. Genus: Paraputo Laing, 1929

83. Paraputo taraktogeni Rao, 1950

Distribution: Assam (Dibrugarh).

31. Genus: Phenacoccus Cockerell, 1893

Phenacoccus sp.

Distribution: Kashmir and Uttarkhand (Dehra Dun).

84. Phenacoccus indicus (Avashti & Shafee, 1980)

Distribution: Karnataka (Mysore, Bengaluru).

85. Phenacoccus quaternus (Green, 1912)

Distribution: Tamil Nadu (Coromandel Coast).

32. Genus: Rastrococcus Ferris, 1954

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Rastrococcus sp.

Distribution: Bihar (Patna).

86. Rastrococcus iceryoides (Green, 1908)

Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Himachal Pradesh, Maharashtra, Kerala, Karnataka, Uttar Pradesh and West Bengal.

87. Rastrococcus invadens Williams, 1986

Distribution: India

88. Rastrococcus mangiferae Ferris, 1954

Distribution: Andhra Pradesh, Karnataka, Odisha and Tamil Nadu.

89. Rastrococcus ornatus Ferris, 1954


90. Rastrococcus spinosus (Robinson, 1918)

Distribution: Odisha (Bhubaneswar).

91. Rastrococcus resinophila Green, 1916

Distribution: Himachal Pradesh, Jammu & Kashmir and Uttar Pradesh.

Tribe: Pseudococcinini

33. Genus: Adolesoma Borchsenius, 1948

92. Adolesoma phragmitidis Borchsenius, 1948

Distribution: West Bengal.

34. Genus: Cataenococcus Ferris, 1955

Cataenococcus sp.

Distribution: Andaman Is.

93. Cataenococcus theaecola Das & Ganguli, 1962

Distribution: Assam and West Bengal.

35. Genus: Chorizococcus McKenzie, 1960

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94. Chorizococcus sorghi Williams, 1970

Distribution: Madhya Pradesh.

36. Genus: Crisicococcus Ferris, 1950

95. Crisicococcus dischidae Ezzat & McConnell, 1959

Distribution: Tripura.

37. Genus: Dysmicoccus Ferris, 1950

96. Dysmicoccus breviceps (Cockerell, 1893)

Distribution: Andhra Pradesh, Karnataka, Kerala, Odisha, Tripura and West Bengal.

97. Dysmicoccus carens Williams, 1970


98. Dysmicoccus cocotis (Maskell, 1890)

Distribution: Kerala.

99. Dysmicoccus cucurbitae Avashti & Shafee, 1986

Distribution: Uttar Pradesh.

100. Dysmicoccus triadus Williams, 1987


101. Dysmicoccus williamsi Avashti & Shafee, 1981


38. Genus: Ferrisia Fullaway, 1923

Ferrisia sp.

Distribution: Odisha (Bhubaneswar, Puri).

102. Ferrisia virgata (Cockerell, 1893)

Distribution: Andhra Pradesh, Bihar, Goa, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Tamil Nadu, West Bengal and Uttar Pradesh.

39. Genus: Indococcus Ali, 1968

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Indococcus sp.


103. Indococcus pipalae Ali, 1968

Distribution: Bihar and Odisha.

40. Genus: Kermicus Newstead, 1897

104. Kermicus wroughtoni Newstead, 1897

Distribution: India

41. Genus: Kiritshenkella Borchsenius, 1948

105. Kiritshenkella sacchari (Green, 1900)

Distribution: Andhra Pradesh, Bihar, Assam, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Tamil Nadu and West Bengal.

42. Genus: Maconellicoccus Ezzat, 1958

106. Maconellicoccus hirsutus Green , 1908

Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Tamil Nadu, Uttar Pradesh.

107. Maconellicoccus perforates (De Lotto 1954)


43. Genus: Paracoccus Ezzat & McConnell, 1956

108. Paracoccus burnerae Ezzat & McConnell, 1956


109. Paracoccus nellorensis Avashti & Shafee, 1984


44. Genus: Planococcoides Ezzat & McConnell, 1956

110. Planococcoides bengalensis Ghosh & Ghose, 1988


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111. Planococcoides robustus Ezzat & McConnell, 1956

Distribution: Andhra Pradesh, Bihar, Gujarat, Karnataka, Kerala, Maharashtra and Uttar Pradesh.

45. Genus: Planococcus Ferris, 1950

Planococcus sp.


112. Planococcus citri Ferris, 1950

Distribution: Karnataka, Kerala and Tamil Nadu.

113. Planococcus ficus Fernald, 1903


114. Planococcus indicus Avashti & Shafee, 1987

Distribution: Kerala (Ernaculum) and Tamil Nadu (Coimbatore: Botanical Garden)

115. Planococcus lilacinus (Cockerell, 1905)

Distribution: Andhra Pradesh, Karnataka, Kerala, Odisha and Tamil Nadu.

46. Genus: Pseudococcus Westwood, 1840

Pseudococcus sp.

Distribution: Andhra Pradesh, Maharashtra and Odisha.

116. Pseudococcus formiceticola (Newstead, 1900)

Distribution: Maharashtra (Konkan).

117. Pseudococcus longispinus (Targioni-Tozzetti, 1867)

Distribution: Andhra Pradesh, Karnataka, Kerala, Tamil Nadu and West Bengal.

118. Pseudococcus saccharicola Takahashi, 1928

Distribution: Tamil Nadu (Coimbatore).

47. Genus: Saccharococcus Ferris, 1950

119. Saccharicoccus sacchari (Cockerell, 1895)

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Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Haryana, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Odisha, Tamil Nadu and Uttar Pradesh.

48. Genus: Trionymus Berg, 1899

120. Trionymus ceres Williams, 1970

Distribution: Madhya Pradesh.

Tribe: Rhizoecini

49. Genus: Geococcus Green, 1902

121. Geococcus citrinus Kuwana, 1923

Distribution: Tamil Nadu (North Arcot Dist.).

122. Geococcus coffeae Green, 1933


50. Genus: Rhizoecus Kunckel d’Herculais, 1878

123. Rhizoecus amorphophalli Betram, 1940


124. Rhizoecus cocois Williams, 1985

Distribution: Kerala (Thiruvananthapuram Dist.).

