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64 Current Herpetol. 23 (2) 2004

Mentawai and Andaman and Nicobar speciesare not conspecific with the Sri Lankanspecies. Dring (1979), in his key to the genusfor Southeast Asia included Taylor's mysorien-sis in the synonymy of kendallii.

nemaspis is a speciose genus of gekkonids,and a large number of species have beendescribed in recent years from southern andsoutheastern Asia (compare Kluge, 1993,2001; Bauer, 2003; to Wermuth, 1966). Wereport here a new species of the genus fromPhuket Island, southern Thailand. This spe-cies has been earlier reported as C. kandianaby Cox et al. (1998) and Cnemaspis sp. inGrossmann and Tillack (2000).

MATERIALS AND METHODSSpecimens were hand-collected. The holo-type was photographed in life and fixed in for-malin, before storage in ethanol. The follow-ing measurements were taken ca. 18 monthsafter collection with a MitutoyoTMdial caliper

(to the nearest 0.1mm): snout-vent length(SVL; from tip of snout to vent); tail length(TL; from vent to tip of unregenerated tail);tail width (TW; measured at base of tail); headlength (HL; distance between posterior edge oflast supralabial and snout-tip); head width(HW; measured at angle of jaws); head depth(HD; maximum height of head, from occiputto throat); ear length (EL; greater ear length);forearm length (FA; distance between thepalm and elbow); eye diameter (ED; greatestdiameter of orbit); eye to nostril distance (E-N; distance between anteriormost point ofeyes and nostrils); eye to snout distance (E-S;distance between anteriormost point of eyesand tip of snout); eye to ear distance (E-E;distance from anterior edge of ear opening toposterior corner of eyes); internarial distance(IN; distance between nares); and interorbitaldistance (IO; shortest distance between orbits).Scale counts and external observations ofmorphology were made using an OlympusSZX9 dissecting microscope. Notes on colourin life are from colour swatches of F. B. Smith(1975; 1981).

Comparative material examined is shown inthe Appendix. Additional sources of informa-tion on character states include: Smith (1935),Nicholls (1949), Dring (1979), Das (1993),Manthey and Grossmann (1997), Das andBauer (1998), Bauer and Das (1998), and Dasand Grismer (2003). Catalogue numbers ofspecimens deposited in the Raffles Museum ofBiodiversity Research, National University ofSingapore, Singapore, are preceded by ZRCinstead of USDZ as formerly proposed byLeviton et al. (1985). The other institutionalabbreviations follow Leviton et al. (1985).

Cnemaspis phuketensis sp. nov.ig. 1

HolotypeZRC 2. 5212, Kathu Fal l s (07 55' N 98 20' E),

Changwat Phuket, Thailand, adult male, T.-M . Leong, collector, 1 September 2002.Paratypes

ZRC 2.5214-18, paratopotypes, other dataas for holotype, except collected 2 September2002; ZRC 2.5233, Manik Falls, ChangwatPhuket, Thailand, Tzi-Ming Leong, collector,4 September 2002.Diagnosis

A small species of Cnemaspis (SVL to 29.1mm), distinguishable from conspecific speciesin showing the following combination of char-acters: two semi-circular supranasals separatedby a single scale; three postnasals borderingnasal; four scale rows separate orbit fromsupralabials; posteriorly, each postmentalbounded by three smooth, rounded and juxta-posed scales; scattered spinose paravertebralrows of tubercles on dorsum; gular andpectoral scales unicarinate; abdominal scalesnot elongated, smooth; tail segmented, withenlarged flattened scales forming whorls, asingle pair of spinose postcloacal spurs present;median subcaudals not enlarged, smooth;supralabials (to midorbit position) 6-7; infrala-bials 6-7; lamellae under toe IV 16-17;midventrals 26-32; and adult males lackpreanal and femoral pores.

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DAS & LEONG-NEW LIZARD FROM SOUTHERN THAILAND 65

FIG. 1. The holotype of Cnemaspis phuketensis sp. nov. (ZRC 2.5212) in life.

