rs469 si09 cephalorhynchus 1988

355
I n t r n a t i o n a l W h a l i n g  C o m m i s s i o n  i o l o g y  o  t h e  G e n u s  e p h  l o r h y n c h u s Ed it e d  b y R o b e r t  L . B r o w n e ll  J r. , U . S .  F is h  a n d W i ld li f e S e r v i c e  P O  o x  7 0  S a n S i m e o n C A  9 3 4 5 2  U S A a n d G re g o ry  P . D o n o v an I n t e r n a ti o n a l  W h a li n g  C o m m i s s i o n T h e  R e d H o u s e  S t a t i o n  R o a d  H is t o n , C a m b r id g e  C B 4  4 N P , U K R e p o r t s  o  t h e  I n t e r n a t i o n a l W h a l in g  C o m m i s s i o n S p e c i a l  I s s u e  

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inid
g
enera
in
the
South
ern Hemi
species.
Gray
lphinus fo r
 
umber ofcontinuing
ts, (ii) life-
meeting, pl
publication of a volume which would  in
clude
We h
ope that th e seventeen papers in cluded in this volum
e, ten
or
iginally  
presentative of th e   wide diversity
of research
are described: females are larger than  ma
les; adult males
  of their
flippers. The
d Hector s dolphin, which
has been kept only in its native
  New   Zeal
has the gre
offTierra del
Brown
Investigations .
waters
around
South
statement. Lockyer,
characteristics
of dolphins that were incidentally caught off Tierra del Fuego.
Details on
Islands
are
presented by Collett and Robineau. Bastida, Lichtschien and
Goodall examine the food habits of this dolphin from waters around Tierra del
Fuego.
Goodall
Two papers by Leatherwood
population estimates fo r C . commersonii in the
northeastern Strait
Cornell
captivity. Evans, Awbrey and
free-ranging Com merson s dolphins,
compare their signals with those
made
papers
papers are based
around
the Kerguelen Islands. The first paper described the external morphology
and
colour
(1984)
related
food
habits
details
on
osteology and osteometrical data. Robineau (1985a and b) described the
distribution of the species around the
Kerguelen
a
geographically
isolated
compared to South
American specimens. Abarnou,
Robineau and Michel
(1986) reported organochlorine pollutants (DDT, PCB and HCB) in samples
from these dolphins but
those found in
similar species from
Brownell and
the two type
a synonym of C. commersonii.
Section II deals with the previously poorly known Chilean dolphin, C .
eutropia.
Goodall
certain aspects of the
and present much new
information from specimens they
al.
clarify
the status of Phocaena obtusata which they believe is a specimen of
C . eutropia.
Best on
Best on
based
collected since 1970. Photographs of fresh whole animals
are presented for the
dolphins
species, all of
respect. The only other recent paper on this species, that
of Rice and Saayman
coloration
and
Hector's
reports
recent
observations of the species
and problems with incidental mortality in set gillnets. Dawson and Slooten
present results
distribution and abundance
individuals. In another paper, Slooten and Dawson provide information
on
caught
last
chapter,
dolphins and
has
our
as much as possible left the style and content to
the authors,
peer review system established for IWC publications. The research
presented here is the first overall examination of the genus Cephalorhynchus;
with time it will be, of course, revised and new information will continue to add
to our
species. With all the new
information on Cephalorhynchus available in this volume, it may be time to
revise the genus, emphasising what is known about the distribution, the
relationship to the Antarctic
items and the coloration and osteology of the four species.
Finally, we would
who
W. E. Evans, D. E.
Gaskin, M . Hall,
S. Hammond, R. J. Harrison, J. E.
Heyning, L. Jones, T. Kasuya, V. Lichtschein, C. Lockyer, J. G. Mead, K. S.
Norris,
W.
B. Reilly, S. H. Ridgway,
D. Robineau, G. J. B. Ross,
T.
A.
Wright,
who either typed, re-typed or proof read
many of the manuscripts.
Contamination
19-29.
L.,
Jr and Praderi, R. 1985. Distribution of Com merson s dolphin, Cephalorhynchus
commersonii and the rediscovery of the type of
Lagenorhvnchus
Inst. Tokyo 36:159-70 .
sur la maturation du squelette post-cranien chez
le dauphin de Commerson [Cephalorhynchus commersonii
(Lacepede, 1804)]
Richardson and
 
rguelen, en parti
culier dans le
  golfe du Morbi
en, pen
.
Robert
Studies   o
Center,
  PO Box 27, Casterton, Victoria 3311, Australia
4 Current Addre
ntic from a
examination of specimens in museums and o ther institutio
ns
in 1972-1984,
boat, s
Tierra
in 1804. It was docum
ented with greatest clar
mes are
Commerson's dolp
Tonina
America between
about 41°30'S and 55°S, at the Strait of La Maire. Their range includes the
eastern Strait of
d Punta Arenas, and
uth
postulated, e
bays, harb
y enter rivers. The
only av ailable abundance
eastern Strait of Magellan, where aerial surveys supported an estimate
of 3 ,211 (SE = 1,168) , but the species does not appear rare anywhere within its broad range.
It prefers
range, and cool temperatures influenced
by
Fuego
nd biology of the species can now be describ
ed
, are described. The first
and 1 4 9 c m (male
) from the Strait of
all
and
alving
takes
in austral
 
COMMERSON S DOLPHINS
length is about 75-80cm. Gestation is
believed to be
brown or
adult
'widow's peak
the flank patch, may be
sufficient to distinguish individuals. The dorsal fin
is rounded
Although
Commerson's
pods,
reported herds have contained
from 1 to over 110 (x = 6.85, SD= 16.5). These dolphins
feed at various
trophic
levels, often in areas disturbed by tidal change. Food items include
coastal schooling fishes and bottom-dwelling invertebrates,
the latter with surprising
including harassment of floating gulls, investigation of human
divers, riding on large swells
and
frequency
suspected high
set for crab and various fishes. Some
sport sniping is known to occur. Legislation
and changes in fishing practices, target species,
and fishing
areas
have
likely reduced the frequency of kills in recent years. About 33 have
been live-captured
and removed to marine zoological parks in Europe and the USA.
INTRODUCTION
different
Commerson's dolphin,
encounter off southeastern South America in
1767 (Laeepede, 1804). It is by far the most widely distributed of the
four. It has
been reported from
southern
Paulian, 1953; Angot, 1954; Pascal, 1901; Robineau, 1984; Robineau and
de
Buffrenil, 1984), and the Falkland Islands (Islas Malvinas), in the
southwest
Atlantic (e.g. Bruce, 1915; Hamilton, 1952; Strange, 1972), and off southern and
southeastern South
the central
Knowledge about this species
or
so
writers primarily
on the validity of the putative species or
its
he
observed
in
'the Bay of the Rio Santa Cruz' and presented details on external
and skeletal
confusion
the
species
and
little
p.
628)
and
from, approximately,
most
contained
little
E
9)
 1988
perform at
published information (e.
g. Fraser, 1
Yanez,
1
948, p. 115; Carcelles, 1948, p. 5; Marelli, 1953, p. 137;
Mann Fischer, 1957,
103; Cab
rera, 19 61, p. 64; Walker, 1968, p. 1120; Morzer Bruyns
, 1971, p. 42;
126).
n a
in southern South America. Mermoz
(1980)
spent
havior
Comodoro Ri
ens and
food, museum specimen
1980;
Ag
  and Venegas, 1978; Sielfeld,
ve provided new
information on distribution,
983, 1984a,
Cephalorhynchus and
pertinen
t
OMMERSON S   DOLPHINS
4, with
erra del Fuego, 1
lantic
(be
76), visits to
on's
do
lphins
was
h
e interviewed  for
summaries of their
operating off southern South
distributed to persons 
f Tierra del Fuego we
re surveyed periodically
d
ion of t
removed, wei
n Us
Lockyer, Smellie, 
and Galeazzi,
lp from M . C
larke on squid bea
ckman are examining p
arasites.
O
n the basis of the amount of fu sion of the vertebral ep
iphyses to the centra
on
cer
vical
fused
on
cer
vical
vertebrae);
Class la,
s in 
a letter
as a polish
erson's dolp
cts as swimm
louines
[the
Falkland
(Lesson
 and Garnot, 1826, p. 181) saw the 'white and bla
ck dolphin of
' at Soledad Bay in the 'Malouines', and considered it to
be
two workers re
sisted recognition of
a new species, citing inadequate evidence. Cuvier (1836, p. 217) wrote: 'we d
not think that, from the little the voyager (Commerson) said, the characteristics
of a new
de
ite animal with black extremities as dubious and
argued
that
scien
new species based solel
p. 128, 261) , wh
o spent several summers
saw w
to  
e
xamine 
Cephalorhynchus   he
apparently knew nothing of this species. Because it was nam
ed only from
Moren
o (1892) reported on his detailed observations of behavior of dolphins
in 'the
six specimens he event
measurements. He named his
ation
he
our
p. 267 ; Dabbene, 1902,
dolphin occurred, Islas de los Estados and Pavon, but it
is
illustrated a  specimen of
t
Stanley
specimen
sity Colle
wrote.
He
  mistaken
arly shows a Commerson's
and not the L. cruciger of Quoy and Gaimard (1824) .
Harm
published
data
urements
specific name, which was based on a good and well-illustra
ted
account of external an d  cranial characters, cannot be accepted. It must be regarded, I think, as a
synonym of Delphinus commersonii , which, although described inadequately
and
without
illustrations,
Table
N a m e
S o u r c e / C o m n e n t s
L e J a c o b i t e
L e M a r s o u i n J a c o b i t e
D a u p h i n  
d e
J a c o b i t e
B r i d l e d d o l p h i n s
P e l f i n b l a n c o
G r o u n d p o r p o i s e
P i e b a l d p o r p o i s e
CoMerson s d o l p h i n
S p r i n g e r
T o n i n a , t o n i n a   o v e r a o
r
d e l p h i n d e C o n m e r s o n
T o n i n a b l a n c a y n e g r a
a n d
d e l f i n b l a n
c o
T u n i n a o v e r a
d e l p h i n d e M a g a l l a n e s
W h i t e b e l t e d
p o r p o
i s e
l d
 