125. Rhizoecus cynodontis Green, 1931

Distribution: Andhra Pradesh (Godavari Dist.) and Tamil Nadu (Coimbatore).

51. Genus: Xenococcus Silvestri, 1924

126. Xenococcus annandalei Silvestri, 1924

Distribution: Andhra Pradesh, Karnataka, Odisha (Berkuda Is. In Chilka lake) and Tamil Nadu.

127. Xenococcus scorpioides De Lotto, 1977


Tribe: Sphaerococcini

52. Genus: Antonina Signoret, 1875

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128. Antonina anceps Green, 1914

Distribution: Maharashtra (Probably Pune).

129. Antonina graminis (Maskell, 1897)

Distribution: Bihar, Delhi, Karnataka, Odisha, Uttar Pradesh and Tamil Nadu.

130. Antonina maritime Green, 1922

Distribution: Tamil Nadu (Coimbatore) and West Bengal.

131. Antonina zonata Green, 1919

Distribution: Bihar and Tamil Nadu (Coimbatore).

53. Genus: Chaetococcus Maskell, 1898

132. Chaetococcus bambusae (Maskell, 1893)


Family: Aclerdidae

54. Genus: Aclerda Signoret, 1874

Aclerda sp.

Distribution: Odisha (Bhubaneswar, Taptapani).

133. Aclerda distorta Green, 1909


134. Aclerda inermis Ayyar, 1936


135. Aclerda ischaemum Ayyar, 1930

Distribution: Tamil Nadu (Coimbatore) and Uttar Pradesh.

136. Aclerda tokionis (Cockerell, 1896)

Distribution: Bihar, Odisha and Tamil Nadu (Salem).

Family: Asterolecaniidae Subfamily: Asterolecaniinae

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55. Genus: Asterolecanium Targioni-Tozzetti, 1868

Asterolecanium sp.

Distribution: Tripura (Agartala) and West Bengal.

137. Asterolecanium longum Green, 1909


138. Asterolecanium minutum Takahashi, 1930


139. Asterolecanium pseudomilliaris Green, 1922

Distribution: Assam and West Bengal.

140. Asterolecanium tenuissimum Green, 1909

Distribution: Kerala (Malabar, Dhoney valley).

56. Genus: Bambusaspis Cockerell, 1902

141. Bambusaspis bambusae (Boisduval, 1869)


Family: Cerococcidae

57. Genus: Asterococcus Borchsenius, 1960

142. Asterococcus ramakrishnai Lambdin, 1983

Distribution: Tamil Nadu (Coimbatore), Maharashtra (Mumbai).

58. Genus: Cerococcus Comstock, 1882

Cerococcus sp.

Distribution: Uttar Pradesh (Aligarh).

143. Cerococcus bryoides (Maskell, 1894)

Distribution: Tamil Nadu (Chennai).

144. Cerococcus indicus (Maskell, 1897)

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Distribution: Andhra Pradesh, Bihar, Karnataka, Maharashtra (Mumbai), Odisha and West Bengal.

145. Cerococcus ornatus Green, 1909


Family: Lecanodiaspididae

59. Genus: Anomalococcus Green, 1902

146. Anomalococcus indicus Ayyar, 1930

Distribution: Karnataka, Odisha and Tamil Nadu.

60. Genus: Cosmococcus Borchsenius, 1959

147. Cosmococcus erythrinae Borchsenius, 1959


61. Genus: Stictacanthus Lambdin & Kosztarab, 1973

148. Stictacanthus azadirachtae Green, 1909

Distribution: South India and Uttar Pradesh.

Family: Coccidae Subfamily: Ceroplastinae

62. Genus: Ceroplastes Gray, 1928

Ceroplastes sp.

Distribution: Delhi, Odisha and Tripura.

149. Ceroplastes actiniformes Green, 1896

Distribution: Andhra Pradesh, Bihar, Maharashtra, Odisha, Uttar Pradesh, Tamil Nadu and West Bengal

150. Ceroplastes ajmerensis (Avasthi & Shafee, 1979)

Distribution: Rajasthan (Ajmer).

151. Ceroplastes alami Avasthi & Shafee, 1986

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Distribution: Uttar Pradesh (Aligarh) and Tamil Nadu (Coimbatore).

152. Ceroplastes ceriferus (Fabricius, 1798)

Distribution: Bihar, Uttar Pradesh, Himachal Pradesh, Maharashtra (Mumbai), Madhya Pradesh, Tamil Nadu (Coimbatore, Tanjore) and West Bengal.

153. Ceroplastes floridensis Comstock, 1881

Distribution: Assam, Bihar, Uttar Pradesh, Himachal Pradesh, Karnataka, Maharashtra (Mumbai), Madhya Pradesh, Odisha, Tamil Nadu (Coimbatore, Tanjore) and West Bengal.

154. Ceroplastes pseudoceriferus (Fabricius, 1798)

Distribution: Bihar, Delhi, Odisha and Uttar Pradesh.

155. Ceroplastes rubens Maskell, 1893

Distribution: Andhra Pradesh (Chicacole), Assam, Kerala, Tamil Nadu (Coimbatore) and West Bengal.

156. Ceroplastes rusci Signoret, 1972

Distribution: Uttarakhand (Dehra Dun).

63. Genus: Vinsonia Signoret, 1972

157. Vinsonia stellifera Westwood, 1871

Distribution: Andhra Pradesh, Karnataka, Kerala, Uttar Pradesh, Maharashtra (Mumbai), Odisha, Tamil Nadu and West Bengal.

Subfamily: Coccinae Tribe: Coccini

64. Genus: Akermes Cockerell, 1902

158. Akermes montanus (Green, 1908)

Distribution: Himachal Pradesh.

65. Genus: Ceronema Maskell, 1895

159. Ceronema koebeli Green, 1909


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66. Genus: Coccus Linnaeus, 1758

Coccus spp.

Distribution: Andhra Pradesh, Karnataka, Tripura, Uttar Pradesh and Odisha.

160. Coccus acutissimus Green, 1896

Distribution: Kerala (Malabar) and Tamil Nadu (Coimbatore).

