Atlas of the Bryophytes of WisconsinJeff Rose
Department of BotanyUniversity of Wisconsin-Madison
07 November, 2016
Contents
Diversity of Bryophytes by County 2
History of Bryology in Wisconsin 3
Atlas Methodology 3
Mosses 4Floristics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4About the Maps & Nomenclature . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Hornworts and Liverworts 52Floristics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 52About the Maps & Nomenclature . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 52
1
Diversity of Bryophytes by County
number of species
under 99 − 4747 − 7373 − 100100 − 146146 − 190190 − 218218 − 254over 254
125
253
126
313
3929
58
41
39
31
92
53218
32
216
283
8
76
63
89
90
190
61
65
141
156
113
99
130
31
26
33
73
32178
91
198
78
27
53
172
43
84
181
70
136
047
135
190
47
65
209
113
42
254
133
88
87
980
77
159
215
100
53
95
146
60
38 7
84
2
History of Bryology in Wisconsin
The earilest collections of Wisconsin bryophytes date back to those of Increase Lapham from the 1830’s. Otherimportant collectors or students of Wisconsin Bryophytes include C. Barnes (WIS), F. Bowers (UWSP), L.Cheney (WIS), G. Conklin (WIS), J. Christy (MPM), R. Evans (WIS), S. Freckmann (UWSP), H. Iltis(WIS), R. Koch (UWL), and M. Nee (WIS). As seen in chart below, the peak activity in brophyte collectingwas between 1960-1980 with over 10,000 collections in that time frame.
year
Col
lect
ions
010
0020
0030
0040
0050
00
1800 1840 1880 1920 1960 2000
Atlas Methodology
Records were downloaded from the Consortium of North American Bryophyte Herbaria websiteand supplemented by the database at the Milwaukee Public Museum (MPM). Currently, 35127 Wisconsincollections are databased. Names were then examined to check for synonomy, remove unidentified specimens,and remove specimens lacking locality data. When possible, dubious specimens at WIS have been checked.However, specimens from other herbria have not been checked unless they are present in duplicate at WIS.In cases where the identifications are highly dubious, these collecteions have not been maped and the speciesexcluded from the flora for the time being.
3
Mosses
Floristics
Mosses (Bryophyta) can be found in all terrestrial and aquatic habitats throughout Wisconsin. Habitatsparticulalry well-suited for large amounts of moss diversity include cliff faces, wetlands (bogs, fens) andparticularly white cedar (Thuja occidentalis) swamps. The distribution and diversity of mosses is far fromuniform throughout the states (see map). This is due to a combiantion of non-uniform distributuion ofhabitats and collection bias towards more “scenic” areas. The problem of collection bias is further hamperedby a lack of bryologists acitvley collecting in Wisconsin in the last several decades. Areas of the highestmoss diversity in the state are (1) the Driftless Area, particularly the Baraboo Hills, (2) the Wisconsin RiverDrainage, (3) the Lake Superior shore, and (4) the Door Peninsula. Conversley, areas of low diversity are(1) West-Central Wisconsin and (2) the Lake Winnebago Area. High divertsity areas (1), (2), and (4) arealso well-known for their high diversity of vascular plant species.
The Baraboo Hills contains species that likely persisted in the Driftless Area during Pleistocene glaciation,but also contains some northern species disjunct from the northern part of the state. Driftless area specialitiesinclude Bryoxiphium norvegicum, Dicranodontium denudatum and Jaffueliobryum spp. Another floristicallyimportant feature of the Driftless Area is several Ozarkian species that enter the state in Grant Countyincluding Forsstroemia trichomitria and Zygodon viridissimus. Associated with the diversity of the Driftlessarea is a noticible increase in diversity along the Wisconsin River (and apparent slight decrease in theAdams/Jeauneau County areas of glacial Lake Wisconsin). This seems to be sometimes associated withspecies that are more widespread in the northern portion of the state but follow the length of the river to theMississippi River. Examples of this pattern include Homalia trichomanoides, Isopterygiopsis muelleriana,Mnium lycopodioides, and Saelania glaucescens.
Door County is signifigant in its mixture of northern and southern habitats, but also supports several GreatLakes dune species sich as Niphotrichum cansecens.
Areas of low diversity are misleading in that the exceptionally low numbers are more than likely due toundercollection (for example, Pepin County likely has at least one moss species), although habitat loss dueto agriculture and/or relative habitat uniformity is likely to be an additional factor. These areas shouldspecifically be targeted for additional records, especially the west-central part of the state, where species ofa slightly more western distribution may yet be found.
About the Maps & Nomenclature
The moss flora of Wisconsin presently contains 415 species in 45 families. Nomenclature follows Flora ofNorth America volumes 27 and 28. The following taxa have been excluded from the flora due to probablemisidentifications (specimens are not at WIS), nomenclatural issues, or problems with georeferencing:
Amphidium mougeotii, Aulacomnium turgidum, Brachelyma subulatum, Brachythecium mildeanum, Bryumsandbergii, Bryum turbinatum, Campylophyllum halleri, Claopodium whippleanum, Dichodontium olympicum,Dicranoweisia cirrata, Dicranum crassinerve, Dicranum elongatum, Dicranum majus, Dicranum ouiscon-sinense, Dicranum spurium, Dicranum strictum, Dicranum tauricum, Grimmia donniana, Hennediellaheimii, Homalothecium aureum, Hygrohypnum smithii, Hypnum circinale, Hypnum ravaudii, Leucobryumantillarum, Mnium hornum, Oncophorus rauei, Orthotrichum affine, Orthotrichum cupulatum, Philonotisgracillima, Philonotis longiseta, Plagiothecium micans, Pogonatum brachyphyllum, Rhytidiadelphus loreus,Scleropodium obtusifolium, Scopelophila ligulata, Sphagnum torreyanum, Sphagnum wilsonii var. quinquefar-ium, Taxiphyllum geophilum, Tetraplodon mnioides, Thamnobryum alleghaniense, Thuidium tamariscinum,Tortula muralis
4
Abietinella abietina(Hedw.) M. Fleisch.