d o l p h i n ( s )
Ccraiierson's orig
inal n
t
( 1 8 1 7
p . 1 6 6 ) ; C u v i e r ( 1 8 3 6 p . 2 1 7 ) .
G r a y 1 8 4 6 ) .
C u n n i n g h a m 1 8 7 1 p . 1 2 8 ) .
L a h i l l e ( 1 9 0 5 p . 6 2 ) q u o t i n g t h e
w o r k
( 1 8 9 2 ) w h o
d i d n o t
a s s i g n a c o m m o n n a m e ;
  M a r e l l i ( 1 9 5 3
p . 1 3 7 ) .
B r u c e 1 9 1 5 ) , c i t i n g n a m e u s e d b y
c r e w
B a l a e n a .
B r u c e 1 9 1 5 ) .
B a n n e r ( 1 9 2 2 p . 6 2 7 ) ; m o s t w i d e l y a c c e p
t e d
a n d
w i d e l y u s e d e g .
B r o w n e l l ,
1 9 7 7 p . 4 8 3 ) .
H a r t ( 1 9 3 5
p . 2 5 3 ) , r e f e r r i n g t o w h a l e r s n a m e s
f o r d o l p hi n s , t h o u g h t
 
a n d L a e e n o r h y n c h u s c r u c i g e r .   o f f
S o u t h G e o r g i a .
C u r r e n t l y i n A r g e n t i n a
a n d S o u t h e r n   C h i l e ; t o n i n a ( f o r d o l p h i n ) ,
o v e r a f o r b l a c k a n d w h i t e , a n d n o t e g g c o l o r e d ,
a s
o m e t i m e s
r e p o r t e d ) .
M a r e l l i ( 1 9 5 3
p . 1 3 7 ) .
 
( M a n n F i s c h e r ,
1 9 5 7 p . 1 0 3 ; A g u a y o
a n d  
1 9 6 7
a y o , 1 9 7 5
p
. 1 , 1 2 4 ) .
A g u a y o ( 1 9 7 5 p . 1 , 1 2 4 ) , f r o m i t s a b u n d a n c
e
i n t h e
S t r a i t o f M a g e l l a n .
.
G i l m o r e ( 1 9 6 9 p . 3 0 7 ) ; R i c e ( 1 9 7 7
p . 9 ) ; W a t s o n ( 1 9 8 4 p . 2 4 8 ) .
G r z i m e k 1 9
7 5 , q u o t e d
i
-
s u g g e s t e d f o r
t h e
g e n u s ( G o o d w i n , 1 9 6 6 ) .
addition,
we are aware of 3 4 9 specimens of Cephalor
hynchus commersonii, 77
from various, mostly
published, sources from December 1 8 7 6 to 1 9 8 4 (Table 4 ,
Fig.
4a)
from
All but eight of the au
thors' specimens were
collected from Tierr
a del Fueg
ere fresh specimens from
e
fir
age program, spectacled porpoises,
20 species of
[Figs
following 19 pages. Text begins again
on p. 29]
as • O
ruce, 1915, discussed in text). For details see T
able 2.
dolphins of f South America, 1
9 4 3
areas •). The d
e made du
ring surveys from
For
details
c
enth
9) 29 06 71
12) — 01 77
13) 01/02 80
P u e r t o V i s s
a r ,
  o f
C o r a o d o r o R i v a d a v i a
2
d a y
s N o f P u e r t o D e s e a d o
C o m o d o r o R i v a
d a v i a
E o f R a w s o n 4 3 ° 1 8 S , 6 4 ° 5 3 W
E o
f R a w s o n 4 3 ° 2 0 S , 6 4 ° 5 5 W
E o f P L o b o s 4 3 ° 4 1 S ,
6 4 ° 5 6 W
2 0
m i P . N o r t e , P e
n i n s u l a
V a l d e s , 4 2 ° O
O S 6 3 ° 3 0 W
S W   o f V i e d m a 4 1 ° 2 6 S
, 6 3 ° O O W
P
. N o r t e , P e n i n
s u l a V a
l d e s
G . S a n J o s e
C o m o d o r o R i v a d a v i a ,
4 5 °
5 2 S , 6 7 ° 2 9 W
C o m o d o r o R i v a d a v i a h a r b o r
C h u b u t & S a n t a
  C r u z
G r o u p & s p e c i m e n h a r p o
o n e d
a n t a C r u z
b y P i e d r a b u e n a ( l o s t l a t e r )
2 s i g h t e d , ' p o r p o i s e s
c o m m o n S h o
r e
h e r e
A c c o m p a n i e d s
h i p a l l
d a y ; s e e n T r a n s p o r
t e
  C h a s i n g f i s
h a t
b a s
e o f p i e r   S h o r e ?
2 a n i m a l s ,
l O l O
1 0 3 5 h r s
H e r o
1 a n i m a l , 1 0 4 0 h r s H e r o
3 s e e n
b y C u n n i n g s 1 0 3 0 h r s H e r o
2 s e e n b y C u m n i n g s , 1 5 2 0 h r s
H e r o
  i n w i n t e r * ( P a y n e
  p c t o
r e
1 s w i m m i n g
n r 3 P . s p i n i p i n n i s S h o r
e
S t u d i e d a l l
J a n u a r y ; m a x h a r b o r S h o r e
c e n s u s 6 1 i n c l . 1 1 c a
l v e s ;
l a r g e
s t g r o u p s e e n = 3 1
C e n s u s o n
2 8 J a n u a r y =
1 8
S h o r e
 
S h o r e ,
b o a t s
S a n t a C r u z , A r g e n t
i n a
S a n t a
 
M t E n t r a n c e
1 6 )
1 7 ) ?
  0 1
' . P a v o n
1 8 ) S p r i n g 2 8 S o f  
C o . R i v a d a v i a
1
9 ) 1 9 0 7 6 9 O f f
C .
4 7 ° 2 5 S
  6 5 ° 3 9 W
2 0
) 1 9
0 7
B l a n
W
2 1 ) 2 1 0 7 6 9 M o u t h o
f R . D e s e a d o
4 7 ° 5 5 S 6 5 ° 4 0 W
2 2 ) 2 1 0 7 6 9 O f f
B . L a u r a , R o c a   B
e l l a c o
4 8 ° 3 5 S 6 6 ° 4 6 W
2 3 )
2 2
0 7
u l i a n
4 9 ° 3 5 S 6 7 ° 3 9 W
2 4 ) 2 3 0 7   6 9 N
o f S a n t a C
r u z
2 5 ) 2 4 0 7 6 9 S
o f R . S a n t a C r u z
5 1 ° 3 5 S 6 8 ° 5 8 W
2 6 ) 2 6 0 7 6 9 M o u t h o f . S a
n t a C r u z
5 1 ° 3 9 S
6 8 ° 2 5 W
2 7 ) 2 6   0 7 6 9 M o u t h o f . S a n t a C r u z
5 1 ° 3 9
2 8 ) 1 3 0 6 7
 
4 9 ° 2 0 S 6 7 ° 4 0 W
2 9 ) 1 5 0 6 7 1 S R o c a S a n
t a C r u z
4 8 ° 5 3 S
  6 6 ° 1 5 W
3 0 ) 1 1 / 1 2 7 8 B . S a n J o r g e ,
b e t w e e n P t o
' H u n d r e d s ' a c c o m p a n y i n g t h e
s h i p R e s a l e s
d
a y & n i g h t ,   L . f l o w e r i
S i
o n e
s p e c i m e n k i l l e d S a n t
a C r u z
b y P i e d r a b u
e n a ( l o s t
  b y c r e w )
L . f l o w e r
i
b y
?
e a r l i e r
o b s e r v a t i o n
o f
M o r e n o ? )
G r o u p j u m p i n
g , 1
h
a r p o o n e d f o r F e u e r l a n d
f o o d
  H e r o
1 a n i m a l , 1 4 1 0 h r s H e r o
2 a n i m a l s , 0 8 3 9 h r s
1 a n i m a l , 1 6 1 5 h r s H e r o
1 2 5 4 h r s H
e r o
l O O S
h r s H e r o
1 a n i m a l , 1 5 0 0 h r s H e r o
2 a n i m a l s ,
1
2 4 1 h r s H e r o
1
a n i m a l ,
1 2 4 5 h r s H e r o
2
1 2 0 0 h r s
H e r o
3
a n i j n a l s , 1 5 4 3 h r s
H e r o
S m a l l g
r o u p s u p t   2 0 , 6
S p e e d - b o a t
D
 