161. Coccus adersi (Newstead, 1917), Ali, 1971


162. Coccus almoraensis Avasthi & Shafee, 1984

Distribution: Bihar and Uttar Pradesh (Almora).

163. Coccus discrepans Green, 1904

Distribution: Andhra Pradesh, Assam, Bihar and Kerala.

164. Coccus gymnospori Sanders, 1909

Distribution: Andhra Pradesh and Maharashtra (Pune).

165. Coccus hesperidium Linnaeus, 1758

Distribution: Andhra Pradesh, Assam, Bihar, Goa, Gujarat, Karnataka, Kerala, Odisha, Jammu & Kashmir, Tripura and West Bengal.

166. Coccus kosztarabi Avasthi & Shafee, 1984


167. Coccus latioperculatum Green, 1922


168. Coccus longulus (Douglas, 1887)

Distribution: Andhra Pradesh, Assam, Karnataka and Odisha.

169. Coccus ophiorrhizae Green, 1896

Distribution: Andhra Pradesh (Kurnool).

170. Coccus ramakrishnae (Ayyar, 1919)

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Distribution: Andhra Pradesh and Himachal Pradesh.

171. Coccus viridis (Green, 1889)

Distribution: Assam, Karnataka, Kerala, Odisha, Tamil Nadu and Tripura.

172. Coccus watti (Green, 1900)

Distribution: Assam, Meghalaya and West Bengal.

173. Coccus radicicola Green, 1921


174. Coccus olivaceum Green, 1922


67. Genus: Eriochiton Maskell, 1887

175. Eriochiton tessellates Signoret, 1873


68. Genus: Eulecanium Cockerell, 1893

Eulecanium sp.


176. Eulecanium capreae (Linnaeus, 1767)

Distribution: Jammu & Kashmir.

177. Eulecanium coryli (Linnaeus, 1758)

Distribution: Jammu & Kashmir and Uttar Pradesh.

69. Genus: Hemilecanium Newstead, 1906

178. Hemilecanium imbricans (Green, 1903)

Distribution: Tamil Nadu (Coimbatore, Nilgiris) and Karnataka (Mysore, Belur).

70. Genus: Maacoccus Tao et al., 1983

179. Maacoccus bicruciatus Tao et al., 1983

23


71. Genus: Marsipococcus Cockerell & Bueker, 1930

180. Marsipococcus marsupiale (Green, 1904)

Distribution: Kerala and Tamil Nadu.

72. Genus: Paralecanium Cockerell & Parrot, 1899

181. Paralecanium expansum expansum (Green, 1896)

Distribution: Karnataka and Tamil Nadu (Chennai).

182. Paralecanium maritimum Fernald, 1903


73. Genus: Parthenolecanium Sulc, 1908

183. Parthenolecanium persicae Fabricius, 1776


74. Genus: Saissetia Deplanche, 1859

Saissetia sp.

Distribution: Odisha (Bhubaneswar, Baripada, Puri ).

184. Saissetia coffeae (Walker, 1852)

Distribution: Andhra Pradesh, Assam, Bihar, Goa, Karnataka, Maharashtra, Manipur, Odisha, Tamil Nadu and West Bengal.

185. Saissetia crassum Ali, 1971


186. Saissetia depressa King, 1902


187. Saissetia filicum Fernald, 1903

Distribution: Karnataka, Odisha and Tamil Nadu.

188. Saissetia miranda De Lotto, 1970

24

Distribution: Tamil Nadu (Palani hills).

189. Saissetia nigra King, 1902

Distribution: Assam, Bihar, Jammu & Kashmir, Tamil Nadu, Uttar Pradesh and West Bengal.

190. Saissetia lieae Cockerell, 1901

Distribution: Assam (Tocklai).

191. Saissetia privigna De Lotto, 1965

Distribution: Andhra Pradesh (Guntur).

75. Genus: Taiwansaissetia Tao et al., 1983

192. Taiwansaissetia formicarii Tao et al., 1983

Distribution: Assam and Karnataka.

Tribe: Pulvinariini

76. Genus: Chloropulvinaria Borchsenius, 1952

193. Chloropulvinaria cellulose Varshney, 1985

Distribution: Bihar, Uttar Pradesh and West Bengal.

194. Chloropulvinaria floccifera (Westwood, 1870)


195. Chloropulvinaria psidii Borchsenius, 1957

Distribution: Andhra Pradesh, Bihar, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu, Tripura and West Bengal.

196. Chloropulvinaria venkatramani Varshney, 1992 [nomen novum]


77. Genus: Micropulvinaria Hodgson, 1968

197. Micropulvinaria maxima thespesiae (Green); Varshney, 1992 [n. comb.]

Distribution: Andhra Pradesh and Tamil Nadu.

78. Genus: Protopulvinaria Cockerell, 1894

25

198. Protopulvinaria mangiferae Steinweden, 1929

Distribution: Bihar, Tamil Nadu and West Bengal.

79. Genus: Pulvinaria Targioni Tozzetti, 1866

Pulvinaria spp.

Distribution: Bihar and Odisha

199. Pulvinaria aligarhensis Avasthi & Shafee, 1980

Distribution: Uttar Pradesh (Aligarh).

200. Pulvinaria burkilli Green, 1908


201. Pulvinaria indica Avasthi & Shafee, 1980


202. Pulvinaria obscura Newstead, 1894


80. Genus: Takahashia Cockerell, 1986

203. Takahashia japonica (Cockerell, 1896)


Subfamily: Filipiinae

81. Genus: Cardiococcus Cockerell, 1903

204. Cardiococcus bivalvata (Green, 1903)

Distribution: Andhra Pradesh and Tamil Nadu.

82. Genus: Ceroplastodes Cockerell, 1893

Ceroplastodes sp.

Distribution: Bihar, Jammu & Kashmir, Odisha and Uttar Pradesh.

205. Ceroplastodes cajani Cockerell, 1900

26

Distribution: Andhra Pradesh, Bihar, Himachal Pradesh, Kerala, Punjab, Tamil Nadu and West Bengal.

206. Ceroplastodes chiton Green, 1909

Distribution: Himachal Pradesh, Uttar Pradesh and West Bengal.

