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Officers and Editors for 2013-2014 President ROBERT D. ALDRIDGE Department of Biology Saint Louis University St. Louis, MO 63103, USA President-Elect AARON BAUER Department of Biology Villanova University Villanova, PA 19085, USA Secretary MARION PREEST Joint Science Department The Claremont Colleges Claremont, CA 91711, USA Treasurer ANN PATERSON Department of Natural Science Williams Baptist College Walnut Ridge, AR 72476, USA Publications Secretary BRECK BARTHOLOMEW P.O. Box 58517 Salt Lake City, UT 84158, USA Immediate Past-President JOSEPH R. MENDELSON III Zoo Atlanta Atlanta, GA 30315, USA Directors (Class and Category) FRANK BURBRINK (2016 R) College of Staten Island, USA ALISON CREE (2016 Non-US) University of Otago, NEW ZEALAND TIFFANY DOAN (2014 R) Central Connecticut State Univ., USA LISA HAZARD (2016 R) Montclair State University, USA TRAVIS LADUC (2014 Mem. at-Large) University of Texas, USA JENNIFER PRAMUK (2014 Cons) Woodland Park Zoo, USA CAROL SPENCER (2014 R) University of California, Berkeley, USA GREGORY WATKINS-COLWELL (2016 R) Yale Peabody Mus. of Nat. Hist., USA Trustee GEORGE PISANI University of Kansas, USA Journal of Herpetology ERIN MUTHS, Co-Editor U.S. Geological Survey Fort Collins, CO 80526, USA GAD PERRY, Co-Editor Texas Tech University Lubbock, TX 79409, USA Herpetological Review ROBERT W. HANSEN, Editor 16333 Deer Path Lane Clovis, CA 93619, USA Contributions to Herpetology KRAIG ADLER, Editor Cornell University Ithaca, NY 14853-2702, USA Facsimile Reprints in Herpetology AARON BAUER, Editor Villanova University Villanova, PA 19085, USA Catalogue of American Amphibians and Reptiles CHRISTOPHER BELL, Co-Editor University of Texas, Austin Austin, TX 78712, USA TRAVIS LADUC, Co-Editor University of Texas, Austin Austin, TX 78758, USA Herpetological Circulars JOHN J. MORIARTY, Editor Three Rivers Park District Plymouth, MN 55441, USA Herpetological Conservation JOSEPH C. MITCHELL, Editor Mitchell Ecol. Res. Service P.O. Box 5638 Gainesville, FL 32627-5638, USA Publisher of Journal of Herpetology, Facsimile Reprints in Herpetology, Herpetological Review, Catalogue of American Amphibians and Reptiles, Herpetological Circulars, Contributions to Herpetology, and Herpetological Conservation Dear Author, Attached please find a gratis pdf file of your article/note published in Herpetological Review. You are receiving this pdf at no charge as a benefit of SSAR membership, and it is for your personal use only (see copyright notice below). Sincerely, SSAR Publications Office Notice warning concerning copyright restrictions: The copyright law of the United States (title 17, United States Code) governs the making of copies or other reproductions of copyrighted material such as PDFs. One of these specific conditions is that the copy or reproduction is not to be "used for any purpose other than private study, scholarship, or research." If a user makes, or later uses, a PDF, copy, or reproduction for purposes in excess of "fair use," that user may be liable for copyright infringement. The Society for the Study of Amphibians and Reptiles (SSAR) holds the copyright to this PDF. SSAR authorizes the author to use this PDF to fill reprint requests for private study, scholarship and research purposes. It is a violation of SSAR's copyright to distribute this PDF via mass emails, or by posting this pdf on any website for download — Except the author's own personal (not business) website / webpage.

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Officers and Editors for 2013-2014 President

ROBERT D. ALDRIDGE Department of Biology Saint Louis University St. Louis, MO 63103, USA

President-Elect AARON BAUER Department of Biology Villanova University Villanova, PA 19085, USA

Secretary MARION PREEST Joint Science Department The Claremont Colleges Claremont, CA 91711, USA

Treasurer ANN PATERSON Department of Natural Science Williams Baptist College Walnut Ridge, AR 72476, USA

Publications Secretary BRECK BARTHOLOMEW P.O. Box 58517 Salt Lake City, UT 84158, USA

Immediate Past-President JOSEPH R. MENDELSON III Zoo Atlanta Atlanta, GA 30315, USA

Directors (Class and Category) FRANK BURBRINK (2016 R) College of Staten Island, USA ALISON CREE (2016 Non-US) University of Otago, NEW ZEALAND TIFFANY DOAN (2014 R) Central Connecticut State Univ., USA LISA HAZARD (2016 R) Montclair State University, USA TRAVIS LADUC (2014 Mem. at-Large) University of Texas, USA JENNIFER PRAMUK (2014 Cons) Woodland Park Zoo, USA CAROL SPENCER (2014 R) University of California, Berkeley, USA GREGORY WATKINS-COLWELL