& C o . R i v a d a v i a
3 1 ) 2 7 8
  P u e r t o D e s e a d o ?
c a p t u r e d , t a k e n
o D u i s b u r g   Z o o
4 c a p t u r e d f o r S u n s h i n e A q u a r i u m
c o n f i s c a t e d i n N e w   Y o r k , 1 5
D e c
T a b l e 2 c o n t
i n u e d
S t r a i t
o f
l a n ,
3 2 ) 2 6 7 2 N e a
r T i e r r a d e l F u e g o
S t r a i t o f M a g e l l
a n
3 3 ) 1 6 7 S t r a i t
o f M a g e l l a n
3 4 ) 1 6 0 1 6 7 ' W s i
d e o f P . D u n g e n e s s
3 5 ) P r e - 1 9 2 2 S t r a i t o f M a
g e l l a n
3 6 ) P r e - 1 9 4 6 C h a n n e l s
A t l a n t i c
c o a s
t
3 7 ) 1 0 1 2 6 8 N E o f B . G e n t e G r a n d e
i n S e c o n d N a r r o w s
5 2
3 8 )
1 2 0 8 6 9 P . A r e n a s 5
3 ° 0 7 S
7 0 ° 5 0 W
3 9 ) 1 5 0 5 S t r a i t o f M a g e l l a n
- 0 6 0 7 7 0
4 0 ) 1 1 0 6 I n o p e n o c e a n n e a r
E
- 1 6 0 7 7 1 e n t r a n c e o f i n S t r a i t
o f M a g e l l a n
4 1 ) P r e - 1 9 7 3 N o l o c a l i t y ( p r o b a b l y
S t r a i t o f M a g e l l a n )
4 2 ) 2 1 0 2 7 3 S t r a i t
o f M a g e l l a n
4 3 ) 1 9 7 3 ?
N o l o c a l i t y
4 4 ) 1 0 0 2 7 8 C . V a l e n t i n , I . D a w s o n
5 3
° 3 3 S 7 0 ° 2 5 W
4 4 a ) l l / 1 2 8 3 N e a r P .
S a r a , N E
S t r a i t
o f M a g e l l a n
44b —
S a r a , N E
-   2 8 4 S t r a i t o f M a g e l l a n
T i e r r a d e l
F u e g o , A r g e n t i n a
4 5 ) 1 0 0 1 6 9 2 E N E
  c o a s t , S o f R . G r a n d e
4 6 )
l o s E s t a d o s
4 7 ) 1 1 / 1 2 4 8   C a n a l d e B e a g l e
4 8 ) P r e - 1 9 5 3 C o a s t s o f T i e r r
a d e l
F u e g o
O b s e r v e d b y P . C o m m e r s o n , m s . t o B o u d
e n s e ,   P
B u f f o n ,   w h o s e n t i t t o l a c e p e d e
  E t o i l e
S i g h t i n g s
r e c o r d e d a s N a s s a u Q
D e l p h i n u s
  b i v i t t a t u s
' b r i d l e d d o l p h i n s ( D e l p h i n u s
N a s s a u R
b i v i t t a t u s ) ' s e e n a r o u n d s h i p
o n ' s e v e r a l s u
b s e q u e n t o c c a s i o n s '
P h o t o   o f 2 b y L i o n e l A d a m s .
S
' H u n d r e d s . .
p l a y e d a r o u n d s
h i p '
' F r e q
u e n t l y s e e n ' T
' h o a r d s . . . a l l d a y l o n g ' . 2 H e r o
U , V ,
s p e c i m e n s c a p t u r e d .   P h o t o s i n W ,
  X
B a r r e t t , A g u a y o . S o u n d s r e c o r d e d .
2   a n i m a l s H e r o
E F
' s e e n o n l y i n c e n t r a l S t r a i t o f H e r o
E
M a g e l l a n '
' s e v e r a l g r o u p s ' H e r o Y
P h o t o s b y F . E r i z e
Z ,
a n
i m a l s , 0 8 3 0 h r s X
2 g r o u p s o f a d o z e n e a c h
C a l y p s o B B
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r S h o r e , E E
c r e w , D u i s b u r g Z o o v e s s e l
S h o a l o b s e r v e d b y   J o s e p h B a n k s E n d e a v o u r F F
i n e v e n i
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s i g h t e d
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a n i m a l s e e n ( + / - 1 0 * l o n g . ) H e r o
a i r e P h o t o b y M e a d C a b o t H e r o
S
e e n b y A . F . F e r n a n
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2 s e e n
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g n e a r t h e b e a c h S h o r e
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1 9 7 5 )
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,
3 s e e n
b y A g u a y o M a t u r a n a .
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w a k e . U S C G
i c e
b r e a k e r G l a c i e r . F i l m b y J . W .
I I
J J ,
D
8 9 2 :
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1 5 7
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a n d / o r 1 9 7 1 : 1 0  
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p c
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Falklan d Qfalvinas) Islands
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m b r o k e
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p o r p o i s e s . T
e m p
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m p
4 0 . 7 F B a l a e n a R R
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s i g h t i n g a s a b o v e )
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c a u g h t
i n
n e t , B a l a e n a R R
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n d e e ,
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a s L a g e n o r h y
n c h u s c r u c i g e r )
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  s e e n s e v e r a l
t i m
T e m p .
4 4 . 6 F ( c i t e d a s
L a e e n o r h y n c h u s c r u c i g e
r )
k
n o w n , c o m m o n e r o n W . S h o r e S S
s i d e
' P r o b a b l y
t h e m o s t c o n m o n S h o r e T T
( d o l p h i n ) '
M
a n y s i g h t i n g s o f d o l p h i n s UU
a r o u n d S o u t
h
c a l l e d
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a n d w h i t e p o r p o i s e s N a s s a u R
t
e a r i n g t h r o u g h t h e w a t e r '
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o i s e s ( p i e b a l d ? ) ' B a l a e n a
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c r u c i g e r )
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( O r i g i n a l l y   c i t e d a s
L a g e n o r h y n c h u s
c r u c i g e r )
' S e
v e r a l p i e b a l d p o r p o i s e s S c o t i a R R
s e e n .
T e m p . 4 4 . 6 F ' ( O r i g i n a l l y
c i t e d a s L a g e n o r h y n c h u s c r u c i g e r )
' S c
h o o l
o f p o r p o i s e s ( p r o b a b l y S c o t i a R R
p i e b a l d )
s e e n i n e v e n i n
g . T e m p .
3 9 . 8 F ' . ( O r i g i n a l l y
  c i t e d
a s
L a g e n o r h y n c h u s
  c r u c i g e r )
' B l a c k o n t h e b a c k a n d w h i t e o n
S c o t i a R R
t h e
' P o r p o i s e s
( p r o b a b l y
p i e b a l d ) '
  S c o t i a R R
( O r i g i n a l l y c i t e d a s
L a g e n o r h y n c h u s c r u c i e e r )
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e b a l d a n d b l a c k S c o t i a R R
p o r p o i s e s '
( O r i g i n a l l y c i t e d a s
L a g e n o r h y n c h u s c r u c i g e r )
S o u r c e
I Mermoz
t , 1 9 8 0
:
268
Z
KK Pine et al.
.
C
EE Thielke, 1 9 8 4 
FF
RR Bruce,  1915
S S Hamilton,
South America by
L o c a l i t y
N o . a n d C o m m e n t s
P l a t f o r m O b s e r v
e r
R i o N e g r o a n d C h u b u t ,
A r g e n t
i n a
1 ) 1 7 0 7 7 8 P u n t a Q u i r o g a
, G o l f o d e S a n
M a t i a s
4 3 ° 4 5 S
6 4 ° 4 0 W
2
— 9 7 9
  V i e d m a , 3 0 k m u p r i v e r
3 ) 1 9 8 0 - 8 4 V i e d m a
a n d   u p R i
o N e g r o
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4 4 ° 5 7 S
6 5 ° 0 1 W
5 ) 1 2 1 1 8 3
C o m o d o r o R i v a d a v i a
6 ) — 1
4 P e n i n s u l a
  V a l d e s , m o u t h o f  
R i o   S a n J o s e
C r u z , A r
g e n t i n a
7 ) 2 0 1 1
7 2 S o f C a b o B l a n c o
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1 1 7 2
2 m i l e s   E I s l a s P i n g u i n o s  
4 7 ° 5 5 S 6 5 ° 4 0
W
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o G a l l e g a s
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o D e s e a d o
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4
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a n t a C r u z
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b o D a n o s o
1 9 m i l e s
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4 8 ° 4 8 S 6 6 ° 5 1
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l e t a  
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° 5 2 W
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o , B .
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a P t o S a n t a
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g e n e s
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f t e n
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t i d a p c )
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2
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s e e n   w i t h i n 1 -
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p l a y e d a r o u
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d 1
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z o d i a c s
c r o s s i n g S t
r a i t o f M a g e l l a n
4
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d e P u e r t o D e s e a d o
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n e s
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R i o S a n t a
C r u z
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r t o S a n J u l i a n
P u e r t o S a n J u l i a n
B a l n e a r i o B c o . J u s t i c i a ,
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t o S a n J u l i a n
R i o G a l l e g
o s 5 1 ° 3 5 S 6 9 ° 0 4 W
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a n
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G a l l e g o s 5 1 ° 3 4 S 6 9 ° 0 4 W
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P u
e r t o D e s e a d o
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L o y o l a , P
u e r t o S a n J u l i a n
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Q u i l l a ,
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P u e r t o D e s e a d o
N o . a n d C o m m e n t s
3 :
a c c o m p a n i e d k a y a k
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a t
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c c o m p a n i e d
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u n u s u a l j u m p s
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l a u n c h
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P o r v e n i r a n d r e t u r n
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n a s
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d o f B a h i a I n u t i l
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a h i a I n u t i l
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2 2 W
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t l a n t i c e n d
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s e e n m o s t
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t r i p s t w i c e a
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C . e u t r o p i a ?
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n  
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4
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o w s
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s t i a n
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a n
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a n i e d
s
  h o
, 3
s e
r a t e ,
h
2 :
s
o
n
1 0 - 1
t e s
u r i n g   2
0 m
i n .
4 : f
, 2
  : g o i n
s e c
  s e
  1 0
  2 , 2
r o u p s
o f 2
, 3 - 4
 