Thuidiaceae
Acaulon muticum(Schreb.) Mull. Hal.
Pottiaceae
Amblyodon dealbatus(Sw.) Bruch & Schimp.
Meesiaceae
Amblystegium serpens(Hedw.) Schimp.Amblystegiaceae
Amphidium lapponicum(Hedw.) Schimp.Orthotrichaceae
Anacamptodon splachnoides(Froel.) Brid.Fabroniaceae
Andreaea rupestrisHedw.
Andreaeaceae
Anomobryum concinnatum(Schrad.) Schimp.
Bryaceae
Anomobryum julaceum(Schrad.) Schimp.
Bryaceae
5
Anomodon attenuatus(Hedw.) Hub.Thuidiaceae
Anomodon minor(Hedw.) Lindb.Thuidiaceae
Anomodon rostratus(Hedw.) Schimp.
Thuidiaceae
Anomodon rugelii(Mull. Hal.) Keissl.
Thuidiaceae
Anomodon tristis(Ces.) Sull. & Lesq.
Thuidiaceae
Anomodon viticulosus(Hedw.) Hook. & Taylor
Thuidiaceae
Aphanorrhegma serratum(Hook. & Wilson) Sull.
Funariaceae
Archidium alternifolium(Dicks.) Mitt.Archidiaceae
Atrichum altecristatum(Ren. & Card.) Smyth & Smyth
Polytrichaceae
6
Atrichum angustatum(Brid.) Bruch & Schimp.
Polytrichaceae
Atrichum crispum(James) Sull.
Polytrichaceae
Atrichum oerstedianum(Mull. Hal.) Mitt.Polytrichaceae
Atrichum tenellum(Rahl.) Bruch & Schimp.
Polytrichaceae
Atrichum undulatum(Hedw.) P. Beauv.
Polytrichaceae
Aulacomnium androgynum(Hedw.) Schwagr.Aulacomniaceae
Aulacomnium heterostichum(Hedw.) Bruch & Schimp.
Aulacomniaceae
Aulacomnium palustre(Hedw.) Schwagr.Aulacomniaceae
Barbula convolutaHedw.
Pottiaceae
7
Barbula unguiculataHedw.
Pottiaceae
Bartramia pomiformisHedw.
Bartramiaceae
Blindia acuta(Hedw.) Bruch & Schimp.
Seligeriaceae
Brachytheciastrum velutinum(Hedw.) Ignatov & Huttunen
Brachytheciaceae
Brachythecium acuminatum(Hedw.) Aust.
Brachytheciaceae
Brachythecium acutum(Mitt.) Sull.
Brachytheciaceae
Brachythecium campestre(Mull. Hal.) Schimp.Brachytheciaceae
Brachythecium erythrorrhizonSchimp.
Brachytheciaceae
Brachythecium laetum(Brid.) Schimp.
Brachytheciaceae
8
Brachythecium rivulareSchimp.
Brachytheciaceae
Brachythecium rotaeanumDe Not.
Brachytheciaceae
Brachythecium rutabulum(Hedw.) Schimp.Brachytheciaceae
Brachythecium turgidum(Hartm.) Kindb.
Brachytheciaceae
Brothera leana(Sull.) Mull. Hal.
Dicranaceae
Brotherella recurvans(Michx.) M. Fleisch.Sematophyllaceae
Bruchia flexuosa(Schwagr.) Mull. Hal.
Bruchiaceae
Bryhnia graminicolor(Brid.) Grout
Brachytheciaceae
Bryhnia novae−angliae(Sull. & Lesq.) Grout
Brachytheciaceae
9
Bryoerythrophyllum recurvirostre(Hedw.) Chen
Pottiaceae
Bryoxiphium norvegicum(Brid.) Mitt.
Bryoxiphiaceae
Bryum argenteumHedw.
Bryaceae
Bucklandiella heterosticha(Hedw.) Bednarek−Ochyra & Ochyra
Grimmiaceae
Buxbaumia aphyllaHedw.
Buxbaumiaceae
Callicladium haldanianum(Grev.) H.A. Crum
Hypnaceae
Calliergon cordifolium(Hedw.) Kindb.
Amblystegiaceae
Calliergon giganteum(Schimp.) Kindb.Amblystegiaceae
Calliergon richardsonii(Mitt.) Kindb.
Amblystegiaceae
10
Calliergonella cuspidata(Hedw.) Loeske
Hypnaceae
Campyliadelphus chrysophyllus(Brid.) R.S. ChopraAmblystegiaceae
Campylium stellatum(Hedw.) C.E.O. Jensen
Amblystegiaceae
Campylophyllum hispidulum(Brid.) Hedenas
Hypnaceae
Ceratodon purpureus(Hedw.) Brid.Ditrichaceae
Cirriphyllum piliferum(Hedw.) Grout
Brachytheciaceae
Clasmatodon parvulus(Hampe) Sull.Fabroniaceae
Climacium americanumBrid.
Climaciaceae
Climacium dendroides(Hedw.) F. Weber & D. Mohr
Climaciaceae
11
Codriophorus acicularis(Hedw.) P. Beauv.
Grimmiaceae
Conardia compacta(Mull. Hal.) H. Rob.Amblystegiaceae
Coscinodon calyptratus(Drumm.) C.E.O. Jensen
Grimmiaceae
Cratoneuron filicinum(Hedw.) Spruce
Amblystegiaceae
Ctenidium molluscum(Hedw.) Mitt.Hypnaceae
Cynodontium polycarpum(Hedw.) Schimp.
Dicranaceae
Cynodontium strumiferum(Hedw.) Lindb.Dicranaceae
Cynodontium tenellum(Schimp.) Limpr.
Dicranaceae
Cyrtohypnum minutulum(Hedw.) W.R. Buck & H.A. Crum
Thuidiaceae
12
Cyrtohypnum pygmaeum(Schimp.) W.R. Buck & H.A. Crum
Thuidiaceae
Dichelyma capillaceum(With.) Myrin
Fontinalaceae
Dichelyma falcatum(Hedw.) MyrinFontinalaceae
Dichelyma pallescensBruch & Schimp.