83. Genus: Dicyphoccus Borchsenius, 1959

207. Dicyphococcus castilloae Borchsenius, 1959


84. Genus: Inglisia Maskell, 1879

208. Inglisia chelonioides Green, 1903

Distribution: Kerala, Maharashtra (Shiroda) and Tamil Nadu (Coimbatore).

85. Genus: Lecanopsis Targioni Tozzetti, 1868

209. Lecanopsis ceylonica Green, 1922


86. Genus: Metaceronema Takahashi, 1955

210. Metaceronema japonica (Maskell, 1897)

Distribution: West Bengal (Darjeeling).

Superfamily: Diaspidoidea Family: Diaspididae

Subfamily: Parlatoriinae Tribe: Leucaspidini

87. Genus: Leucaspis Signoret, 1869

211. Leucaspis manii (Borchsenius, 1964)

Distribution: Meghalaya.

212. Leucaspis riccae Targioni Tozzetti, 1881

Distribution: Andhra Pradesh (Hyderabad).

88. Genus: Lopholeucaspis Balachowsky, 1953

27

Lopholeucaspis sp.

Distribution: Karnataka (Anekal).

213. Lopholeucaspis cockerelli (Grandpre & De Charmoy, 1899)


214. Lopholeucaspis excoecariae Borchsenius, 1964


215. Lopholeucaspis japonica (Cockerell, 1897)

Distribution: Karnataka (Vasanthapura).

216. Lopholeucaspis menoni Borchsenius, 1964

Distribution: Rajasthan, Uttar Pradesh and West Bengal.

89. Genus: Radionaspis Ferris, 1942

217. Radionaspis indica (Marlatt, 1908)

Distribution: Maharashtra (Pune).

90. Genus: Salicicola Lindinger, 1905

218. Salicicola indiaeorientalis (Lindinger, 1911)

Distribution: Himachal Pradesh (Shimla).

Tribe: Parlatoriini

91. Genus: Apteronidia Berlese, 1895

219. Apteronidia blanchardi (Targioni Tozzetti, 1892)

Distribution: Andhra Pradesh (Hyderabad).

92. Genus: Cryptoparlatoria Lindinger, 1905

220. Cryptoparlatoria pini Takahashi, 1938

Distribution: Himachal Pradesh (Manali).

93. Genus: Genaparlatoria MacGillivray, 1921

28

221. Genaparlatoria pseudaspidiotus (Lindinger, 1905)

Distribution: Andhra Pradesh (Godavari Dist.).

94. Genus: Greeniella Cockerell, 1897

222. Greeniella cornigera (Green, 1896)


223. Greeniella dentata (Lindinger, 1911)


224. Greeniella ferreae (Rutherford, 1914)


225. Greeniella viridis (Lindinger, 1911)


95. Genus: Gymnaspis Newstead, 1898

Gymnaspis sp.

Distribution: Kerala (Kattuthurai).

226. Gymnaspis affinis Ayyar, 1930

Distribution: Tamil Nadu (Tirunelveli).

227. Gymnaspis cassida Hall & Williams, 1962


228. Gymnaspis ficus Green, 1919


229. Gymnaspis mangifera Ayyar, 1924

Distribution: Tamil Nadu (Tirunelveli)

230. Gymnaspis ramakrishnai Green, 1919


29

96. Genus: Neoleucaspis Green, 1926

231. Neoleucaspis parallela Green, 1926


97. Genus: Parlaspis McKenzie, 1945

Parlaspis spp.

Distribution: Karnataka and Kerala.

232. Parlaspis papillosa (Green, 1919)

Distribution: Kerala (Palghat).

98. Genus: Parlatorearum Lindinger, 1911

Parlatorearum sp.

Distribution: Andhra Pradesh (Cuddapah).

99. Genus: Parlatoreopsis Lindinger, 1911

233. Parlatoreopsis chinensis (Marlatt, 1908)

Distribution: Tamil Nadu (Coonoor, Coimbatore).

100. Genus: Parlatoria Targioni Tozzetti, 1868

Parlatoria sp.

Distribution: Karnataka and Tamil Nadu.

234. Parlatoria artocarpi Green, 1919

Distribution: Kerala (Peria Ghat).

235. Parlatoria atalantiae Green, 1905


236. Parlatoria boycei MacKenzie, 1952


237. Parlatoria camelliae Comstock, 1883

30


238. Parlatoria cinerea Hadden in Doane & Hadden, 1909

Distribution: West Bengal and South India.

239. Parlatoria crotonis Douglas, 1887

Distribution: Bihar

240. Parlatoria crypta McKenzie, 1943

Distribution: Andhra Pradesh, Delhi, Karnataka, Maharashtra and Punjab.

241. Parlatoria marginalis McKenzie, 1945

Distribution: Delhi.

242. Parlatoria morrisoni McKenzie, 1943


243. Parlatoria mytilaspiformis Green, 1899


244. Parlatoria oleae (Colvee, 1880)

Distribution: Bihar, Kashmir, Maharashtra, Rajasthan and West Bengal.

245. Parlatoria orientalis Rao, 1953


246. Parlatoria pergandii Comstock, 1881

Distribution: Bihar, Karnataka, Kerala and Tamil Nadu (Coimbatore, Mundathorai, Tirunelvelly)

247. Parlatoria proteus (Curtis, 1843)

Distribution: Karnataka, Tamil Nadu (Coimbatore, Nilgiris), Tripura and West Bengal.

248. Parlatoria vateriae Green, 1919

Distribution: Kerala (Travancore, Kollam).

249. Parlatoria ziziphi (Lucas, 1853)

31

Distribution: Tamil Nadu (Coimbatore, Ramanathapuram, Tanjore) and West Bengal.

101. Genus: Tamilparla Takagi, 1987

250. Tamilparla smilacis Takagi, 1987

Distribution: Tamil Nadu (Coonoor, Nilgiris).