, r i g a n d
r o p
s a l
l   o v
b a
h
r y n
S h o r
F u e g
u i n o
F u e g
u i n o
o
S
t e r
O i l
R
Y a
5 1
c h ,
  P a r
a m o
O f f
R i o
C u l
l e n
 
g u
O
5
ISSUE 9) 1988
3
N o .
B . S a n
  S e b a s t i a n
C a l e t a L a M i s i o n
5 3 ° 4 1 S 6 7 ° 4 9 W
M o u t h o f R i o G r a n d e
N o .
o f
2 A d u l t w i t h
S e v e r a l : w e n t
2 - 3
y o u n g
a f t e r
  a
Y a k t e
m i
Y a
O b s e r v e r
6 5
6 5
6 0
1 0 5 ) 1 4 0 2 8 2 P u n t a S i n a i , E s t a n c i a S a r a
1 0 6 ) 1 3 0 3 8 2 R i o C u l
l e n S .
1 4
0 3 8 2 P u n t a d e
A r e n a s ,
P e n i n s u l a P a r a m o
1 0 8 )
2 7
0 3 8 2 R i o G r a n d e ,
n e a r m o u t h
1 0 9 ) 1 0 1 0 8 2 B . S a n S e b a s t i a n
5 3 ° 1 6 S 6 8 ° 1 6 W
1 1 0 ) 2 3 - B . S a n S e b a s
t i a n
2 6 1 1 8 2 5 3 ° 1 7 S  
6 8 ° 1 4 W
1 1 1 )
1
8 1 2 8 2 P u e s t o 1 7 , E s t a n c i a S a r a
1 1 2 ) 3
8 2 C a b o P e n a s
1 1 3 ) 1 1 0 1 8 4 P u n t a  
M a r i a
1 1 4 ) 1 3 0 1
8 4 R a n c h o T a d i c h
1 1 5 )
C h o r r i l l o s
B . S a n S e b a s t i a n
1 1 6 ) 1 4
 
B . S a n S e b a s t i a n
1 1 7 ) 1 4 0 1 8 4 L o s C h o r r i l l o s ,
B .
S a n S e b a s t i a n
1 1 8 )
0 1
0 8 4 R i o G r a n d e ,
E s t a c i o n A C A
1 1 9 ) 1 4 0 3 8 4
E o f P e n i n s u l a P a r a m o
1 2 0 )
1 4 0 3 8 4 3 k m S o f R i b C u l l e n
1 2 1 ) 3 0 0 3 8 4 M o u t h o f R i o L
u z
1 2 2 ) 0 7 0 4 8 4   R a d a R i o
  G r a n d e
1 2
3 ) 0 8 0 4 8 4 R i o G r a n d e
1 2 4 ) 2 8 0 4 8 4 O f f s h o r e o f R i o C u l l e n
1 2 5 ) 1 0 0 5 8 4 B . S a n S e b a s t i a n
D r a k e Passage
1 2 6 ) 2 7 7 W e l l S o f C a p e H o r n
1 2 7
C a p e
H o r n
Falklan d M a l v i n a s )
I s l a n d s
1 2 8 ) 0 7 1 2 7 7 C a r c a s s I s l a n d ,
5 1 ° 3 0 S 4 7 ° 4 0 W
1 2 9 )
0 9 1 2 7 7 C a
r c a s s I s l a n d ,
1
3 0 )
2 0
0 1 7 8 C a r c a s s I s l a n d ,
1 3 1 ) 3 0 1 2 7 8 C a r c
a s s I s l a n d
1 3 2 ) 2 0 1 2
7 9
1 3 3 ) 2 8 1 2 7 9
P o r
t S t a n l e y H a r b
o r
0 3 0 1 8 0 P o r t
S t a n l e y
1 3 5 ) 2 8 0 1 8 0 M a l v i n a s 5 1 ° 3 0 S 5 7 ° 4 0 W
1 3 6 ) ? 0 1 8 0 M a l v
i n a s , 2 0 m i l e s E .
5 2 ° 5 3 S 5 7 ° 5 0 W
1 3 7 )
1 2 1 2 8 0 P o r t S t a n l e y
s c h o o l
o f
1 : n e a r
s h o r e . R e s p i r a t i o n
r a t e r e c o r d e d
1 2 - 1 5 : g r o u p s o f 3 - 4 , s u r f i n g
i n w a
v e s
7 - 1 2 : g r o u p s o f t w o , i n r i v e r
c h a n n e l s a t l o w
t i d e
3 - 4 : 2 , 2
p s o f 2 - 3   w i t h
L a g e n o r h
y n c h u s a u s t r a l i s
2 - 4 : s u r f i n g
i n
0
  o f 1 , 2 , 3  
1 : s e e
2
n e a r
1
n e a r
s h o r e
4 - 6 : s w i m m i n g i n b r e a k e r s
1
s h o r e
1 :
i n t h e b r e a k e r s
1 : a p p r o a c h e d b o a t , v e r y
c u r i o u s
2 :
a p p r o a c h e d b o a t
6 :
c r o s s e d b o w t o w a r d s h o r e
3 : p l a y
i n g a r o u n d s h i p
2 : s w i m m i n g N ,
f a s t , a h e a d
a l o n g s i d e b o w
3
3 - 5 :
s e v e r a l g r o u p
s f o l l o w e d
z o d i a c s
3 0 - 4 0 : e s c o r t i n g z o d i a c s ,
6 h o u r s
5 :
e s c o r t e d b o a t s
1 2 : e s c o r t e d z o d i a c s , 2
h o u r s .
P h o t
1
3 :
  2 a n i m a l s ; l a t e r
1 f o l l o w e d
l a u n c h
1 0 0 : s c h o o l
s o f
5 - 2 5
a
a p e r i o d o f 4 d a y s
4 :
h i p
4 F o l l o w e d z o d i a c s
S h o r e
S h o r e
S h o r e
S h o r e
Y a k t e m i
Y a k t e m i
S h o r e
S h o r
M o t o r b o a t
M o t o r b o a t
H e r o
L i n d b l a d
E x p l o r e r
W o r l d
D i
6 6
W o r l d
D i s c o v e r e r
W o r l d
D i s c o v e r e r
L i n d b l a d
E x p l o
r e r
B r a n s f i e l d
7 9
8 0
8 1
8 2
7 7
1 4
1 4
B r a n s f i e l d 1 4
J o h n B i s c o e 8 3
J o h n B i s c o e 8 4
J o h n B i s c o e 8 5 ,
South tlantic O cean
1 1
0 1 9 N o f B u r d w o o d B a n k
5 3 ° 3 5 S 5 7 ° 2 0 W
3
S e e n f o r 1 m i n
8 6
Sp
erica
by
Total
blished.
Other
notations
 with mandibles.
,
N
onico,
Puerto
ty, Portland,
Oregon; SM
-F, Senckenberg
es
arious sour c
activ ities
  7
g o
h
e   S
m o n t
  M a f
a s   N
o l e t
,   t i
P a V
a m o ,
t i a n
s t l a
a s t i
a s t l a
  n K m   5 -
a n  
t l a
n K m
  K
a s t l
a s t l a
  n
t i a j
n  
; s k
n e s
- 9
i a j i
K m 8 - 9
n a s
Paramo K m 55
Paramo K m 55
PaVamo K m 55
Punta Marfa
Punta 
st
B.
Near
San
San Sebastian
.5
Paramo Km 56
Paramo 56.5
San
1
  S
o T
.5
S
C
abo
Penas
S
S
Cabo
Pen
as
S
Punta  Mari a 2 K m N
Punta Marfa
Rfo S a n Pablo S E
Rfo
Rfo Grande M
Paso de las Cholgas
S
A
A
A
ds of Comme
available
si
4 (Figs 2
can
be
supplemented by
been
coast
abo
of Magellan, waters
occurrence is not
d at
ata) (Brow nell and Praderi, 1985) cannot
be used
with any
old specimen
drift
many
miles
No Com merson's dolphins
coast
d
el
d
el
The no
s
h
401). However, w e
o
LPHINS
in an area atypical for this species, and the entry in the
published table reads
. W e have
the
Vied
ma (41°26'S, 63°00'W) in June 1 9 7 1 and
in Gol
, Table 2),
Commers
and
Within the Strait o
w son, and
on the eastern
rth of Porvenir. [A
 from
Commerson's dolphins
Sec
Leatherw
ood,
Kast
here are 
Desolation by J.T. Polking
kland (Malvinas)
ilton, 1952; and
The souther
solated
enco
, in February 1 9 6 6
(Aguayo and Torres, 1 9
67; A
guayo, 1
9 7 2 and 1 9 7 8
(Table 3
tip of
here
waters
merican
m
none
ommerson's do
lp hins
at South
rs (at u
riding on the pressur
econd-hand report h
bservations at the
  island. For example,
h
Georgia,
23
Novem
rch 1 9 1 3 , has never
been eported to
d Comm
a Iniitil northwa
ce (Leatherwoo
e there aresevera
at
th
ere
may
 water
littoral
spec
ies
waters of
 
ounts. W e regar
d th ree recent
reports of this sp
the species
 found
Comme
earshore. More
 the harbor at Co
ment
 