Fontinalaceae
Dichodontium pellucidum(Hedw.) Schimp.
Dicranaceae
Dicranella cerviculata(Hedw.) Schimp.
Dicranaceae
Dicranella heteromalla(Hedw.) Schimp.
Dicranaceae
Dicranella palustris(Dicks.) A.C. Crundwell
Dicranaceae
Dicranella rufescens(With.) Schimp.
Dicranaceae
13
Dicranella schreberiana(Hedw.) Hilf.Dicranaceae
Dicranella staphylinaH. WhitehouseDicranaceae
Dicranella subulata(Hedw.) Schimp.
Dicranaceae
Dicranella varia(Hedw.) Schimp.
Dicranaceae
Dicranodontium denudatum(Brid.) E. Britton
Dicranaceae
Dicranoweisia crispula(Hedw.) MildeDicranaceae
Dicranum bonjeaniiDe Not.
Dicranaceae
Dicranum condensatumHedw.
Dicranaceae
Dicranum flagellareHedw.
Dicranaceae
14
Dicranum fulvumHook.
Dicranaceae
Dicranum fuscescensTurner
Dicranaceae
Dicranum montanumHedw.
Dicranaceae
Dicranum muehlenbeckiiBruch & Schimp.
Dicranaceae
Dicranum ontarienseW.L. PetersonDicranaceae
Dicranum polysetumSw.
Dicranaceae
Dicranum scopariumHedw.
Dicranaceae
Dicranum undulatumSchrad.
Dicranaceae
Dicranum viride(Sull. & Lesq.) Lindb.
Dicranaceae
15
Didymodon fallax(Hedw.) R.H. Zander
Pottiaceae
Didymodon rigidulusHedw.
Pottiaceae
Didymodon tophaceus(Brid.) LisaPottiaceae
Didymodon vinealis(Brid.) R.H. Zander
Pottiaceae
Diphyscium foliosum(Hedw.) D. MohrBuxbaumiaceae
Distichium capillaceum(Hedw.) Bruch & Schimp.
Ditrichaceae
Distichium inclinatum(Hedw.) Bruch & Schimp.
Ditrichaceae
Ditrichum flexicaule(Schwagr.) Hampe
Ditrichaceae
Ditrichum heteromallum(Hedw.) E. Britton
Ditrichaceae
16
Ditrichum lineare(Sw.) Lindb.Ditrichaceae
Ditrichum pallidum(Hedw.) Hampe
Ditrichaceae
Ditrichum pusillum(Hedw.) Hampe
Ditrichaceae
Drepanocladus aduncus(Hedw.) Warnst.Amblystegiaceae
Drepanocladus longifolius(Mitt.) Broth.
Amblystegiaceae
Drepanocladus polygamus(Schimp.) HedenasAmblystegiaceae
Drepanocladus sordidus(Mull. Hal.) Hedenas
Amblystegiaceae
Drummondia prorepens(Hedw.) E. BrittonOrthotrichaceae
Elodium blandowii(F. Weber & D. Mohr) Eckel
Thuidiaceae
17
Elodium paludosum(Aust.) Broth.Thuidiaceae
Encalypta ciliataHedw.
Encalyptaceae
Encalypta proceraBruch
Encalyptaceae
Encalypta rhaptocarpaSchwagr.
Encalyptaceae
Entodon brevisetus(Hook. & Wilson) Lindb.
Entodontaceae
Entodon challengeri(Paris) CardotEntodontaceae
Entodon cladorrhizans(Hedw.) Mull. Hal.
Entodontaceae
Entodon seductrix(Hedw.) Mull. Hal.
Entodontaceae
Ephemerum cohaerens(Hedw.) HampeEphemeraceae
18
Ephemerum crassinervium(Schwagr.) Hampe
Ephemeraceae
Ephemerum spinulosumBruch & Schimp.Ephemeraceae
Eucladium verticillatum(Hedw.) Bruch & Schimp.
Pottiaceae
Eurhynchiastrum pulchellum(Hedw.) Ignatov & Huttunen
Brachytheciaceae
Eurhynchium praelongum(Hedw.) Schimp.Brachytheciaceae
Eurhynchium striatum(Schreb.) Schimp.Brachytheciaceae
Fabronia ciliaris(Brid.) Brid.
Fabroniaceae
Fissidens adianthoidesHedw.
Fissidentaceae
Fissidens bryoidesHedw.
Fissidentaceae
19
Fissidens bushii(Cardot & Ther.) Cardot & Ther.
Fissidentaceae
Fissidens dubiusP. Beauv.
Fissidentaceae
Fissidens elegansBrid.
Fissidentaceae
Fissidens exilisHedw.
Fissidentaceae
Fissidens fontanus(Bach. Pyl.) Steud.
Fissidentaceae
Fissidens grandifronsBrid.
Fissidentaceae
Fissidens minutulusSull.
Fissidentaceae
Fissidens obtusifoliusWilson
Fissidentaceae
Fissidens osmundioidesHedw.
Fissidentaceae
20
Fissidens subbasilarisHedw.
Fissidentaceae
Fissidens taxifoliusHedw.
Fissidentaceae
Fontinalis antipyreticaHedw.
Fontinalaceae
Fontinalis dalecarlicaBruch & Schimp.
Fontinalaceae
Fontinalis hypnoidesHartm.
Fontinalaceae
Fontinalis novae−angliaeSull.
Fontinalaceae
Fontinalis sphagnifolia(Mull. Hal.) Wijk & Margad.
Fontinalaceae
Fontinalis sullivantiiLindb.
Fontinalaceae
Forsstroemia trichomitria(Hedw.) Lindb.
Leucodontaceae
21
Funaria americanaLindb.
Funariaceae
Funaria flavicansMichx.
Funariaceae
Funaria hygrometricaHedw.
Funariaceae
Gemmabryum caespiticum(Hedw.) J.R. Spence
Bryaceae
Gemmabryum dichotomum(Hedw.) J.R. Spence & H.P. Ramsay
Bryaceae
Grimmia laevigata(Brid.) Brid.