(2016 R) Yale Peabody Mus. of Nat. Hist., USA

Trustee GEORGE PISANI University of Kansas, USA

Journal of Herpetology ERIN MUTHS, Co-Editor U.S. Geological Survey Fort Collins, CO 80526, USA GAD PERRY, Co-Editor Texas Tech University Lubbock, TX 79409, USA

Herpetological Review ROBERT W. HANSEN, Editor 16333 Deer Path Lane Clovis, CA 93619, USA

Contributions to Herpetology KRAIG ADLER, Editor Cornell University Ithaca, NY 14853-2702, USA

Facsimile Reprints in Herpetology AARON BAUER, Editor Villanova University Villanova, PA 19085, USA

Catalogue of American Amphibians and Reptiles

CHRISTOPHER BELL, Co-Editor University of Texas, Austin Austin, TX 78712, USA TRAVIS LADUC, Co-Editor University of Texas, Austin Austin, TX 78758, USA

Herpetological Circulars JOHN J. MORIARTY, Editor Three Rivers Park District  Plymouth, MN 55441, USA

Herpetological Conservation JOSEPH C. MITCHELL, Editor Mitchell Ecol. Res. Service P.O. Box 5638 Gainesville, FL 32627-5638, USA

Publisher of Journal of Herpetology, Facsimile Reprints in Herpetology, Herpetological Review, Catalogue of American Amphibians and Reptiles, Herpetological Circulars, Contributions to Herpetology, and Herpetological Conservation

 

Dear Author,

Attached please find a gratis pdf file of your article/note published in Herpetological Review. You are receiving this pdf at no charge as a benefit of SSAR membership, and it is for your personal use only (see copyright notice below).

Sincerely,

SSAR Publications Office Notice warning concerning copyright restrictions: The copyright law of the United States (title 17, United States Code) governs the making of copies or other reproductions of copyrighted material such as PDFs. One of these specific conditions is that the copy or reproduction is not to be "used for any purpose other than private study, scholarship, or research." If a user makes, or later uses, a PDF, copy, or reproduction for purposes in excess of "fair use," that user may be liable for copyright infringement. The Society for the Study of Amphibians and Reptiles (SSAR) holds the copyright to this PDF. SSAR authorizes the author to use this PDF to fill reprint requests for private study, scholarship and research purposes. It is a violation of SSAR's copyright to distribute this PDF via mass emails, or by posting this pdf on any website for download — Except the author's own personal (not business) website / webpage.

Herpetological Review 44(3), 2013

NATURAL HISTORY NOTES 517

area of open woodland along a permanent wetland at El Questro Wilderness Park in the east Kimberley region of Western Australia (15.980556°S, 127.458889°E). Upon approaching the animal, we noticed a juvenile V. panoptes (SVL: 192 mm, TL: 479 mm) in the mouth of the male. The male dropped the juvenile before fleeing a distance of approximately 30 m. The juvenile V. panoptes was dead upon discovery and exhibited wounds consistent with being mouthed by another monitor lizard.

On 8 June 2001, we observed a large V. panoptes on the bank of the Daly River near Oolloo Crossing, Northern Territory, Australia (14.004391°S, 131.240014°E). The lizard was a male, judging by its size (ca. 60 cm), and was in the act of swallowing a sub-adult V. panoptes that was about one-third the size of the cannibal. The head and front legs were inside the larger lizard. The smaller lizard was still struggling, raking at the head of the larger lizard with its hind legs. The larger lizard appeared to be struggling to swallow its prey, and would shake it from side-to-side occasionally. Once the prey stopped struggling, the larger monitor slowly swallowed it until just a portion of tail (ca. 15 cm) was left hanging out of its mouth. At this stage the larger lizard appeared to have difficulty walking, and we did not observe the last portion of the tail being swallowed. During the latter part of the swallowing, a third V. panoptes walked past that was a similar size to the one that was eaten.