 
ISSUE 9) 1988
Off mainland Argentina (Mermoz, 1980), around the Falklands (Harmer, 1922;
Hamilton, 1952) and off Kerguelen (Angot, 1954)
Commerson's
along the edges of kelp beds (Macrocystis and
Durvilled). Lahille (1899) reported Commerson's dolphins at Isla Pavon in the
Rio
Santa Cruz. Records detailed in Table 3 confirm the occurrence of these
dolphins well upstream in the Rios Grande (Tierra del Fuego), Gallegos,
Deseado, San
latter case, animals apparently have
been observed making forays as far as 30km from the sea, beyond the town of
Viedma. All the
(1954)
described
an
incident at Kerguelen on 16 February, 1952 in which a
group
of
3 0 animals
beached themselves but returned to sea on the next wave; such
behavior has
of dolphins
a receding tide in a narrow tidal estuary southwest of
Rio
Gallegos,
meandering Rio San Martin, indicate that the animals do not
always
successfully
In
general,
off
South
prefer
areas
where
the
temperatures are influenced by the cool
Malvinas
current.
allopatric
Chilean dolphin, C. eutropia, by contrast, seems to prefer the western
side
Cameron, 1988;
Commerson's dolphins are 4-9°C in summer near Cape Horn,
annually average
11.5°C near Peninsula Valdes (Gorden and Goldberg, 1970) and have been
measured to be as high as 13-16°C in November in Bahia San Jorge
(Gewalt,
1979).
(1892), Bruce (1915) and
Harmer (1922) and detailed
comments on body form by Harmer (1922). Inevitably, some authors who
copied
presented descriptions and illustrations.
and Bonner (1980) do not show the characteristic rounded fin
or
flippers,
other
illustrations
bear
little
resemblance to the living animals. With the significant numbers of observations
and
Size
Prior
were
a 136.3cm male (USNM 550156 Spotte et al., 1979) and
a 147.3cm female
112
South
MMN (SPECIAL ISSUE 9 ) 1 9 8 8 3 5
American
Strait of M ag
an those from elsewhere. In fact, the
largest specimens we are aware
 of from South America
a 152cm female (Fig . 5, upper
right),
bo
th
c
 Duisburg Zoo
herwood
Rivadavia (Mermoz,
nd
N
ishiwaki,

from
6.5cm, fem
Tota
l., 1 9 8 1 , 1 9 8
8): Class 1 males averaged
108.7cm,
males averag
are
us Cephalorhync
hus   Comm
 to be consistently
 del Fuego. The maximum weights
 
2kg 
(males)
specime
ns
from
reported on
3.26-3.74
 (Corn
ell 
e
ly
th
ree
th
ick.
Head
 is rounded
described
the tip of the snout. There is no ell d
efined beak, but
the upper. The blow
1.2cm in length (n 
M M
and
t
o
r t  
k e n o t c h
  T
L )  
r i o r   m a r g
i n o f
o l e
a n g
i o n o
p e r  
a n t .
i n s
n o f   d
  f i n
  p
  f i n
c u s
c e n
g e n
i t
r o
u t h
e a r
F l
a
o r
  l e n g
  w i d t h
D o
s e  
h e
 
h
 
)
y
s s ,
l a c e
D i a m
e t e r
o f h e
n e y e
g t h  
 
o l e   w i d
t h
g e
  s
g e n i
t ,   m a l
merson's dolphins
from Tier
g e
M e a n S t a n d a r d d e v i a t i o
n C
o e f f i c i e n t o f v
a r i a t i o n
Fema
M a
l e s R N P s p e c i m e n s )
C l a s s
2
1 4
C l a s s 3
1 9
64.3-73.
on's dolphin
 Fuego
e m e n t
R a n g e
M e a n
S t a n d a r d d e
v i a t i o n
C V
B
l u b b e r s k i n
M u s c
o n e
)
L u n g , r i g h t
L i v e r
S p l e e n
F a l s e p a n c r e a s
P a n c
r e a s
K i d n e y , l e f t
K i d n e y , r i g h t
I n t e s t i n e s
M a m m a r y g l a n d , l e f
t
n d ,   r i
L e f t t e s t e s
R i g h t t e
s t e s
9 , 2 5 0 - 2 4 , 4 6 2
7 , 7 8 0 - 1 3 , 5 9 0
2 , 8 5 0 - 9 , 0 6 0
1 5 0 -
3 0 0
2 9 0 -
9 5 5
7 5 0 - 1 , 5 9 0
5 - 4 5
6 0 - 3
4 5 - 5 0 0
4 0 - 4
1 0 , 5 1 7
. 4
2 1 9 .
1 7 . 7
1 9 8 .
4 , 7 8 7 . 0
2 , 0
4 0 . 3
r lobate,
appear directl
y correlated
gree of
 p. 87) ;
wh
lehole, 1963 , p. 214); and
 black and white (
  M
plashed w ith black on the head, back an
d tail' and
ery
sm
all
female
. Although
 
 gray spots
e and descri
ented
a
detailed
called 'artwo
0 ; M arc
uzzi and Pilleri,
Adults
with a
ego; in only one o
f the
was
t
(Robineau, 1984) was seldom seen in
specimens from Tierra
by a
with a
highly feathered
by
as a progressively
the
black, as
a patch around
the genital region (see detailed discussion below). The rest of the body is white
(Fig. 5 left).
sharply demarcated ventrally, slightly less so laterally. As
noted
by
Moreno
(1892)
and
white zones
may be slightly streaked or feathered but without a transitional gray zone.
The
black
regions
or lighter than the dark areas of C.
eutropia,
as alleged
by Watson (1981, pp. 249-50). There is no visible eye patch
of the kind that can be distinguished in species such as Peale's dolphin,
Lagenorhynchus australis and C. eutropia. There is
no visible flipper stripe,
black
might
be
axillary
patch
in
a lobe of black.
along
the border between black and white on the back of
the head, varies
 
c f
10 cm
  to scale.
dimorphic, mal
tricted marking. Goodall an
nce of such
l
ISSUE 9) 19
 
his pa
A sm
all proportion
 sm all circula
variable genita
l patch
 the
anterior
h
alf
d
posteri
en illus
than
la
ltz
(1973
nd no animals with
ind
illus
trated
by
circumstances surroundin
me measuremen
 
n females than in males (Leatherwood
and W alker,
 o
is short
in males
tches
females
and
Norris
wledge  of sexually dimorphi
, it is being used to distinguish between the sexes i
n
obs
example, J.T. Polkinghorn was able to
determine that a
lled on its
of his
irregula
r
interior
edge
was dark.
Some tongues were whitish with scattered black spots. The upper
portions of the
The penis is
of
the
dark
the
death,
near-term male fetus from a female taken
incidentally in fishing nets. The
fetus' color pattern consisted of black zones and
a white throat patch
smaller than
on adults.
the
center
third
stripe (Fig. 5, right).
described,
neonates of C. commersonii
darker gray and black.
that 'the
grayish and then the
being at
Gewalt (1981) reported young
Punta Sara, Chile in
frequently
signs of shading. A
m otions and
close association with an adult) we observed at the First Narrows
on 3 1 January 1984 displayed essentially
all elements of adult coloration but as
gray on black or
San
Cornell, 1985).
Commerson's
stranded animals begin to
darken soon after exposure
color
and
data from studies of
strandings and photographs must
are subjective
entire
of markings.
World began to light
distinguish from adults except by
his
sm
although he
the
 appeared to be
 
e can
than
whi
te
on
g
ere
d
etermined
t
juveniles. The largest sp
l
a
han 117cm long and one year old or less. Locky
er
 lon
female s
n that
age (Fig.
5, middle).
ia
(Goodall
, Norris
ing on the
all white Commerson's
Leatherwood, 1 9 8 2 ) and no
 oth
 
 
on
ro Rivadavia. W e have found
 
d by
detected on close examination by a
darkening in the
c
apture.
Bit
e scars were present on five of 1 1 specimens of C . co
mmersoni i fr
of skin of a
Commerson's dolphin which he described as a soft outer layer of
1.5mm
and 355353) and found no evidence
of
cones.
of deep-sea fishes
aximize the detection
that the
brain of
a 50
was
SOOccm . From these estimates, the authors asserted that the species has a
relatively high brain weight
Lagenorhynchus and
highest degree
ii from Tierra
left kidney, 256-396
ventral
(16-30mm, x = 21), mid-lateral (14-25mm, x = 20) and mid dorsal
anterior to the dorsal fin (18-35mm,
x = 25). Intestines of 1 3 animals were
7.0
to
length.
Parasites
L
C
hile
e have found Comm
rnal
parasites
n part, reflect the
nets; so, they were not
beach-cast animals which may have stranded because they were ill. W e have
found worms
e
chial tubes, in on
lungs, stomach wall
rld f
ISSUE 9) 1988
Duisburg, Germany in 1978 died
of 'mycosis of the
1979; pers.
treatment en route to Japan died
of 'verminous
fourth animal suffered from
by
transport,
unable to use her flukes fo
r
al., 1979) but measured only 117cm at
 death (measured by
d bottlenose
 
and 
of
Plata
C.
eutropi
a
these last two spec
ther
ent
in
of 280-30
(Goodall,
unpublished).
Measurements
(classes 2 and 3)

23, range = 249-
285, x = 268.7, S = 9.3, CV = 0.03); Class 2 (n = 40, range = 26
9-300, x =
3
3
en measured
only 127.6cm
pparently  th
e smallest
sexually matur
sexually matur
moment,
hysicall
n t
M e a
 