Grimmiaceae
Grimmia muehlenbeckiiSchimp.
Grimmiaceae
Grimmia olneyiSull.
Grimmiaceae
Grimmia ovalis(Hedw.) Lindb.Grimmiaceae
22
Grimmia piliferaP. Beauv.
Grimmiaceae
Grimmia plagiopodiaHedw.
Grimmiaceae
Grimmia teretinervisLimpr.
Grimmiaceae
Gymnostomum aeruginosumSm.
Pottiaceae
Hamaticaulis vernicosus(Mitt.) Hedenas
Amblystegiaceae
Haplocladium microphyllum(Hedw.) Broth.Thuidiaceae
Haplocladium virginianum(Brid.) Broth.Thuidiaceae
Hedwigia ciliata(Hedw.) P. Beauv.
Hedwigiaceae
Herzogiella striatella(Brid.) Z. Iwats.
Hypnaceae
23
Herzogiella turfacea(Lindb.) Z. Iwats.
Hypnaceae
Homalia trichomanoides(Hedw.) Schimp.
Neckeraceae
Homalotheciella subcapillata(Hedw.) Broth.
Brachytheciaceae
Homomallium adnatum(Hedw.) Broth.
Hypnaceae
Hygroamblystegium varium ssp. noterophilum(Sull. & Lesq.) Vanderp. & Hedenas
Amblystegiaceae
Hygroamblystegium varium ssp. varium(Hedw.) Monk.
Amblystegiaceae
Hygroamblystegium varium var. humile(P. Beauv.) Vanderp. & Hedenas
Amblystegiaceae
Hygrohypnum eugyrium(Schimp.) LoeskeAmblystegiaceae
Hygrohypnum luridum(Hedw.) Jenn.
Amblystegiaceae
24
Hygrohypnum ochraceum(Turner) Loeske
Amblystegiaceae
Hylocomiastrum pyrenaicum(Spruce) M. Fleisch.
Hylocomiaceae
Hylocomium splendens(Hedw.) Schimp.Hylocomiaceae
Hymenostylium recurvirostre(Hedw.) Dix.Pottiaceae
Hyophila involuta(Hook.) A. Jaeger
Pottiaceae
Hypnum cupressiformeHedw.
Hypnaceae
Hypnum curvifoliumHedw.
Hypnaceae
Hypnum faurieiCardot
Hypnaceae
Hypnum imponensHedw.
Hypnaceae
25
Hypnum lindbergii(Mitt.) Hedenas
Hypnaceae
Hypnum pallescens(Hedw.) P. Beauv.
Hypnaceae
Hypnum pratenseKoch
Hypnaceae
Imbribryum miniatum(Lesq.) J.R. Spence
Bryaceae
Isopterygiopsis muelleriana(Schimp.) Z. Iwats.Plagiotheciaceae
Isopterygiopsis pulchella(Hedw.) Z. Iwats.Plagiotheciaceae
Jaffueliobryum raui(Aust.) Ther.Grimmiaceae
Jaffueliobryum wrightii(Sull.) Ther.
Grimmiaceae
Leptobryum pyriforme(Hedw.) Wilson
Bryaceae
26
Leptodictyum riparium(Hedw.) Warnst.Amblystegiaceae
Leskea gracilescensHedw.
Leskeaceae
Leskea obscuraHedw.
Leskeaceae
Leskea polycarpaEhrh.
Leskeaceae
Leskeella nervosa(Brid.) LoeskeLeskeaceae
Leucobryum albidum(Brid.) Lindb.Dicranaceae
Leucobryum glaucum(Hedw.) Angstr.
Dicranaceae
Leucodon brachypusBrid.
Leucodontaceae
Leucodon julaceus(Hedw.) Sull.
Leucodontaceae
27
Lindbergia brachyptera(Mitt.) Kindb.Leskeaceae
Meesia triquetra(L.) Angstr.Meesiaceae
Meesia uliginosaHedw.
Meesiaceae
Micromitrium megalosporumAust.
Ephemeraceae
Mnium lycopodioidesSchwagr.Mniaceae
Mnium marginatum(Dicks.) P. Beauv.
Mniaceae
Mnium spinosum(Voit) Schwagr.
Mniaceae
Mnium spinulosumBruch & Schimp.
Mniaceae
Mnium stellareHedw.
Mniaceae
28
Mnium thomsoniiSchimp.
Mniaceae
Myurella julacea(Schwagr.) Schimp.
Theliaceae
Myurella sibirica(Mull. Hal.) Reimers
Theliaceae
Neckera pennataHedw.
Neckeraceae
Niphotrichum canescens(Hedw.) Bednarek−Ochyra & Ochyra
Grimmiaceae
Oncophorus virens(Hedw.) Brid.Dicranaceae
Oncophorus wahlenbergiiBrid.
Dicranaceae
Orthotrichum anomalumHedw.
Orthotrichaceae
Orthotrichum elegansSchwagr.
Orthotrichaceae
29
Orthotrichum obtusifoliumBrid.
Orthotrichaceae
Orthotrichum ohioenseSull. & Lesq.
Orthotrichaceae
Orthotrichum pumilumSw.
Orthotrichaceae
Orthotrichum pusillumMitt.
Orthotrichaceae
Orthotrichum sordidumSull. & Lesq.
Orthotrichaceae
Orthotrichum speciosumNees
Orthotrichaceae
Orthotrichum stellatumBrid.
Orthotrichaceae
Orthotrichum strangulatumP. Beauv.
Orthotrichaceae
Oxyrrhynchium hians(Hedw.) Loeske
Brachytheciaceae
30
Palustriella falcata(Brid.) Hedenas
Amblystegiaceae
Paraleucobryum longifolium(Ehrh.) LoeskeDicranaceae
Philonotis fontana(Hedw.) Brid.
Bartramiaceae
Philonotis marchica(Hedw.) Brid.
Bartramiaceae
Physcomitrium collenchymatumGier
Funariaceae
Physcomitrium hookeriHampe
Funariaceae
Physcomitrium immersumSull.