Description of holotypesmall species of Cnemaspis (snout-ventlength 29.0mm); snout elongate, large (HL/

SVL ratio 0.18), narrow (HW/SVL ratio 0.16),depressed (HD/HL ratio 0.66), distinct fromneck; lores sloping and interorbital regionflattened; snout long (E-S/HW ratio 0.87),longer than eye diameter (ED/E-S ratio 0.43);scales on snout and forehead warty, undermagnification revealed as tubercles that areraised towards posterior of each scale; scaleson snout larger than those on occipital region;eye large (ED/HL ratio 0.32); 'extra-brillarfringes' (of Underwood, 1954) indistinct; pupilround; elongated supraciliaries on top half oforbit; ear-opening deep, slit-like, its greatestdiameter vertically, fairly narrow (EL/E-Sratio 0.13); eye to ear distance greater thandiameter of eyes (E-E/ED ratio 1.59); no ridgeof tubercles along mandible or from posteriorof orbit to postero-venter of tympanum, orfrom antero-dorsum of tympanum to nape;rostral completely divided by a simple rostralgroove which meets anterior of snout; rostralless than half as deep as wide (rostral width=

1.3mm/rostral depth=0.5mm; width/depthratio 2.6); contacted posteriorly by two nostrilsand two semi-circular supranasals that areseparated by a single scale; ventro-posteriorly,rostral is in contact with supralabial I; nostrilsoval, situated within nasals, and oriented later-ally; nasals reduced, in broad contact withsupralabial I; anterior nasal larger than poste-rior nasal; three postnasals bound nasal; fourscale rows separate orbit from supralabials;mentals subtriangular, deeper than wide, pairedpostmentals that are semicircular, smaller thanmental and separated by a single scale; poste-riorly, each postmental is bounded by threesmooth, rounded and juxtaposed scales; tonguenarrowly elongate, with a weak median cleft.ody slender, elongate (A-G/SVL ratio 0.51);scale size does not decrease dorsally afterthorax; ventrally, scales do not decrease insize from chin region to gular, pectoral andabdominal regions; scales on dorsum at mid-body unicarinate, smaller than those of ven-trum at same level; scales along vertebralregion not differentiable from adjacent scales;scattered spinose tubercles on paravertebral

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region not arranged in rows; pectoral andabdominal scales not elongate, smooth; spinousprocesses on lateral surface of body; scales onmanus and pes smooth, rounded; scales oninner surface of forearm, distal aspect of upperarm, dorsal surface of thighs, tibia; upper armand forearm unicarinate.orelimbs moderately long, slender; hind-limbs relatively short; tibia short (TBL/SVLratio 0.18); no shield-like subtibial scales;digits elongate, all bearing claws that areslightly recurved; subdigital scansors entire,unnotched; no fragmented basal lamellae, sixenlarged scansors at base of digits, which aremore than twice width of other scansors;lamellae under digit IV of pes 16 (includingthe enlarged basal ones); interdigital webbingabsent; relative length of digits (finger): 3>2>4>5>1; (toe): 3>4>5>2>1.

Original tail long, tip missing, preservedportion of tail longer than snout-vent length(TL/SVL ratio 1.21); tail tip blunt, tail baseswollen, segmented, with enlarged flattenedscales with a median keel forming whorls, asingle pair of spinose postcloacal spur present;tail with a distinct pair of furrow laterally;median subcaudals not enlarged, smooth; scaleson postanal region and at proximal part of tailbase and on rest of subcaudals smooth; hemi-peneal swelling at tail base, preanal depressionpresent.

Scutellation.See Table 1.Colouration (in life)orsum olive (# 30), with sinuous, darkgrayish brown (# 20) markings on nape, axilla,torso and inguinal regions, labials dark-barred;spinose tubercles on flanks chamois (# 123D);limbs and tail dark-banded; a dark grayishbrown (# 20) canthal stripe, commencing fromsnout, traversing the orbit region and extend-ing to axilla; throat and undersurface of tailwith fine hair brown (# 119A) mottlings; restof venter unpatterned cream; iris buff yellow(# 53); pupil black.Measurements (in mm).

ee Table 2.Sexual dimorphism

RC 2.5212, 2.5214-17 are adult males(presence of hemipeneal swelling at tail base,with preanal depression; ZRC 2.5218 and2.5233 are adult females (lack of hemipenealswelling or preanal depression). Preanal andfemoral pores are absent in males. Femaleslack obvious endolymphatic sacs that arevisible externally in many adult female geckos.Etymology

Latin implying an inhabitant of PhuketIsland, in southern Thailand.