Although our 2001 observation may reflect scavenging, the prey in our 2012 observation was definitely alive. Moreover, there are observations of V. panoptes capturing and eating other moni-tor species (Christian 1995. Herpetol. Rev. 26:146; Rhind and Doody 2011. Herpetofauna 41:64–65). Benefits of cannibalism not involving offspring or eggs include nutrition, reduction of competition for resources, or reproductive competition (Polis 1981, op. cit.). In tropical ecosystems in Northern Austra-lia, V. panoptes plays an important role as a key predator and is responsible for the regulation of a number of prey species (Blamires 2004. Copeia, 2004:370–377; Doody et al. 2006. Wildl. Res. 33:349–354; Doody et al. 2009. Anim. Conserv. 12:46–53; Doody et al. 2012a. Herpetol. Rev. 43:339–340; Doody et al. 2012b. Herpetol. Rev. 43:491–492; Doody et al. 2013. Biol. Invas. 15:559–568; Webb and Manolis 2010. Freshwater Crocodile, Crocodylus johnstoni, Status Survey and Action Plan. Crocodile Specialist Group, Darwin). Large male V. panoptes may likewise consume conspecifics frequently, but our observations are not sufficient to determine the frequency of cannibalism or its costs-benefits in this top predator.

DAVID RHIND, School of Anatomy and Developmental Biol-ogy, Monash University, Clayton, Victoria 3800, Australia (e-mail: [email protected]); J. SEAN DOODY, Department of Ecology and Evolution-ary Biology, University of Tennessee, 569 Dabney Hall, Knoxville, Tennessee 37996-1610, USA (e-mail: [email protected]); RACHEL PRITCHARD, c/o Narrawong Post Office, Narrawong, Victoria 3285, Australia (e-mail: [email protected]); COLIN McHENRY, Department of Anatomy and De-velopmental Biology, Monash University, Clayton, Victoria 3800, Australia (e-mail: [email protected]).

ZOOTOCA VIVIPARA (Common Lizard). RECORD WEIGHT. On 25 June 2011 a very heavy adult female specimen of Zoo-toca vivipara was found on the island of Hiddensee near Vitte (54.563816°N, 13.112562°E) at the German coastline of the Bal-tic Sea underneath a dry-docked sailing boat. The female Z. vi-vipara reported here weighed 10.0 g (live weight), which makes it the heaviest gravid female specimen of the Common Lizard ever recorded (Fig. 1). The examined Z. vivipara measured 157

mm in total length (SVL = 72 mm; tail length = 85 mm). Most adult specimens of Z. vivipara express a total length of 110–140 mm with a maximum length of approximately 180 mm (Günther and Völkl 1996. In R. Günther [ed.], Die Amphibien und Reptilien Deutschlands, pp. 588–600. Fischer, Jena). Female individuals typically have a SVL measuring between 45 and 70 mm, seldom more than 75 mm (Günther and Völkl, 1996, op. cit.). Few reports regarding body weight data have been published for Z. vivipara.

FRANZISKA JÄHRLING (e-mail: [email protected]), ANJA JUNGHANNS (e-mail: [email protected]), and FALK ORT-LIEB (e-mail: [email protected]), Zoological Institute and Museum Grei-fswald, Johann Sebastian Bach-Straße 11/12, 17489 Greifswald, Germany.

SQUAMATA — SNAKES

AGKISTRODON CONTORTRIX (Copperhead). DIET. Agkis-trodon contortrix is a widely-distributed viperid species that occurs in the eastern and central United States and northern Mexico. The species’ diet is known to include many species of small mammals, birds, snakes, lizards, frogs, salamanders, and invertebrates (Campbell and Lamar 2004. The Venomous Rep-tiles of the Western Hemisphere, Volume I. Cornell University Press, Ithaca, New York. 475 pp.). On 19 April 2012, at 2040 h, a male A. contortrix (SVL ca. 63.4 cm; Auburn University Museum [AUM] 39901) was collected injured on State Road 503 near CR 20 in Jasper Co., Mississippi, USA (32.13914°N, 89.06058°W; datum WGS84). The snake died overnight. While curating the specimen, the remains of an adult Plestiodon inexpectatus (Southeastern Five-lined Skink; AUM 39902) were found in the stomach. The lizard had been ingested headfirst. To our knowledge, this is the first record of A. contortrix consuming P. inexpectatus (Campbell and Lamar, op. cit.).

We thank Craig Guyer for verifying the records and Scott Pey-ton for assistance in procuring collecting permits.

BRIAN FOLT (e-mail: [email protected]) and DAVID LAURENCIO (e-mail: [email protected]), Auburn Museum of Natural History, Depart-ment of Biological Sciences, 331 Funchess Hall, Auburn University, Ala-bama 36849, USA.

BOGERTOPHIS SUBOCULARIS (Trans-Pecos Ratsnake). OVER-WINTERING BEHAVIOR. Colubrid snakes in North America, especially those inhabiting higher latitudes, typically hibernate during the cold winter months (Ernst and Ernst 2003. Snakes of

FiG. 1 Gravid adult female Zootoca vivipara with a record weight of 10.0 g (found on the island of Hiddensee, Germany).