Funariaceae
Physcomitrium pyriforme(Hedw.) Hampe
Funariaceae
Plagiomnium ciliare(Mull. Hal.) T.J. Kop.
Mniaceae
31
Plagiomnium cuspidatum(Hedw.) T.J. Kop.
Mniaceae
Plagiomnium drummondii(Bruch & Schimp.) T.J. Kop.
Mniaceae
Plagiomnium ellipticum(Brid.) T.J. Kop.
Mniaceae
Plagiomnium medium(Bruch & Schimp.) T.J. Kop.
Mniaceae
Plagiomnium rostratum(Schrad.) T.J. Kop.
Mniaceae
Plagiopus oederianus(Sw.) H.A. Crum & L.E. Anderson
Bartramiaceae
Plagiothecium cavifolium(Brid.) Z. Iwats.
Plagiotheciaceae
Plagiothecium denticulatum(Hedw.) Schimp.Plagiotheciaceae
Plagiothecium laetumSchimp.
Plagiotheciaceae
32
Plagiothecium latebricolaSchimp.
Plagiotheciaceae
Platydictya confervoides(Brid.) H.A. CrumAmblystegiaceae
Platydictya jungermannioides(Brid.) H.A. CrumAmblystegiaceae
Platydictya minutissimum(Sull. & Lesq.) H.A. Crum
Amblystegiaceae
Platydictya subtilis(Hedw.) H.A. CrumAmblystegiaceae
Platygyrium repens(Brid.) Schimp.
Hypnaceae
Pleuridium subulatum(Hedw.) Rabenh.
Ditrichaceae
Pleurozium schreberi(Willd.) Mitt.
Hylocomiaceae
Pogonatum urnigerum(Hedw.) P. Beauv.
Polytrichaceae
33
Pohlia annotina(Hedw.) Lindb.
Bryaceae
Pohlia bulbifera(Warnst.) Warnst.
Bryaceae
Pohlia cruda(Hedw.) Lindb.
Bryaceae
Pohlia elongataHedw.
Bryaceae
Pohlia filum(Schimp.) Mertensson
Bryaceae
Pohlia lescuriana(Sull.) OchiBryaceae
Pohlia melanodon(Brid.) A.J. Shaw
Bryaceae
Pohlia nutans(Hedw.) Lindb.
Bryaceae
Pohlia obtusifolia(Vill.) L.F. Koch
Bryaceae
34
Pohlia proligera(Kindb.) S.O. Lindberg
Bryaceae
Pohlia wahlenbergii(F. Weber & D. Mohr) A.L. Andrews
Bryaceae
Polytrichastrum alpinum(Hedw.) G.L. Sm.
Polytrichaceae
Polytrichastrum formosum(Hedw.) G.L. Sm.
Polytrichaceae
Polytrichastrum longisetum(Sw.) G.L. Sm.Polytrichaceae
Polytrichastrum ohioense(Ren. & Card.) G.L. Sm.
Polytrichaceae
Polytrichastrum pallidisetum(Funck) G.L. Sm.Polytrichaceae
Polytrichum communeHedw.
Polytrichaceae
Polytrichum juniperinumHedw.
Polytrichaceae
35
Polytrichum piliferumHedw.
Polytrichaceae
Polytrichum strictumBrid.
Polytrichaceae
Pseudobryum cinclidioides(Hub.) T.J. Kop.
Mniaceae
Pseudocalliergon trifarium(F. Weber & D. Mohr) Loeske
Amblystegiaceae
Pseudocalliergon turgescens(T. Jensen) LoeskeAmblystegiaceae
Pseudocampylium radicale(P. Beauv.) Vanderp. & Hedenas
Amblystegiaceae
Pseudoleskea patens(Lindb.) Kindb.Leskeaceae
Pseudoleskeella catenulata(Brid.) Kindb.Leskeaceae
Pseudotaxiphyllum distichaceum(Mitt.) Z. Iwats.
Hypnaceae
36
Pseudotaxiphyllum elegans(Brid.) Z. Iwats.
Hypnaceae
Pterigynandrum filiformeHedw.
Pterigynandraceae
Ptilium crista−castrensis(Hedw.) De Not.
Hypnaceae
Ptychostomum cernuum(Hedw.) Hornsch.
Bryaceae
Ptychostomum creberrimum(Taylor) J.R. Spence & H.P. Ramsay
Bryaceae
Ptychostomum cyclophyllum(Schwagr.) J.R. Spence
Bryaceae
Ptychostomum pallens(Sw.) J.R. Spence
Bryaceae
Ptychostomum pallescens(Schleich.) J.R. Spence
Bryaceae
Ptychostomum pendulumHornsch.Bryaceae
37
Ptychostomum pseudotriquetrum(Hedw.) J.R. Spence & H.P. Ramsay
Bryaceae
Ptychostomum weigelii(Spreng.) J.R. Spence
Bryaceae
Pylaisia intricata(Hedw.) Schimp.
Hypnaceae
Pylaisia polyantha(Hedw.) Schimp.
Hypnaceae
Pylaisia selwyniiKindb.
Hypnaceae
Pylaisiadelpha tenuirostris(Bruch & Schimp.) W.R. Buck
Hypnaceae
Rauiella scita(P. Beauv.) Reimers
Thuidiaceae
Rhabdoweisia crispata(Dicks.) Lindb.
Rhabdoweisiaceae
Rhizomnium appalachianumT.J. Kop.
Mniaceae
38
Rhizomnium magnifolium(Horik.) T.J. Kop.
Mniaceae
Rhizomnium pseudopunctatum(Bruch & Schimp.) T.J. Kop.
Mniaceae
Rhizomnium punctatum(Hedw.) T.J. Kop.
Mniaceae
Rhodobryum ontariense(Kindb.) Paris
Bryaceae
Rhynchostegium aquaticumA. Jaeger
Brachytheciaceae
Rhynchostegium serrulatum(Hedw.) A. JaegerBrachytheciaceae
Rhytidiadelphus subpinnatus(Lindb.) T.J. Kop.Hylocomiaceae
Rhytidiadelphus triquetrus(Hedw.) Warnst.Hylocomiaceae
Rhytidium rugosum(Ehrh.) Kindb.