Subfamily: Odonaspidinae Tribe: Rugaspidiotini

102. Genus: Mangaspis Takagi & Kondo, 1997

251. Mangaspis Bengalurunsis Takagi & Kondo, 1997


103. Genus: Natalaspis MacGillivray, 1921

252. Natalaspis simplex (Green, 1899); Ben-Dov & Takagi, 1974


104. Genus: Ramachandraspis Rao, 1953

253. Ramachandraspis fenestrata Rao, 1953


Tribe: Odonaspidini

105. Genus: Berlesaspidiotus MacGillivray, 1921

254. Berlesaspidiotus crenulatus Ben-Dev, 1988

Distribution: Tamil Nadu (Chennai, Palani).

106. Genus: Froggattiella Leonardi, 1900

255. Froggattiella penicillata (Green, 1905); Rutherford, 1915


107. Genus: Odonaspis Leonardi, 1897

Odonaspis spp.


32

256. Odonaspis paucipora Ben-Dov, 1988

Distribution: West Bengal (Kolkata).

257. Odonaspis saccharicaulis (Zehntner, 1897); Fernald, 1903

Distribution: Karnataka, Odisha and Tamil Nadu (Coimbatore).

Subfamily: Aspidiotinae Tribe: Aspidiotini

108. Genus: Abgrallaspis Balachowsky, 1948

258. Abgrallaspis cyanophylli Balachowsky, 1948

Distribution: Karnataka (Bengaluru Kerala and Tamil Nadu (Coimbatore).

109. Genus: Aonidia Targioni - Tozzetti, 1868

259. Aonidia crenulata Green, 1900


260. Aonidia indica Green, 1919


261. Aonidia loranthi Green, 1896


262. Aonidia tentaculata Green, 1919


110. Genus: Aonidiella Berlese & Leonardi, 1895

263. Aonidiella andersoni (Laing, 1929); McKenzie, 1938


264. Aonidiella aurantii (Maskell, 1879)

Distribution: Andhra Pradesh, Assam, Bihar, Delhi, Kanataka, Kerala, Odisha, Punjab and Tamil Nadu.

265. Aonidiella citrine Nel, 1933

33


266. Aonidiella comperei McKenzie, 1937

Distribution: Maharashtra (Mumbai, Colaba).

267. Aonidiella inortata McKenzie, 1938


268. Aonidiella misrai Laing, 1929


269. Aonidiella orientalis McKenzie, 1937

Distribution: Bihar, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Punjab, Tamil Nadu, Tripura and West Bengal.

111. Genus: Aspidiella Leonardi, 1898

270. Aspidiella hartii (Cockerell, 1895)

Distribution: Maharashtra (Pune) and Tamil Nadu (Coimbatore).

271. Aspidiella sacchari MacGillivray, 1921 [comb. new.]


112. Genus: Aspidiotus Bouche, 1833

Aspidiotus spp.

Distribution: Andhhra Pradesh, Odisha and Karnataka.

272. Aspidiotus destructor Signoret, 1869

Distribution: Andhra Pradesh, Kerala, Jammu & Kashmir, Maharashtra (Mumbai), Rajasthan, Tripura, West Bengal and Lakshadweep Is.

273. Aspidiotus hederae (Vallot, 1829)


274. Aspidiotus pandani (Boisduval, 1868)


34

113. Genus: Borchseniaspis Zahradnik, 1959

275. Borchseniaspis palmae Zahradnik, 1959


114. Genus: Brainaspis MacGillivray, 1921

276. Brainaspis kellyi MacGillivray, 1921


115. Genus: Clavaspis MacGillivray, 1921

Clavaspis sp.


116. Genus: Chrysomphalus Ashmead, 1880

277. Chrysomphalus aonidum (Linnaeus, 1758)

Distribution: Andhra Pradesh, Assam, Bihar, Kerala, Jammu & Kashmir, Maharashtra (Mumbai), Madhya Pradesh, Gujarat, Tamil Nadu, Tripura and West Bengal.

278. Chrysomphalus dictyospermi (Morgan, 1889)

Distribution: Andhra Pradesh, Assam, Meghalaya, Tamil Nadu and Tripura.

279. Chrysomphalus diversicolor (Green, 1923)

Distribution: Karnataka and Tamil Nadu.

117. Genus: Cryptopalratoreopsis Borchsenius, 1947

280. Cryptopalratoreopsis euphorbiae Hall & Williams, 1962


281. Cryptopalratoreopsis longispina (Takahashi, 1939)


282. Cryptopalratoreopsis spinosissima (Lindinger, 1911)

Distribution: India (Central India).

283. Cryptopalratoreopsis targionipsis (Lindinger, 1911)

35


118. Genus: Diaspidiotus Berlese & Leonardi, 1896

284. Diaspidiotus pseudocamelliae Borchsenius, 1966


119. Genus: Duplaspidiotus MacGillivray, 1921

285. Duplaspidiotus tesseratus (Grandpre & de Charmoy, 1899)

Distribution: Karnataka and Maharashtra (Mumbai, Lonavala).

120. Genus: Hemiberlesia Cockerell, 1897

286. Hemiberlesia lataniae (Signoret, 1869)

Distribution: Assam, Odisha, Karnataka, Maharashtra (Mumbai), Tamil Nadu, Tripura and West Bengal.

287. Hemiberlesia rapax (Comstock, 1881)

Distribution: Assam, Odisha, Karnataka, Tamil Nadu, Tripura and West Bengal.

121. Genus: Lindingaspis MacGillivray, 1921

Lindingaspis sp.


288. Lindingaspis floriadana Ferris, 1942


289. Lindingaspis fusca McKenzie, 1943

Distribution: Maharashtra (Pune) and South India.

290. Lindingaspis rossi (Maskell, 1891)

Distribution: Andhra Pradesh, Maharashtra (Pune) and Madhya Pradesh.

122. Genus: Melanaspis Cockerell, 1897

Melanaspis sp.


36

291. Melanaspis glomerata (Green, 1903)

Distribution: Andhra Pradesh, Bihar, Odisha, Gujarat, Tamil Nadu and Uttar Pradesh.

292. Melanaspis smilacis (Comstock, 1883)


123. Genus: Metaspidiotus Takagi, 1957

293. Metaspidiotus sp.

Distribution: Karnataka and Goa.

124. Genus: Morganella Cockerell, 1897

294. Morganella cueronsis (Cockerell, 1899)

Distribution: Tamil Nadu and South India.