C V
a s a l   l e n g t
h
. 2  
t h  
2 5
  1 3
0 1 3 7 .
  w i d t
l e n g t h
2 5
 
a x i l l a e
w i d t
 
a e w i d t h
  a t m i d
l e n g t h 2
5 2
 
3 / 4 l
2 5
4  
u m t i p t o
  e
n a r e s
2 5
8 4 . 0
9
R
o s t r u m t i p t o i n t e r
n a l n a
r e s
. 2 1
1 0 P r e o
 
  1 2 2 .
o r b i t a l
w i d t h
2 3
1 4
0 . 0
  w i d t h
 
n a r e s
w i d t h  
2 6
. 9
o m a t i c
w i d t h
2 5
  1 4 1 .
s t w i d
o f
s 2
e t a l
2 4
 
e h e i g
h t
2 5
9 9 .
1
  l e n g t h
2 5
. 6 1 0
a l f o s s a
l e n g t h
L
  f o s s a l e
n g t h R
2 4
  5 8
l
f
2
5
6 4 0 .
l f o s s a
w i d t h R
2 4
h   L
2 5
. 0 4 3 . 4
l e n g t h
  R
b i t a l p
r o c e s s l
e n g t h L
2 5
i t a l p r o
c e s s l e n
g t h R
n a r e s w i d t h
2 5
h L  
1 9  
2 0
2 5
1 2 1 2
o w l e n g t h
2 5
 
2 3
3 3
L R
  t o o t h r o w l e n g t h
2 3
1 1 6 . 3
a u s l
2
3
 
2 3
2 1 8 . 6
t L
2 1
b u l a r   f o s s a
l e n g t h
  L 2 2
l a r f o s s a l
e n g t h R
2
  L  
0
n g t h R
2
5
 
. 6
g t h R
 
ted in h
hynchus
in
such
t
s 3 animals
nes
rom Kergue
anial skelet
s reported by
ertebral
first tw o 
always and four
e anticl
inal vertebra
, 1988) . T
che
Usuall
  tuburcle.
The
imals do not
= 41
  (L = 
3
2.67,
 right
s
point
posteriorly
nd 3 fe
(or 6); IV:
-3) . This differs slightly
pecies fuse
1988) .
ales by 5-6 years a
nd
about
he  samplewas lactatin
 the sma
5, p. 1 33
 
sex u
had
 its ov
aries and
were immature
ars 
an
have been found
. A fu
, (mother RNP 
634) w
as long as its mother) and w eighed
 7.27kg(12 .4
as in the ri
rooves were presen
t on eac
e
m
each side of the f
ace, just below and anterior
 to
  the  tipof
and
March
or
February
in a Northern emisp
ribed an
in animal
83 a
mals he captured
 a
rea
f
eed
ottom (IWC,
)
rtant
prey,
mmerson's
do
these conclusio
ns are
hallows, in
 and anchovies
turbed area, or ot
. 6 ; Mer
 has
b
een 
f
n open
o extrica
h, 1 9 8 3 was unable to
return to
eral on th
hen the
eral on th
hen the
bab ly t h e squid,
L o l i g o s a n p a
u l e n s i s s e e B a s t i d a
e t a l . ( 1 9 8 8 ) .
 
 
t e d   s p
e c i e s
C o m
e
y n c h u s
  e u t r o
p i a
F e b .  
i n i p i
n n i s ( 3
)
n i p i n n
i s   (
i n s ,
s h a
?   ( 3 )
F
L
h y n c h
u s a u s
t r a l i s
 
J a n
l i s ( 1 )
a l i s
  f u l m
a r s ( 2
l i s
c e n s
m m e r s o
n s *
t a n t ,
n o i n
*
i n t
o n s  
e r a c t i
o n s
s o n s
  d
s , S t r a
i t
n
  a b o a
t e m i *
i n g
e   a
j u s
e b r e
a t  
u l a
o
f
k m s
l y s e
  t o g e t h
e r  
t e d
y t o g
,
w a k e
t
d
 
, 1
  N o r r i s
 
J .
r n
w o o d
L e a t
d
D o l p
d g e
  o n
i n g
ere were nume
r a
Gallina, pe
merson's
dolphins
gulls, La
rus dominicanus
e resting near the rig after su
rfeiting with food
ing
breached on it, landing
how (1930,
p. 137)
and othe
rs have
d aerial acr
es 
sport them selves
they do app
the bow
20, and modern
an and inland
They can maneuv
h areas
 species
i
lphins sometim
he observed played around the ship at great
speed (Lacepede, 1804) . Moreno (1892) described how they escorted the Resales
'day and nig
themselves to
of his vessel; Cabrer
them aro
und anchored
Scott and Dotte Larsen (Table 3, Sightings 52
and
76, respectively) saw them at the bow w ave of the Navarius and the MS
Lindblad Explor
er. Dolphins
from the
they
landed
passe
). One
2) is the
ough they so
observed
 
sonar was operating at
pers.
comm.). Commerson's dolphins appear to have responded to sounds of motors,
whistling, or shouting on
Pascal (1981) found that
was
stopped,
a
of Commerson's
produced
red
ith
inelife
Aquarium
100kHz, probably
ertoire includes
 stereotyped, pulsed
referred
and
animals
that
ins
housed
 with
at Duisburg Zoo. The former authors
 
cies with
The
sou
s
be
hose of
 most other
uding Commers
on's dolp
Chilean  channe
ne animal w
n and lived lar
gely on the m
More
80;
Sielfeld
1978; Venegas and
Sielfeld, 1979; and others). Fishermen feel that dolphin meat does not
quickly
deteriorate
thought
to
dy
t,
olphins, in
There are
no reliable estimates on numbers of dolphins killed for bait for crab
tr
fishing takes place mainly
e
onii
estimated
f Magella Chile. Such captures
difficult to control.
harpooning dolphins and w e found
harpoons, some with blood indica
ting
ab boats visited. The centolla in this region has been
overfished
and
to cento
rab fishing takes place in the eastern Strait of Magellan,
where these dolphins are abundant
(L
indiscrim
d for sport'
accounts
1988b).
Accidental/incide
ntal 
takes
hins
in
1 ) Incidental entanglement in
from boats
hins,
dolphin (Goodall, 1978) . Since 1976, onl
y
traps
dolphins cannot enter
(Estancia An
Mitchell,
19
and that
is
scal
e
at
t Isla
 
ECIAL ISSUE 9)
that
such
mbers, but fishermen in Bahi'a Imitil caught an estimated 30
dolphins
and C
species
occurr
ing
in
and Peale's
  reports
o
Commerson's d
and occasionally river
mouths with extreme
and Cameron,
perpendicular t
o the
in areas where
ly sloping. The nets
(Fig.
13
merged
these nets. Any
ishermen find them are left on
the flats to swim away on the next tide. A minimum of 69 Commer
son's
n drowned in robalo nets in Argentine Tierra
del
aters, especially in Bahi'a Imitil and along the eastern S
trait of
(Goodall and
Cameron, 1980) .
that mortality continues;
  In some areas,
 other fish takes plac
deployed from small boats (usually rowboats). This small-scale fishery employ
s
ing
circling a
circle; and two boats stretching a net between them. These
methods are seldom used in Tierra
del Fuego
but
of fish takes place
mmerson's and other
of
overboard, some ha
ve gone to
a fish meal plant and a few have been donated
to scien
tific institutions.
(Table 5). Data
composition
e fishing season
The specim ens show
imals were taken durin
ppears tha
t dolphins
ccording to
 and the
 full m
more dolphins
e first than in
the last half; for
ere w ere twice
Bla
veh i
g this species, three
ed
).
AL ISSUE 9) 1988
on
5
into
two hours
ent
rmit allowing th
e
nd the dolp
have died during capture
operations (Gewalt, 1979; pers. comm., 1984). Four animals, one male and
th
transferred to  Mystic Marinelife
Aquarium, where they lived a few hours, eight days (the male), and tw o and one
half years, respecti
at Co
were granted a
permit to captu
ublic
sentim
ent
ornell
.
uego.
ish. Res.
es mari
nt de m
a
el ano 19
78. Puertos de 
ile y U ru
. Ed it
Baker, A .
331^.
Barren,
f dolp
anuscript) .
l.
auna mari
aceos en agua
D
ata
on
incheira
ommer
. K
 
  B ioaco
8 . Letter fro
18 67
iferos
y
18
27,
9 ,
8 3 3 .
Tomo 4 , parte
onnees su
r la
. Co
ntributi
on
a
 
Ete rovi
c, F.
E
 bottom
with
s
Association, Mystic,
family Delphin
Zool. I: 171-82.
merson's dolph
oo Duisburg, Chronik Ge
55pp.
Duisberg.
Z
oologische
 
so uth
central Chile, early winter 1970. Antarct. J. US 6(1): 10-11.
Goodall,
del Fuego
5 05 .
ns stranded on Tierra del Fuego. Sci.
Rep.
Wha
les
chus
 
2
5 .
Goodall,
R
hynchus
commer
 Dec.
1979:
Goodall, R.N.P. and Cameron, I .S. 1980. Exploitation of small
  cetaceans off southern South
Gooda
985. Review of
the  food habits
Condy and  R.M
pos ium on Antarctic
Whal. C om m n 37: 399-40.
Goodall. R.N.P., Galeazzi,
Sobral, A.P. 1984. Studies on Commerson's
dolph
Cameron, I.S. 19
aleazzi, A.R., Oporto, J.A. and Cameron, I.S. 1988. On the
Chilean dolph
na
obtusata
, with
a
comparison
pedia. 8 : 324-7.
Map Folio Series, Folio 13, Americ
an
Geographical
Gra
nd Gray, J.E. (eds.)
ebus and T
ek's
d, S . 1982. Two sightings of white pilot whales, Globicep
hala
mela
ena,
a
eans.
(2): 338-43.
the Falkland
4(66):