Hylocomiaceae
39
Rosulabryum capillare(Hedw.) J.R. Spence
Bryaceae
Saelania glaucescens(Hedw.) Broth.Ditrichaceae
Sanionia uncinata(Hedw.) Loeske
Amblystegiaceae
Sarmetypnum exannulatum(Schimp.) HedenasAmblystegiaceae
Schistidium agassiziiSull. & Lesq.Grimmiaceae
Schistidium apocarpum(Hedw.) Bruch & Schimp.
Grimmiaceae
Schistidium appalachianumH. H. Blom.
Grimmiaceae
Schistidium atrofuscum(Schimp.) Limpr.
Grimmiaceae
Schistidium confertum(Funck) Bruch & Schimp.
Grimmiaceae
40
Schistidium crassitheciumH.H. Blom
Grimmiaceae
Schistidium dupretii(Ther.) W.A. Weber
Grimmiaceae
Schistidium pulchrumH.H. Blom
Grimmiaceae
Schistidium rivulare(Brid.) Podp.Grimmiaceae
Schistidium virideH.H. Blom & C. Darigo
Grimmiaceae
Schistostega pennata(Hedw.) F. Weber & D. Mohr
Schistostegaceae
Schwetschkeopsis fabronia(Schwagr.) Broth.
Fabroniaceae
Sciuro−hypnum curtum(Lindb.) Ignatov
Brachytheciaceae
Sciuro−hypnum oedipodium(Mitt.) Ignatov & Huttunen
Brachytheciaceae
41
Sciuro−hypnum plumosum(Hedw.) Ignatov & Huttunen
Brachytheciaceae
Sciuro−hypnum populeum(Hedw.) Ignatov & Huttunen
Brachytheciaceae
Sciuro−hypnum reflexum(Starke) Ignatov & Huttunen
Brachytheciaceae
Sciuro−hypnum starkei(Brid.) Ignatov & Huttunen
Brachytheciaceae
Scorpidium cossonii(Schimp.) Hedenas
Scorpidiaceae
Scorpidium revolvens(Sw.) RubersScorpidiaceae
Scorpidium scorpioides(Hedw.) Limpr.Scorpidiaceae
Seligeria campylopodaKindb.
Seligeriaceae
Seligeria donniana(Sm.) Mull. Hal.Seligeriaceae
42
Seligeria pusilla(Hedw.) Bruch & Schimp.
Seligeriaceae
Seligeria recurvata(Hedw.) Bruch & Schimp.
Seligeriaceae
Sematophyllum demissum(Wilson) Mitt.
Sematophyllaceae
Sphagnum andersonianumR.E. Andrus
Sphagnaceae
Sphagnum angustifolium(Warnst.) C.E.O. Jensen
Sphagnaceae
Sphagnum bartlettianumWarnst.
Sphagnaceae
Sphagnum capillifolium(Ehrh.) Hedw.Sphagnaceae
Sphagnum centraleC.E.O. JensenSphagnaceae
Sphagnum compactumLam. & DC
Sphagnaceae
43
Sphagnum contortumSchultz
Sphagnaceae
Sphagnum cuspidatumEhrh.
Sphagnaceae
Sphagnum fallaxH. Klinggr.
Sphagnaceae
Sphagnum fimbriatumWilson
Sphagnaceae
Sphagnum flexuosumDozy & Molk.Sphagnaceae
Sphagnum fuscum(Schimp.) H. Klinggr.
Sphagnaceae
Sphagnum girgensohniiRussow
Sphagnaceae
Sphagnum henryenseWarnst.
Sphagnaceae
Sphagnum imbricatumHornsch.
Sphagnaceae
44
Sphagnum inundatumRussow
Sphagnaceae
Sphagnum lescuriiSull.
Sphagnaceae
Sphagnum magellanicumBrid.
Sphagnaceae
Sphagnum majus(Russow) C.E.O. Jensen
Sphagnaceae
Sphagnum obtusumWarnst.
Sphagnaceae
Sphagnum palustreL.
Sphagnaceae
Sphagnum papillosumLindb.
Sphagnaceae
Sphagnum platyphyllum(Lindb.) Warnst.Sphagnaceae
Sphagnum pulchrum(Lindb.) Warnst.Sphagnaceae
45
Sphagnum recurvumP. Beauv.
Sphagnaceae
Sphagnum ripariumAngstr.
Sphagnaceae
Sphagnum rubellumWilson
Sphagnaceae
Sphagnum russowiiWarnst.
Sphagnaceae
Sphagnum squarrosumCrome
Sphagnaceae
Sphagnum subfulvumSjors
Sphagnaceae
Sphagnum subsecundumNees
Sphagnaceae
Sphagnum subtile(Russow) Warnst.
Sphagnaceae
Sphagnum tenerumSull. & Lesq.
Sphagnaceae
46
Sphagnum teres(Schimp.) Angstr.
Sphagnaceae
Sphagnum warnstorfiiRussow
Sphagnaceae
Sphagnum wulfianumGirg.
Sphagnaceae
Splachnum ampullaceumHedw.
Splachnaceae
Splachnum sphaericumHedw.
Splachnaceae
Straminergon stramineum(Dicks.) HedenasAmblystegiaceae
Syntrichia papillosa(Wilson) Jur.Pottiaceae
Syntrichia ruralis(Hedw.) F. Weber & D. Mohr
Pottiaceae
Taxiphyllum deplanatum(Bruch & Schimp.) M. Fleisch.
Hypnaceae
47
Tetraphis pellucidaHedw.
Tetraphidaceae
Thelia asprella(Schimp.) Sull. & Lesq.
Theliaceae
Thelia hirtella(Hedw.) Sull. & Lesq.
Theliaceae
Thelia lescuriiSull.
Theliaceae
Thuidium delicatulum(Hedw.) Schimp.
Thuidiaceae
Thuidium recognitum(Hedw.) Lindb.Thuidiaceae
Timmia megapolitanaHedw.