295. Morganella longispina (Morgan, 1889)


125. Genus: Neofurcaspis Green, 1926

296. Neofurcaspis andamanensis Green, 1926

Distribution: Andaman Is.

126. Genus: Octaspidiotus MacGillivray, 1921

297. Octaspidiotus stauntoniae (Takahashi, 1933)

Distribution: Karnataka and Goa.

298. Octaspidiotus australiensis (Kuwana, 1931)


299. Octaspidiotus calophylli var. symplocos (Green, 1924)

Distribution: Tamil Nadu (Nilgiris, Coonoor, Kotagiri).

300. Octaspidiotus tamarindi (Green, 1919)

Distribution: Karnataka (Nandi) and Tamil Nadu (Coimbatore).

37

301. Octaspidiotus tripurensis Takagi, 1984

Distribution: Tripura (Agartala).

127. Genus: Parrottia MacGillivray, 1921

Parrottia sp.

Distribution: Karnataka (Devarayanadurga).

302. Parrottia moorei (Green, 1896)


128. Genus: Pseudaonidia Cockerell, 1897

Pseudaonidia spp.


303. Pseudaonidia duplex (Cockerell, 1896)


304. Pseudaonidia trilobitiformis (Green, 1896)

Distribution: Andhra Pradesh, Bihar, Kerala, Tamil Nadu and West Bengal.

129. Genus: Quadraspidiotus MacGillivray, 1921

305. Quadraspidiotus perniciosus (Comstock, 1881)

Distribution: Andhra Pradesh, Bihar, Himachal Pradesh, Jammu & Kashmir, Karnataka, Kerala, Maharashtra (Mahabaleshwar), Meghalaya, Odisha, Punjab, Sikkim, Tamil Nadu, Uttar Pradesh and West Bengal.

130. Genus: Semelaspidus MacGillivray, 1921

306. Semelaspidus artocarpi (Green, 1896)

Distribution: Karnataka (Bengaluru, Coorg) and Maharashtra (Mahabaleshwar).

131. Genus: Temnaspidiotus MacGillivray, 1921

307. Temnaspidiotus excisus (Green, 1896)

Distribution: South India and Tripura (Agartala).

38

Subfamily: Diaspidinae Tribe: Protodiaspidini

132. Genus: Anaimalaia Takagi, 1995

308. Anaimalaia scabra Takagi, 1995

Distribution: Tamil Nadu (Anaimalai Hills).

Tribe: Kuwanaspidini

133. Genus: Kuwanaspis MacGillivray, 1921

309. Kuwanaspis annandalei (Green, 1919)

Distribution: Bihar (Parasnath Hills).

134. Genus: Xiphuraspis Borchsenius & Williams, 1963

310. Xiphuraspis spiculata Borchsenius & Williams, 1963

Distribution: Kerala (N. Malabar).

135. Genus: Coronaspis MacGillivray, 1921

311. Coronaspis malabarica Takagi, 1999

Distribution: Kerala (Periyar Tiger Reserve) and Tamil Nadu (Anaimalai Hills).

Tribe: Lepidosaphidini

136. Genus: Acanthomytilus Borchsenius, 1947

312. Acanthomytilus sacchari (Hall, 1923)

Distribution: Bihar, Tamil Nadu (Coimbatore), Uttar Pradesh and West Bengal.

313. Acanthomytilus vermiformis (Takahashi, 1931)

Distribution: Kerala (Dhony Valley).

137. Genus: Andaspis MacGillivray, 1921

Andaspis sp. (nr. leucophloeae Rao)


314. Andaspis artocarpi Borchsenius, 1967

39


315. Andaspis betulae (Borchsenius, 1967)

Distribution: Meghalaya (Shilong).

316. Andaspis laingi Rao, 1952

Distribution: Tamil Nadu (Coimbatore, Makkalidurga).

317. Andaspis leucophleae Rao, 1952

Distribution: Tamil Nadu (Coimbatore, Chennai) and Madhya Pradesh.

318. Andaspis meliae (Green, 1919)

Distribution: Andhra Pradesh, Karnataka (Mandya), Tamil Nadu (Coimbatore) and West Bengal (Kolkata).

319. Andaspis numerate Brimblecombe, 1959


320. Andaspis retrusa (Green, 1919)


138. Genus: Aonidomytilus Leonardi, 1903

321. Aonidomytilus albus (Cockerell, 1893)

Distribution: Karnataka (Bengaluru), Tamil Nadu (Coimbatore, Salem, Puttukottai) and Puducherry.

139. Genus: Cornuaspis MacGillivray, 1921

322. Cornuaspis beckii (Newmann, 1869)

Distribution: Assam, Karnataka, Kerala and West Bengal.

323. Cornuaspis piperis (Green, 1908)

Distribution: Western Ghats; Kerala (N. Malabar, Wynaad, Travancore).

140. Genus: Eucornuaspis Borchsenius, 1963

324. Eucornuaspis pinnaeformis Borchsenius, 1966

40


325. Eucornuaspis pseudomachili Borchsenius, 1964


141. Genus: Howardia Berlese & Leonardi, 1896

326. Howardia biclavis (Comstock, 1883)

Distribution: Karnataka, Kerala (Wynaad) and Tamil Nadu (Chennai, Anaimalai Hills).

142. Genus: Insulaspis Mamet, 1950

327. Insulaspis citrine Borchsenius, 1964

Distribution: New Delhi.

328. Insulaspis gloverii (Packard, 1869)

Distribution: Bihar, Karnataka and West Bengal.

329. Insulaspis mcgregori (Banks, 1906)

Distribution: Kerala (Thiruvananthapuram Dist.).

330. Insulaspis pallidula Williams, 1969

Distribution: Andhra Pradesh (Godavari Dist.).

331. Insulaspis tapleyi (Williams, 1960); Borchsenius, 1963

Distribution: Karnataka and Uttar Pradesh.

332. Insulaspis tokionis (Kuwana, 1902); Borchsenius, 1963


143. Genus: Lepidosaphes Shimer, 1868

333. Lepidosaphes cornutus Green in Ayyar, 1937

Distribution: Kerala and Tamil Nadu (North Arcot Dist., South Arcot Dist., Tanjore, Salem).