TABLE1. Squamation data of the holotype (ZRC 2.5212) and paratypes of Cnemaspis phuketensis sp.nov. See text for details. Abbreviations: +=present; -=absent; M=male; F=female; SL=supralabial;IL=infralabial; IO=interorbital; T4=lamellae under toe IV of pes; MV=midventral; PA=preanal depression

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TABLE 2. Measurements (in mm) of the holotype (ZRC 2.5212) and paratypes of Cnemaspis phuketen-sis sp. nov. See text for details. Abbreviations: O=original tail; R=regenerated tail; SVL=snout-vent length;TL=tail length; FA=forearm length; TBL=tibia length; A-G=axilla to groin distance; HL=head length;HW=head width; HD=head depth; ED=eye diameter; E-E=eye to ear distance; E-S=eye to snout dis-tance; E-N=eye to nostril distance; IO=interorbital distance; EL=ear length; and IN=internarial distance.

Natural history noteshe holotype was taken from a leaf of an

herb, 5 cm above substrate; the paratypes fromeither earth banks beside a small stream orfrom tree trunks at chest level, during the day.Leong et al. (2003) recorded the followingadditional saurian species as sympatric: Dracotaenioptera Gunther, 1861 (at Kathu Falls),and Cyrtodactylus oldhami (Theobald,1876)(at both Kathu and Manik Falls).ComparisonsThe new species from Phuket, Thailand, iscompared with congeners from Thailand. Onlyopposing suites of characters are listed. Cne-maspis affinis (Stoliczka, 1870), distribution:southern Thailand and Peninsular Malaysia:SVL to 48.0mm; a dark marking at axilla; noenlarged tubercles on tail; and dorsal surfacewith five transverse yellow bands; C. chan-thaburiensis Bauer & Das, 1998, distribution:

Chanthaburi Province, Thailand: SVL to 41.0mm; a ridge of tubercles border anteriormargin of ears and another from ear to nape; aseries of white paravertebral markings betweennape and tail; and belly mottled light brown;C. kumpoli (Taylor, 1963), distribution: Trangin southern Thailand: SVL o 52.0mm; suprala-bials 11; posterior supralabials with longitudi-nal keels; and three postcloacal spurs; and C.siamensis (Smith, 1925), distribution, Thailandand northern Peninsular Malaysia: SVL to39.7 mm; forehead with keeled scales; suprala-bials 9-11; infralabials 8-10; paravertebraltubercles in 12 or 14 rows; and ventrals andsubcaudals tricarinate.The new Thai species is next compared with

other south-east Asia congeners. C. argusDring, 1979, distribution: Gunung Lawit innorthern Terengganu, Malay Peninsula: SVLto 65.3mm; fourth and fifth fingers subequal;supralabial VIII in midorbital position; and

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yellow radiating lines from the orbits; C.boulengerii Strauch, 1887, distribution: ConSon Island, Vietnam: SVL o 66.0mm; suprala-bials 8-10; infralabials 7-8; a series of 6-7shield-like subtibial scales; subcaudal scalesalmost as wide as tail, and large black nuchaland shoulder spots; C. dringi Das & Bauer,1998, distribution: Kapit District, centralSarawak, East Malaysia (Borneo): SVL to45.5 mm; five postnasals; postcloacal spursabsent; toe IV lamellae 20; flanks with distinctwhite patches; and ventrals heavily pigmented;C. flavolineata (Nichols, 1949), distribution,the northern Malay Peninsula: SVL to 46.7mm; yellow line along back; paired, elongatepostmentals; and lamellae under toe IV 28;C. gordongekkoi Das, 1993, distribution, Lom-bok Island, Lesser Sundas, Indonesia: SVL to73.0mm; lamellae under toe IV 22-23; suprala-bials at midorbital position 9; infralabials 10;interorbitals 20, and throat scales smooth; C.kendallii (Gray, 1845), distribution: Borneo,the Riau Archipelago and peninsular Malaysia:SVL to 58.0mm; tubercles on dorsum keeled;postnasals six; postmentals bounded by 4-5scales posteriorly; two rows of tubercles runposteriorly from orbit towards tympanum; andmedian subcaudal scales raised; C. nigridia(Smith, 1925), distribution, Bau and GunungGading, western Sarawak, East Malaysia(Borneo) as well as Natuna Island: SVL to69.8mm; supranasals in wide contact; post-mentals bounded by four scales posteriorly;postnasals five; two rows of tubercles runposteriorly from orbit towards tympanum; andtail without dark bands; and C. timoriensis(Dumeril & Bibron,1836), distribution, Timor,without specific information, and may beeither West Timor, Republic of Indonesia, orthe newly-independent nation of East Timor:SVL to 35mm; dorsum with no enlargedtubercles; supralabials five; lamellae under toeIV 12; and dorsal surface reddish-brown, witha series of brown paravertebral spots.wo further species of the genus have beenrecently described from the Seribuat Archipel-ago, off the east coast of Peninsular Malaysia(Das and Grismer, 2003). One of these,