Co
3 8 + 3 plates.
Hart, T.J. 1935. On the diatom s of the skin film of whales, and their p
ossible bearing on proble
out of wate
mmal. 52: 2 2 2 .
I
L,
SC/28/Rep.3
nt. W hal. Commn 28: 7 9-82 .
Interna
e
on
Kamming
rsma, H. 19
phalorhynchus
comm
deration of re
5: 1-103.
1 0 3 -7.
n geografica de
L
lenas de nuestros m
tle-nosed dolphins (Tursiops
rn Califor
nia, Baja
t w hale dolp
ceans. Plenum Publ. C
orp., New York. 438pp.
Leatherw ood, S., Kastelein
Paper
S
Leather
Strait of
Magellan, January
-February 1
98 4.
rvations of C
ary 1984.
Les
aux observes d
onde
ciences
 
s Anne'es 1
s
l
re
., 1 9 5 :
 
Marc
oography of
Pilleri  (ed.).
n, C ephalorhyn
Whales Res. Inst., Tokyo (3 2 ): 149-53 .
Mitchell, E.
smaller cetaceans.
. 1973.
actuales de la Repiiblica
a.
 Whales and
.
Niella, R .
r.
Rev
ipeds. University of Tokyo
3 -
2 0 4 . In: S.H. Ridgway (ed.) Mammals of the Sea,
Biology an
d Medic
cetace
Hero Cruise
nos. Acta . Zool.
-32 .
Pa
Comm erson
aux lie
Mission Kerguelen
1951. Mem.
Inst. Sci.
UCN Newslette
s
nce
on
 the
tion of
C ephalorhynch
.
m the sea, m ainlan
d and i
5-14.
Plii
l Ulls
rvettes de S
Physicienne
Zool: iv
 Canadian
Rice, D.W. 1977. A list of m arine mammals
 of
Report
N
Zool. 62 :
 
 en partic
orbihan.
 
551-9 . 
S
hochi,
d.). Inve
ions d
e delfi
nidos en
a conserva
tion y
m an
rs on's d
est d
p. 
Thielke,
9 .
tlantico Sur. E d
am mals
teristic
al
61
1977. Underw
ater sounds
of Ceph
lzjager im
F eu
erland. lagadabent
ener e
ines Ube
of the
in. N
atural Histo
rson s 
aerial and vessel surv
 
Magellan) o
Marta eastward to the entra
nce to the Strait. They ap
peared to be most abundant in and near
the
n
slightly
lowe
t at
re seen as individ
t
ground swells,
n,
Comm
cidentally in nets and are harpooned by
fishermen for us
cies
Commerson's
dolphins, Chilean and
  agreeme
nts
with
sts from the Chilean
biology and
ommittee
IN SOUTHERN CHILE
ith place names mention
ed in the text.
azzi, Leatherwood, Miller,
in
and
D M
Between 20 January and 1 9 Febru
ary 1984 an expedition was made to southern
Chile (Fig. 1 . Data on
Commerson's dolphins were collected by  the
 following
means (see Table 1 and Fig. 2): (1) Aerial surveys, at 90
knots and 500ft, in a
P
non-amphibious
aircraft
with
about
(a)
en the First and
show n in
all
areas
safe
ly
accessible
(relative cove
rage of such reconnaissance surveys is shown in Fig. 2B). (2)
Flights aboard tw in-engine Dauphin and Bolkow BO-105-C helicopters,
operating at 60 to 90 knots at 300 to 1,000ft
altitude between Punta Catalina
ls in the eastern
8
to
in ca
as
 
S c h e d u l e o
f a c t i v i t i e s d
u r i n g e x p e d i t
i o n t o S o u t h
e r n C
  L o c a
t i o n
2 0 J a n 1 9 8
4 S h o r e v e h i c
l e )
S e n o O t w a y n
o r t h e a s t s i d e )
2 1 J a
n 1 9 8
t *   E s t
r e c h o d e M a g
a l l a n e s
S t r a i t o f M a g
e l l a n ) ,
  b
u n t a A r e n a s
a n d P r i r o e r a
A n g o s t u
r a   F i r s t
N a r r o w s )
2 2 J a n 1 9 8 4 S h o r e v e h i c l
e ) R o a
o m P u n t a
A r e n a s   t o F u e r t a B u l l n e z
2 3 J a n 1 9 8 4
  F e r r y
F i r s t   N a r r o w s
b e t w e e
n P u n t a D e l
g a d a a n d T i e r r a d e l
F u e g o )
2 4 J a n
1 9 8 4 A i r c r a f t *
F i r s t N a
t o
a A r e n a s
2 5 J a n 1 9 8 4 A i r c r a f t * S t r
a i t o f   M a g e l l a n b e t w e e
n P u n t a A r e n a s a n d F i
r s t   N a r r o
w s )
4 B o a t S e
g u n d a A n g o s t u
r a
N
a r r o w s ) f r o m b a s e a t S a
n
2 7 J a n 1 9 8 4
S h o r e
v e h i c l e ) P u n t a A r e n
a s t o C a b
o P o s e s
i o n
o t ) P o s e s i o n
H e l i c o p t e
r * * C a b o P o s e s i o n t o
P u n t a C a t a l i n a
2 8 J a n  
1 9 8 4 H e l i c
o p t e r * * S t r a i t o f M a g e
l l a n e a s t o f
  F i r s t N a
r r o w s )
2 9   J a n 1 9 8 4
H e l i c o p t e r * * * S t r a i
t
a s t o f F i r s t
  N a r r o w s )
3
4 A i r c r a f t
* S t r a
i t o f M a g e l l a n b e t w
e e n P u n t a A r e n a s a n d F
i r s t N
a r r o w s )
3 1 J
a n 1
9 8 4
F i r s t N
 
  D e
l g o d a a n d T i e r r a d e l F u e g
o )
3 1 J a n
1 9 8 4 S h o r e ( f o o
t ) P u n t a D e l g
a d a
1
  F
e b 1 9 8 4 A i r c r a f t *
  S t a i t
o f M a g e l l a n b e t w e e
n P u n t a A r e n a s a n d
)
2 F e b 1 9 8 4 F e r r y
B e t w e e n P
u n t a A r e n a s a n d P o r v e n
i r
  W e s t o
f T i e r r a d e l F u e g o B
a h i a I n u t i l t o S e c o
n d
3 F
e b
  P u
n t a A r e n a s t o S a n G
r e g o r i o S e c o n d N a r r o w s )
A i r
c r a f t *
S t r a i t o f M a g e l l
a n P u n t a A r e n a s
  C a b o F r o w a r d )
a n d S e n o O t w a y
4 F e b 1 9 8 4
B o a t  
e l l a n e a s t
o f F i r s t
N a r r o w s )
5 F e
S
a n G r e g o r i o t o I s l
a M a r t a
9
- 1 5 F e b 1 9 8 4 B o a t
S t r a i t o f M a g e l l a n
  w e s t o f P u
n t a   A r e n
a s )
  o
77
g
5
LU
HERD SIZE
rn i
Fig. 4. Distribution of C. commersonii herd sizes including all observations from aircraft, helicopter,
small
boat
p. 205) collected specimens from Bahia Inutil and reported that
fishermen there
We
failed to see any
Commerson's dolphins on our one ferry crossing from Punta Arenas to
Porvenir, 2 February,
reconnaissance
Isla
and
off Cabo San Vicente, at the southwest end of the Second
Narrows
(Fig.
3B).
east to the
entrance of the Strait (Fig. 2). Within those areas, Commerson's dolphins were
seen
concentrated in and near
coverage (Fig.
found in larger
The
population in that stretch (Fig. 3A, 3B) was estimated to contain 3,161 animals,
SD = 900
narrows we found
Commerson's dolphins in
all segments of the entrance bay to which we were
able to
and
companies servicing the oil rigs
maintained
sightings logs for March-June in the eastern Strait. They reported
seeing
Commerson's dolphins on 10 of 43 flights in March, 8
of
in
in June.
shore
ct the
facts th
se
areas
mmerson s 
erved them sur
of shore, ridi
nts as
vast m ajority,
, some
 interviews,
 tha
the m any
resented with
olphins i
s i
stern Stra
the ac
tivity apparent
lly unchanged
sin ce
  belonging
CIAL ISSUE 9) 1988
for crab in southern
meat of dolphins to that of seals, fish or
te r
restrial m
bait, in order
than
that the
  even if dolphin
indicating recent use,
Further,
harpooning dolphins, mostly for
shifting more each season
from centolla to centollon
western Strait we surmise the pressure on remaining Commerson's dolphin
populations in Chile will lessen. However, given that the centollon fishery
operates essentially ye
ar-round and in
areas difficult to
con
cern
erson's
dol
er
species
summarized below,
hus sp., encou
or
because d
b), individuals not positively
ined 1
st (11 of 16) within or
coastw
ard
found in the company
of
Commerson's dolphins but came to a moving vessel to ride its bow
w
olphins. Two new borns,
estimated to be less
d, one at the
February
and
ta on 5 February.
Animals identified by fishermen as Peale's dolphins are killed by rifle or
hand-
ear the
f
Isla
Santa
Inez,
the.
fishermen reportedly took 'hundreds' in 1980 and 1981 and the
reafter
ISSUE 9) 1988
t of M ag ellan.
D a t e
2 2 J u l 1 9 8 4
2 6
1 O c t 1 9 8 4
2 O c t 1 9 8 4
4 O c t 1 9 8 4
1 8 O c t
1
2 N o v 1 9 8 4
4 N o v 1 9 8 4
L o c a t i o n
O f f I
N e a r
P u e r t o N a t a l e s
S e n o U l t i m a E s p e r a n z a
C
a b o S a n Y s i d r o
5 3 ° 3 2 ' S   7 0 5 5
 ' W
5 2 ° 5 2 ' S , 7 1 ° 0 0 ' W
5 3 ° 5 5 ' S , 7 1 °
1 0 ' W
5 3 ° 2 5 ' S , 7 1 ° 1 0 ' W
5 3
5 3 ° 4 7   S , 7 3 ° 2 0 ' W
5 3 ° 5 2 ' S , 7 3 ° 2 2 ' W
5 3 ° 5 7 ' S , 7 7 ° 2 2 ' W
5 4 ° 0 7 ' S , 7 3 ' 0 5 ' W
5 4 ° 0 9 ' S , 7 2 ° 4 9 ' W
5 4 ' 0 7 ' S , 7 2 ° 1 0 ' W
5 3 ° 5 5 ' S , 7 0 - 5 5 ' W
N u m b e r
3
W . S i e l f e l d
W . S i e l f e l d 2
R . M a t u s
S . L e a t h e r w o o d
E d u a r d o C a r
l o s  
E
d u a r d o C a r l o s  
 