334. Lepidosaphes malicola Borchsenius, 1947

Distribution: Jammu & Kashmir (Anantnag).

41

335. Lepidosaphes ulmi (Linnaeus, 1758)


144. Genus: Neopinnaspis McKenzie, 1949

336. Neopinnaspis travancorensis (Lindinger, 1911)


145. Genus: Velataspis Ferris, 1937

337. Velataspis serrulata Ganguli, 1958


Tribe: Diaspidini

146. Genus: Africaspis MacGillivray, 1921

338. Africaspis chionaspiformis (Newstead, 1910)

Distribution: Tamil Nadu (Coimbatore, Tirunelveli).

147. Genus: Aulacaspis Cockerell, 1893

Aulacaspis spp.

Distribution: Karnataka and Kerala.

339. Aulacaspis gudalura Ferris, 1956

Distribution: Tamil Nadu (Nilgiris).

340. Aulacaspis hedyotidis Takagi, 1985


341. Aulacaspis herbae Hall, 1946


342. Aulacaspis litzeae (Green, 1896)


343. Aulacaspis loranthi Cockerell, 1902

42

Distribution: Bihar (Parasnath Hill).

344. Aulacaspis madiunensis (Zehtner, 1898)

Distribution: Andhra Pradesh and Tamil Nadu (Chengalpattu, North & South Arcot Dist., Tanjore, Tiruchirapalli).

345. Aulacaspis malayala Varshney, 2002

Distribution: Kerala (Periyar Tiger Reserve).

346. Aulacaspis mangiferae (Newstead, 1908)

Distribution: Andhra Pradesh, Karnataka and Tamil Nadu (Tirunelveli).

347. Aulacaspis orientalis (Green, 1922)

Distribution: Tamil Nadu (Courtallum, Tirunelveli).

348. Aulacaspis rosae (Bouche, 1833)

Distribution: Tamil Nadu (Courtallum, Tirunelveli).

349. Aulacaspis tubercularis (Newstead, 1908)

Distribution: Karnataka, Kerala and Uttar Pradesh.

350. Aulacaspis vitis (Green, 1896)

Distribution: Assam, Bihar, Himachal Pradesh, Odisha, Karnataka and Tamil Nadu.

148. Genus: Chionaspis Signoret, 1869

Chionaspis sp.


351. Chionaspis engeddensis Bodenheimer, 1924

Distribution: Punjab.

352. Chionaspis furfura (Fitch, 1857)

Distribution: Jammu & Kashmir.

353. Chionaspis himalaica Takagi, 1985


43

354. Chionaspis ramakrishnai Rao, 1953


355. Chionaspis salicis (Linnaeus, 1758)

Distribution: Himalayas and Jammu & Kashmir.

149. Genus: Contigaspis MacGillivray, 1921

356. Contigaspis coimbatorensis Borchsenius & Williams, 1963

Distribution: Karnataka and Tamil Nadu (Coimbatore).

357. Contigaspis indigoferae (Hall, 1928)

Distribution: Tamil Nadu (Hosur).

150. Genus: Dentachionaspis MacGillivray, 1921

358. Dentachionaspis centripetalis Rao, 1953

Distribution: Jammu & Kashmir (Srinagar).

151. Genus: Diaspis Costa, 1828

Diaspis sp.

Distribution: Odisha and Tamil Nadu.

359. Diaspis boisduvali Signoret, 1869

Distribution: Tamil Nadu (Nilgiris).

360. Diaspis echinocacti (Bouche, 1833)

Distribution: Karnataka, Kerala, Maharashtra (Pune), Odisha, Tamil Nadu and Uttar Pradesh.

152. Genus: Duplachionaspis MacGillivray, 1921

361. Duplachionaspis divergens (Green, 1899)

Distribution: Kerala, Punjab and Tamil Nadu.

153. Genus: Fiorinia Targioni- Tozzetti, 1868

Fiorinia spp.

44

Distribution: Andhra Pradesh and Tripura.

362. Fiorinia fioriniae (Targioni Tozzetti, 1867)


363. Fiorinia fletcheri Laing, 1929


364. Fiorinia frontecontracta Green, 1919

Distribution: Maharashtra (Mumbai).

365. Fiorinia himalaica Takagi, 1975

Distribution: Himachal Pradesh and Uttar Pradesh.

366. Fiorinia japonica Kuwana, 1902

Distribution: West Bengal (Kolkata).

367. Fiorinia multipora Lindinger, 1911


368. Fiorinia pakistanensis Ahmad, 1970

Distribution: Kashmir.

369. Fiorinia plana Green, 1919

Distribution: Tamil Nadu (Coimbatore, Tirunelveli).

370. Fiorinia proboscidaria Green, 1920


371. Fiorinia sapindi Green, 1919

Distribution: Maharashtra (Pune).

372. Fiorinia theae Green, 1900

Distribution: Assam, North India and West Bengal (Darjeeling).

154. Genus: Greenaspis MacGillivray, 1921

45

373. Greenaspis decurvata (Green, 1903)

Distribution: Kerala, Odisha, Tamil Nadu and West Bengal.

374. Greenaspis elongata (Green, 1896)


155. Genus: Lineaspis MacGillivray, 1921

375. Lineaspis caroli (Green, 1919)


156. Genus: Myrtaspis Takagi, 1999

376. Myrtaspis syzygii (Takagi, 1985)


157. Genus: Pinnaspis Cockerell, 1892

Pinnaspis sp.

Distribution: Karnataka (Thalghatpura, Pannayanpalaya, Shivganga).

Pinnaspis sp. [nr. strachani (Cooley, 1899)].

Distribution: West Bengal (Barrackpore) and Karnataka (Gokarna).

377. Pinnaspis aspidistrae (Signoret, 1869)

Distribution: Andhra Pradesh, Karnataka (Shivganga), Kerala (Cochin, Malabar), Madhya Pradesh, Maharashtra (Janjira Estate), Tamil Nadu (Coimbatore, Nilgiris) and Tripura.

378. Pinnaspis buxi (Bouche, 1851)


379. Pinnaspis dracaenae (Cooley, 1899)

Distribution: Kerala (South Malabar, Vadanapalle).