Cnemaspis limi from Pulau Tioman, PahangState, can be differentiated from the new Thaispecies in being significantly larger-SVL to88.2mm; supranasals in contact; supralabials11-14; and scales on forearm smooth. Thesecond Seribuat species, C. baueri, is fromPulau Aur, Johor State, and differs from thenew species in being much larger, SVL to 64.9mm; fourth toe longer than fifth; lamellaeunder toe IV 26-27; supralabials 11-13; scaleson forearm smooth; and caudal bands absent.

ffinities of the new Thai Cnemaspis remainunknown, for lack of a phylogenetic hypothe-ses for the genus. Several distinct and appar-ently unrelated species, once allocated to C.kandiana, are now known, the association withthis Sri Lankan endemic is for sharing spinoseflanks, comprising tuberculate scales, keeledgular scales, and the presence of preanal andfemoral scales.

ACKNOWLEDGMENTShe Raffles Museum of BiodiversityResearch

(through a Raffles Museum Fellowship to thefirst author) and Institute of Biodiversity andEnvironmental Conservation, UniversitiMalaysiaSarawak supported manuscript preparation.For permitting us to examine material undertheir care, we thank Edwin Nicholas Arnold,and Colin John McCarthy (BMNH), KarlaKishinami (BPBM); Alan Leviton, Jens Vin-dum, and the late Joseph Bruno Slowinski(CAS), Harold Knight Voris, Robert FrederickInger, and Alan Resetar (FMNH), John Ever-ett Cadle, the late Ernest Edward Williams,and Jose Paul Ovidio Rosado (MCZ), RobertoPoggi, and Giuliano Doria (MSNG), DavidAuth, and Frederick Wayne King (UF), KelvinKok Peng Lim, Peter Kee Lin Ng, Heok HuiTan, and Chang Man Yang (ZRC); Ronald IanCrombie, William Ronald Heyer; and GeorgeRobert Zug (USNM), Axel Groenveld, andLeobertus van Tuijl (ZMA) and J. R. B.Alfred, and Shyamal Kumar Chanda (ZSI).Finally, we are grateful to Aaron M. Bauer andtwo anonymous reviewers for reading themanuscript.

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LITERATURE CITEDABDULALI,H. 1955. Extension of range of the

izard Cnemaspis kandiana (Kelaart). J. Bom-ay Nat. Hist. Soc. 53: 134.ANNANDALE, . 1909. Report on a small collec-

ion of lizards from Travancore. Rec. Indianus. 3: 253-257.

BAUER, A. M. 2002. Two new species ofnemaspis (Reptilia: Squamata: Gekkonidae)

rom Gund, Uttara Kannada, India. Mitt.amburg. Zool. Mus. Inst. 99: 155-167.

BAUER, A. M. AND I. DAS. 1998. A newnemaspis (Reptilia: Gekkonidae) from South-

astern Thailand. Copeia 1998: 439-444.BOULENGER, . A. 1885. A list of reptiles and

atrachians from the island of Nias. Ann. Mag.at. Hist., Ser. 5, 16: 388-389.

BOULENGER, . A. 1890. A list of the reptiles andatrachians collected by Dr. E. Modigliani onereinu (Sipora), Mentawei Islands. Ann. Mus.iv. Stor. Nat. Genova Ser. 2, 14: 613-618.

CHAN-ARD, ., W. GROSSMANN, . GUMPRECHT,NDK. D. SCHULZ. 999. Amphibians and rep-

iles of Peninsular Malaysia and Thailand: anllustrated checklist/Amphibien and Reptiliener Halbinsel Malaysia and Thailands: eine

llustrierte Checkliste. Bushmaster Publications,urselen.