Timmiaceae
Tomentypnum nitens(Hedw.) Loeske
Brachytheciaceae
Tortella fragilis(Hook. & Wilson) Limpr.
Pottiaceae
48
Tortella humilis(Hedw.) Jenn.
Pottiaceae
Tortella inclinata(R. Hedw.) Limpr.
Pottiaceae
Tortella tortuosa(Hedw.) Limpr.
Pottiaceae
Tortula acaulon(With.) R.H. Zander
Pottiaceae
Tortula cernua(Hub.) Lindb.Pottiaceae
Tortula mucronifoliaSchwagr.
Pottiaceae
Tortula obtusifolia(Schwagr.) Mathieu
Pottiaceae
Tortula plinthobia(Sull. & Lesq.) Broth.
Pottiaceae
Tortula truncata(Hedw.) Mitt.Pottiaceae
49
Trematodon ambiguus(Hedw.) Hornsch.
Bruchiaceae
Trichostomum spiraleGrout
Pottiaceae
Trichostomum tenuirostre(Hook. & Taylor) Lindb.
Pottiaceae
Ulota crispa(Hedw.) Brid.
Orthotrichaceae
Ulota hutchinsiae(Sm.) Hammar
Orthotrichaceae
Warnstorfia fluitans(Hedw.) Loeske
Amblystegiaceae
Warnstorfia pseudostraminea(Mull. Hal.) Tuom. & T.J. Kop.
Amblystegiaceae
Weissia controversaHedw.
Pottiaceae
Weissia muhlenbergiana(Sw.) W.D. Reese & B.A.E. Lemmon
Pottiaceae
50
Hornworts and Liverworts
Floristics
Coming soon!
About the Maps & Nomenclature
Nomenclature follows Schuster (1966-1992, Hepaticae and Anthocerotae of North America East of the Hun-drenth Meridian vols. I-VI), sypplemented by Soderstrom et al. (2016, Phytokeys 59:1-828)
The hornwort and liverwort flora presently contains 139 species in 32 families.
The following taxa have been excluded from the flora due to probable misidentifications, nomenclaturalissues, or problems with georeferencing:
Diplophyllum andrewsii, Frullania dilatata, Jungermannia caespiticia, Riccia curtisii, Riccia gougetiana
52
Anastrophyllum hellerianum(Nees) R.M. Schust.Anastrophyllaceae
Anastrophyllum michauxii(F. Weber) H. BuchAnastrophyllaceae
Anastrophyllum minutum(Schreb.) R.M. Schust.
Anastrophyllaceae
Aneura pinguis(L.) Dum.
Aneuraceae
Anthoceros macouniiM. Howe
Anthocerotaceae
Anthoceros punctatusL.
Anthocerotaceae
Asterella tenella(L.) P. Beauv.Aytoniaceae
Athalamia hyalina(Sommerf.) S. Hatt.
Cleveaceae
Barbilophozia attenuata(Nees) Loeske
Anastrophyllaceae
53
Barbilophozia barbata(Schreb.) Loeske
Anastrophyllaceae
Barbilophozia kunzeana(Hub.) K. Muller
Anastrophyllaceae
Bazzania trilobata(L.) Gray
Lepidoziaceae
Blasia pusillaL.
Blasiaceae
Blepharostoma trichophyllum(L.) Dum.
Pseudolepicoleaceae
Calypogeia fissa(L.) Raddi
Calypogeiaceae
Calypogeia integristipulaSteph.
Calypogeiaceae
Calypogeia muelleriana(Schiffn.) K. MullerCalypogeiaceae
Calypogeia neesiana(C. Massal. & Carestia) K. Muller
Calypogeiaceae
54
Calypogeia sphagnicola(Arnell & J. Perss.) Warnst. & Loeske
Calypogeiaceae
Calypogeia suecica(Arnell & J. Perss.) K. Muller
Calypogeiaceae
Cephalozia bicuspidata(L.) Dum.
Cephaloziaceae
Cephalozia catenulata(Hub.) Lindb.
Cephaloziaceae
Cephalozia connivens(Dicks.) Lindb.
Cephaloziaceae
Cephalozia lacinulata(J.B. Jack) SpruceCephaloziaceae
Cephalozia lunulifolia(Dum.) Dum.
Cephaloziaceae
Cephalozia macounii(Aust.) Spruce
Cephaloziaceae
Cephalozia pleniceps(Aust.) Lindb.
Cephaloziaceae
55
Cephaloziella byssacea(Roth) Warnst.
Cephaloziellaceae
Cephaloziella elachista(J.B. Jack) Schiffn.Cephaloziellaceae
Cephaloziella hampeana(Nees) Schiffn.
Cephaloziellaceae
Cephaloziella rubella(Nees) Warnst.
Cephaloziellaceae
Chiloscyphus coadunatus(Sw.) R.M. Schust. & J.J. Engel
Lophocoleaceae
Chiloscyphus pallescens(Ehrh.) Dum.
Lophocoleaceae
Chiloscyphus polyanthos(L.) Corda
Lophocoleaceae
Cladopodiella fluitans(Nees) Jarg.
Cephaloziaceae
Cololejeunea biddlecomiae(Aust.) A. EvansLejeuneaceae
56
Conocephalum salebrosumSzweyk., Buczkowska & Odrzykoski
Conocephalaceae
Dichiton integerrimum(Lindb.) H. Buch
Cephaloziellaceae
Diplophyllum apiculatum(A. Evans) Steph.
Scapaniaceae
Fossombronia foveolataLindb.
Fossombroniaceae
Frullania asagrayanaMont.
Jubulaceae
Frullania bolanderiAust.
Jubulaceae
Frullania brittoniaeA. Evans
Jubulaceae
Frullania eboracensisGott.
Jubulaceae
Frullania inflataGott.
Jubulaceae
57
Frullania oakesianaAust.
Jubulaceae
Frullania planaSull.