380. Pinnaspis exercitata (Green, 1896)

Distribution: Karnataka (Coorg), Odisha, Tamil Nadu (Coimbatore, Chennai) and West Bengal.

381. Pinnaspis fici (Green, 1908)

46


382. Pinnaspis minima (Green, 1908)


383. Pinnaspis strachani (Cooley, 1899)

Distribution: Andhra Pradesh, Bihar, Karnataka (Bengaluru, Devanahally, Bellari, Vasanthapura, Hoskote, Nandi, Koramangala), Kerala (Kondayur), Jammu & Kashmir, Tamil Nadu (Coimbatore, Nilgiris, Marudamali) and West Bengal.

384. Pinnaspis theae (Maskell, 1891)

Distribution: North India (Assam) and West Bengal (Darjeeling).

385. Pinnaspis uniloba (Kuwana, 1909)

Distribution: Assam and Tamil Nadu (Chennai, Pudukottai).

158. Genus: Poliaspis Maskell, 1880

386. Poliaspis cycadis Comstock, 1883

Distribution: Maharashtra (Kolhapur).

159. Genus: Pseudaulacaspis MacGillivray, 1921

Pseudaulacaspis sp.

Distribution: Karnataka (Kodigehalli, Namadachiluve).

387. Pseudaulacaspis barberi (Green, 1908)

Distribution: Andhra Pradesh (Visakhapatnam, Alamanda), Karnataka (Bengaluru, Bellari) and Tamil Nadu (Tanjore Dist.).

388. Pseudaulacaspis cockerelli (Cooley, 1897)

Distribution: Andhra Pradesh (Godavari Dist.), Karnataka (Bengaluru, Lal Bagh, Koramangala), Kerala (Malabar, Wynaad, Kondayur), Maharashtra (Pune), Tamil Nadu (Hosur, Nilgiris) and West Bengal (Kolkata).

389. Pseudaulacaspis gynandropsidis (Green, 1922)

Distribution: Karnataka (Bengaluru, Jalahalli).

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390. Pseudaulacaspis manni (Green, 1907)

Distribution: Assam, Himachal Pradesh (Kangra), Tamil Nadu (Hosur) and West Bengal (Kolkata, Darjeeling).

391. Pseudaulacaspis megaloba (Green, 1899)


392. Pseudaulacaspis pentagona pentagona (Targioni- Tozzetti, 1886)

Distribution: Assam (Maungep, Sonapari), Karnataka (Byadarahalli, Vasanthapura), Meghalaya (Shilong) and Uttar Pradesh (Chaubattia).

393. Pseudaulacaspis pentagona theae (Maskell, 1896)


394. Pseudaulacaspis polygoni (Green, 1899)

Distribution: Karnataka (Mallandur).

395. Pseudaulacaspis pusa (Rao, 1953)


396. Pseudaulacaspis subcorticalis (Green, 1905)

Distribution: Karnataka (Jayanthigrama, Kuppehalli, Kobbinahalli).

397. Pseudaulacaspis tenera (Green, 1922)

Distribution: Karnataka (Karkada).

398. Pseudaulacaspis varicosa (Green, 1896)

Distribution: Karnataka (Coorg, Sidapur) and Tamil Nadu (Mundathurai, Nilgiris, Tirunelveli).

399. Pseudaulacaspis venue (Menon & Khan, 1961)

Distribution: Himachal Pradesh (Kangra, Palampur).

160. Genus: Pseudoparlatoria Cockerell, 1892

400. Pseudoparlatoria parlatorioides (Comstock, 1883)


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161. Genus: Rutherfordia MacGillivray, 1921

401. Rutherfordia major (Cockerell, 1894)

Distribution: Kerala (Periyar).

162. Genus: Sinistraspis MacGillivray, 1921

402. Sinistraspis unilateralis (Newstead, 1913)

Distribution: Kerala (Thiruvananthapuram).

163. Genus: Tenuiaspis MacGillivray, 1921

403. Tenuiaspis minuta (Green, 1896)


164. Genus: Thysanofiorinia Balachowsky, 1954

404. Thysanofiorinia leei Williams, 1971

Distribution: Tripura (Agartala) and Uttarakhand (Dehra Dun).

405. Thysanofiorinia nephelii (Maskell, 1897)

Distribution: Karnataka (Coorg, Siddapur) and West Bengal (Krishnanagar).

165. Genus: Trullifiorinia Leonardi, 1906

406. Trullifiorinia geloniae (Green, 1900)

Distribution: Tamil Nadu (Ooty).

166. Genus: Unaspis MacGillivray, 1921

Unaspis sp.

Distribution: Karnataka (Nandi).

407. Unaspis acuminate (Green, 1896)

Distribution: Kerala (Chalikudi, Malabar, Peria Ghat) and Tamil Nadu (Madurai).

408. Unaspis atricolor (Green, 1922)

Distribution: Andhra Pradesh (Hyderabad) and Tamil Nadu (Coimbatore, Dharmapuri).

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409. Unaspis permutans (Green, 1899)

Distribution: Kerala (Malabar, Peria Pass).

REFERENCES

Varshney R. K. 1992. A Check list of Scale insects and mealy bugs of South Asia, (Part-1), Rec. zool. Surv. India, Occ. Paper, 139: 152. Varshney R. K. 2002. A Check list of Scale insects and mealy bugs of South Asia, (Part-2), Rec. zool. Surv. India, Occ. Paper, 191: 147. Williams D. J. 1978. The anomalous ant-attended mealy bugs (Homoptera: Pseudococcidae) of South-east Asia, Bull. Br. Mus. nat. (Ent.), 37 (1): 1-72. Y. Ben-Dev, C.J.Hodgson and D.R.Miller. 1997. Changes and Comments on the Taxonomy and Nomenclature of Some Taxa in the Families Coccidae, Eriococcidae and Pseudococcidae (Homoptera: Coccoidae). Phytoparasitica, 25(3): 199-206. Internet Source : http://www.sel.barc.usda.gov/SCALENET/SCALENET.HTM

scale insects and mealy bugs (insecta: homoptera: coccoidea)

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