L e a l
E d u a r d o T / » a V
 
L e a l
  L e a l
_
I d e n t i f i e d b y
S .
L e a t h e r w o o d
f r o m p h o t o s  
a t H M R C
A e r i a l o b s e r v a t
i o n
I d e n t i f i e d b y
S .
L e a t h e r w o o d
_
_
_
_
_
_
_
_
_
-
species w e asse
mbled during and subsequent
to our expedition are
  flippers, flukes
, photos
boats have fishe
ortant
seaso
might fear
lled annually
(Goodall and Jordan, 1986) .
shallows just east
ring a beach walk on 27 Janua
ry and a 4.57m
ip of Punta
examined a
a recent
of 3 1 came
en Rio
terfly in the western end o
f
ute
single male, ca 7m
sla Marta on
group
of three at 52°29'S, 73°42 'W in the northwestern
Strait of Magellan on 1 1
February
The surveys rep
from Sea
  Research Center. W e thank the following
 
Goldsberry of
planning field research;
skillful piloting and
Matus
help with
Duisberg Zoo; Simon
and Maurio Rivas of ENAP, Dave Knapton, Eduardo Arevalo and
Carlos Gonzales of Servicio Aereo Littoral and Patricio
Matamaura
of
Helimares
for
eastern Strait; Captain
Francisco Gallardo for time aboard the ferry Patagonia; Gary Glaxner of
Gulf
Fleet
Jorge Jordan for time aboard the
crab tender M/V
REFERENCES
Aguayo L,
A. 1975. Progress report on small cetacean research in Chile. J.
Fish.
Antarctic
Mammals,
American Geographical Society, New York.
Cornell, L.H., Antrim, J.E., Asper, E.D. and Pincheira, B.J. 1988. Commerson's
dolphins
Thompson, P.O. and Jehl, J. 1971. Bioacoustics of marine mammals
off Argentina: R/V Hero cruise 71-3. Antarct. J. U.S. 6(6): 266-8.
Cunningham, R.O. 1871. Notes on the natural history of the Strait of Magellan
and west coast of
Edmonston and Douglas, Edinburgh. 517pp.
Gilmore, R.M. 1971. Observations on marine mammals
and
birds
off
central Chile, early winter 1970. Antarct. J. U.S.
6(1): 10-11.
Goodall, R.N.P.
Sci. Rep. Whales Res. Inst.,
Tokyo 30: 197-230.
Exploitation of small cetaceans off southern South
America. Rep. int. Whal. Commn
30: 445-50.
Goodall, R.N.P. and Jordan, J.D. 1986. Review of fisheries in the Magellan region of Chile.
Document SC/38/SM21 presented to the IWC Scientific Committee, May 1986. (Unpublished).
9pp.
1979. Preliminary report on
of small cetaceans of f
southern South America and the Antarctic Peninsula. Paper SC/31/SM2 presented to the IW C
Scientific Committee,
June 1979.
Goodall, R.N.P., Galeazzi, A.R., Leatherwood, S., Miller, K.W., Cameron, I.S., Kastelein, R.A.
and Sobral, A.P.
Tierra del Fuego, 1976-1984, with a
review of available information
South
Harmer,
S.F. 1922. On Commerson's dolphin and other species of Cephalorhvnchus. Proc. Zoo/.
Soc. London 1922:
627-38 + 3 plates.
January-February
1984.
Document SC/36/SM8 presented
46pp.
dolphin. Oceans 18(6): 46-9.
 
83
in this volume).
Mermoz, J.F. 1980. A brief report on the behavior of Commerson's dolphin, Cephalorhynchus
commersonii, in Patagonian shores. Sci. Rep. Whales Res. Inst., Tokyo
(32): 149-53.
los Mamiferos de Chile. (Manuscript). 18pp.
Sielfeld
K.,
Chile.
Atalah,
rnanejo de mami'feros marines
marines
Sielfeld W, K., Venegas, C.C. and Atalah, A.G. 1977b.
Consideraciones acerca del
sounds
of
 
1988 85
Dolphin
  Cephalorhynchus
S.G. Brown
ABSTRACT
Fifteen
sighting
records
South
American coastal waters and off the Falkland Islands made from vessels of the 'Discovery
Investigations' in the years
and
their behaviour. External measurements and other information from a captured specimen
are recorded. Statements on the occurrence of the species
around South Georgia are
INTRODU TION
Hemisphere
during
marking
voyages by the vessels Discovery (1925-27), RRS Discovery II (1929-39 ) and
RRS
also
some
whale marking cruises using chartered whale catchers in the waters around
South Georgia
of all
cetaceans sighted. The records of the large whale species have been
used
in
several
publications
but
most
paper now in preparation on the
sightings of small cetaceans. For this note it is only necessary to state that
where
possible detailed notes were made of the external appearance, colour, estimated
size, numbers and behaviour of the
animals. These notes were
animals. These notes were
and
Commerson's
enable a definite identification
. AMERICA S . GEORGIA
confirmed are reported here. A few additional sightings lacking sufficient
details bu
t whi
ch either
may have
been omitted.
D I S T R I B U T I O N
Fifteen records of Commerson's dolphin
seen from RRS William Scoresby an
d
RRSDiscovery //between April 1931 and March 1 9 3 4 are listed in Table
1 .
One
(observati
American coastal waters. Seven sighting
s
(No
s
and two (Nos
seen just off the 
Five sightings were in Argentinian inshore waters to
the
Jan
off Puerto Gallegos north of
the Strait, and two (Nos 8 and 9) on 2
February
del Fu ego, to the south. All 1 5 records
were in
ns give
als seen but
it is clear in two cases (Nos 2 and 5) and possibly in a third (No. 7 when at
least
o
the observation, the mean school
size
B E H A V IO UR
Thirteen
the behaviour of the dolphins.
For 12 records notes
time.
the vessel
us
r
a
One of the five or six dolphin
s encountered on 10
5)
was
d (see below). While it was in the water harpooned,
Rayner notes that:
panions failed to show as much interest in the condition
of the wounded
htened from the ship. After getting under way
, they still accompanied the
jumped clear
 
o n  
1
2
3
4
5
6
7
D a t e / t i m e
2 3 A p r i l 1 9 3 1
( 1 0 . 4 0 h r s )
1 8 S e p t e m b
e r 1 9 3 1
( O Q . O O h r s )
1 8 S e p t e m b e r 1
9 3 1
s )
3 D e c e m b e r 1 9 3 1
( 1 3 . 5 0 h r s )
1 0 J a n u a r y
 
r s )
1 1 J a n u a r y 1 9 3 2
( 1 5 . 3 0 h r s )
1 2
3 2
3 5 h r s )
P o
5 2 ° 4
( S t
a i t o f M a g e l l a n )
5 2 °
( A p p r o
a c h i n g S e c o n d
N a
r r o w s , S t r a i t
o f
M a g e l l a n )
S t r a i t o f M a g e l l a n
5 1 ° 2 6 ' S 5 9 ° 0 8 ' W
( O f
f F a l k l a n d I s . )
5 1
( O f f P u e r t o
G a l l e g o s
A r g e n t i n a )
5 1 ° 3 9 ' S 6 8 ° 5 5 ' W
( P u e r t o G a l l e g
o s )
5 1 ° 2 7 ' S
6 8 ° 5 1 ' W
( O f f P u e r t o G a l l e g o s )
N o . o f
1
2
2 0
a t
B e h a v i o u r
B e h a v i o u r
B e
B e h a v i o u r
S p e c i m e n ,
B e h a v i o u r
B e h a v i o u r
B e h a v i o u r
V e s s e l
W S
-
(09.55hrs)
(11.20hrs)
(16.00-16.15hr
s)
(19.20hrs)
15 2 0 M a r c h
1 9 3 4
(Off San Sebastian
S t r a i t o f M a
g e l l a n
( A p p r o a c h i n g f i r s t
N a
t o D u n g
e n e s s )
S t r a i t o f M a g e