Jubulaceae
Frullania ripariaHampe
Jubulaceae
Frullania selwynianaPearson
Jubulaceae
Geocalyx graveolens(Schrad.) NeesGeocalycaceae
Harpanthus drummondii(Taylor) Grolle
Geocalycaceae
Harpanthus scutatus(F. Weber & D. Mohr) Spruce
Geocalycaceae
Jamesoniella autumnalis(DC) Steph.
Jungermanniaceae
Jungermannia lanceolataL.
Jungermanniaceae
58
Jungermannia schraderiMart.
Jungermanniaceae
Lejeunea cavifolia(Ehrh.) Lindb.Lejeuneaceae
Lepidozia reptans(L.) Dum.
Lepidoziaceae
Lophocolea bidentata(L.) Dum.
Lophocoleaceae
Lophocolea heterophylla(Schrad.) Dum.
Lophocoleaceae
Lophocolea minorNees
Lophocoleaceae
Lophozia alpestris(Schleich.) Evans
Jungermanniaceae
Lophozia badensis(Gott.) Schiffn.
Jungermanniaceae
Lophozia bicrenata(Schmidel) Dum.
Jungermanniaceae
59
Lophozia collaris(Mart.) Dum.
Jungermanniaceae
Lophozia excisa(Dicks.) Dum.
Jungermanniaceae
Lophozia gillmanii(Aust.) R.M. Schust.Jungermanniaceae
Lophozia heterocolpos(Thed.) M. Howe
Jungermanniaceae
Lophozia incisa(Schrad.) Dum.
Jungermanniaceae
Lophozia longidens(Lindb.) Macoun
Jungermanniaceae
Lophozia marchica(Nees.) Steph.
Jungermanniaceae
Lophozia porphyroleuca(Nees) Schiffn.
Jungermanniaceae
Lophozia ventricosa(Dicks.) Dum.
Jungermanniaceae
60
Lunularia cruciata(L.) Dum.
Lunulariaceae
Mannia fragrans(Balb.) Frye & L. Clark
Aytoniaceae
Mannia pilosa(Hornem.) Frye & L. Clark
Aytoniaceae
Mannia triandra(Scop.) GrolleAytoniaceae
Marchantia polymorphaL.
Marchantiaceae
Marsupella emarginata(Ehrh.) Dum.
Gymnomitriaceae
Metzgeria conjugataLindb.
Metzgeriaceae
Moerckia hibernica(Hook.) Gott.
Pallaviciniaceae
Mylia anomala(Hook.) A. Gray
Jubulaceae
61
Nardia geoscyphus(De Not.) Lindb.
Jungermanniaceae
Notothylas orbicularis(Schwein.) Sull.Notothyladaceae
Nowellia curvifolia(Dicks.) Mitt.
Cephaloziaceae
Odontoschisma denudatum(Nees) Dum.
Cephaloziaceae
Pallavicinia lyellii(Hook.) Gray
Pallaviciniaceae
Pellia epiphylla(L.) CordaPelliaceae
Pellia megasporaR.M. Schust.
Pelliaceae
Pellia neesiana(Gott.) Limpr.
Pelliaceae
Phaeoceros laevis(L.) Prosk.
Anthocerotaceae
62
Plagiochila porelloides(Torr.) Lindenb.Plagiochilaceae
Porella pinnataL.
Porellaceae
Porella platyphylla(L.) Pfeiff.
Porellaceae
Preissia quadrata(Scop.) Nees
Marchantiaceae
Ptilidium ciliare(L.) HampePtilidiaceae
Ptilidium pulcherrimum(Weber) Hampe
Ptilidiaceae
Radula complanata(L.) Dum.
Radulaceae
Radula obconicaSull.
Radulaceae
Reboulia hemisphaerica(L.) Raddi
Aytoniaceae
63
Riccardia latifrons(Lindb.) Lindb.Aneuraceae
Riccardia multifida(L.) Gray
Aneuraceae
Riccardia palmata(Hedw.) Carruth.
Aneuraceae
Riccia beyrichianaHampe
Ricciaceae
Riccia bifurcaHoffm.
Ricciaceae
Riccia cavernosaHoffm.
Ricciaceae
Riccia fluitansL.
Ricciaceae
Riccia frostiiAust.
Ricciaceae
Riccia membranaceaGott., Lindenb. & Nees
Ricciaceae
64
Riccia sorocarpaBisch.
Ricciaceae
Riccia sullivantiiAust.
Ricciaceae
Ricciocarpos natans(L.) CordaRicciaceae
Scapania apiculataSpruce
Scapaniaceae
Scapania brevicaulisTaylor
Scapaniaceae
Scapania curta(Mart.) Dum.
Scapaniaceae
Scapania cuspiduligera(Nees) K. MullerScapaniaceae
Scapania glaucocephala(Taylor) Aust.Scapaniaceae
Scapania gymnostomophilaKaal.
Scapaniaceae
65
Scapania irrigua(Nees) Nees
Scapaniaceae
Scapania kauriniiRyan
Scapaniaceae
Scapania mucronataH. Buch
Scapaniaceae
Scapania nemorea(L.) Grolle
Scapaniaceae
Scapania paludicolaLoeske & K. Muller
Scapaniaceae
Scapania paludosa(Mull. Frib.) Mull. Frib.
Scapaniaceae
Scapania subalpina(Nees) Dum.
Scapaniaceae
Scapania umbrosa(Schrad.) Dum.Scapaniaceae
Scapania undulata(L.) Dum.
Scapaniaceae
66
Solenostoma atrovirens(Dum.) Mull. Frib.
Jungermanniaceae
Solenostoma cordifolium(Dum.) Steph.
Jungermanniaceae
Solenostoma crenuliformisSteph.
Jungermanniaceae
Solenostoma fossombronioides(Aust.) R.M. Schust.Jungermanniaceae
Solenostoma gracillimumR.M. Schust.
Jungermanniaceae
Solenostoma hyalina(Lyell) Mitt.
Jungermanniaceae
Solenostoma pumila(With.) K. Muller
Jungermanniaceae
Solenostoma sphaerocarpum(Hook.) Steph.
Jungermanniaceae
Trichocolea tomentella(Ehrh.) Dum.
Trichocoleaceae
67