COX, M. J., P. P. VAN DIJK, J. NABHITABHATA,ND K. THIRAKHUPT. 998. A photographicuide to snakes and other reptiles of Peninsularalaysia, Singapore and Thailand. New Holland

ublishers (UK) Ltd., London.DAS, I. 1993. Cnemaspis gordongekkoi, a new

ecko from Lombok, Indonesia, and the bioge-graphy of Oriental species of Cnemaspis

Squamata: Sauria: Gekkonidae). Hamadryad8: 1-9; P 1.1.

DAS, I. 2005. A revision of the genus Cnemaspistrauch, 1887 (Sauria: Gekkonidae) from theentawai Archipelago and adjacent archipela-

os off Sumatra, Indonesia, with the descriptionf four new species. J. Herpetol. In press.

DAS, I. ANDA. M. BAUER. 998. Systematics andiogeography of Bornean geckos of the genusnemaspis Strauch, 1887 (Sauria: Gekkonidae),ith the description of a new species. Raffles

Bull. Zool. 46: 11-28.DAS, I. AND L. L. GRISMER.2003. Two new

pecies of Cnemaspis Strauch, 1887 (Sauria:ekkonidae) from the Seribuat Archipelago,ahang and Johor States, West Malaysia.erpetologica 59: 546-554.

DE ROOIJ, N. 1915. The reptiles of the Indo-ustralian Archipelago. Vol. I. Lacertilia, Chelo-

ia, Emydosauria. E. J. Brill, Leiden.DRING, . C. 1979. Amphibians and reptiles from

orthern Trengganu, Malaysia, with descriptionsf two new geckos: Cnemaspis & Cyrtodactylus.ull. British Mus. Nat. Hist. (Zool.) 34: 181-241.

GROSSMANN, . ANDF. TILLACK.000. Bemerkun-en zur Herpetofauna des Khao Lak, Phangga, thailandische Halbinsel Teil 1: Einfuhrung;mphibia; Reptilia: Sauria. Sauria, Berlin 22(4):3-38.

KLUGE,A. G. 1993. Gekkonoad lizard taxonomy.nternational Gecko Society, San Diego.

KLUGE,A. G. 2001. Gekkotan lizard taxonomy.amadryad 26: 1-209.

LEONG,T. M., T. CHANARD,ANDY. CHUAYNKERN.003. Additional anuran and saurian records forhuket, south Thailand. Nat. Hist. J. Chula-ongkorn Univ. 3: 17-21.

LEVITON,A. E., S. C. ANDERSON, . H. GIBBS,E.EAL,AND C. E. DAWSON. 985. Standards inerpetology and ichthyology. Part I. Standardymbolic codes for institutional resource collec-

ions in herpetology and ichthyology. Copeia985: 802-832.

MANTHEY, . ANDW. GROSSMANN.997. Amphib-en & Reptilien Sudostasiens. Natur and Tier

erlag, Munster.NICHOLLS,L. 1949. A new gekkonid from the

alay Peninsula. Bull. Raffles Mus. 19: 47-49.ROSLER,H. 1981. Bemerkungen zur Geographis-

hen Verbreitung der Gattung CnemaspisStrauch 1887); Anmerkungen zur Systematikon C. kandiana (Kelaart 1852); Allgemeineberlegungen zu ihrer Biologie. Sauria, Berlin981: 7-14.

SMITH,F. B. 1975. Naturalist's color guide. Parts Ind II. American Museum of Natural History,ew York.SMITH,F. B. 1981. Naturalist's color guide. Part

II. American Museum of Natural History, New

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70 Current Herpetol. 23(2) 2004

York.SMITH, M. A. 1935. The fauna of British India,

ncluding Ceylon and Burma. Reptilia andmphibia. Vol. IL-Sauria. Taylor and Francis,ondon.

TAYLOR, E. H. 1963. The lizards of Thailand.niv. Kansas Sci. Bull. 44: 687-1077.

THAKUR, S. 1998. Turtles and lizards of theahyadri. J. Ecol. Soc., Pune 11: 38-40.

THOMAS, J. AND P. S. EASA. 1997. Additions tohe reptile fauna of Silent Valley, Kerala. Cobra7: 31-33.

UNDERWOOD, G. 1954. On the classification andvolution of geckos. Proc. Zool. Soc. London24: 469-492.

WERMUTH, H. 1966. Liste der rezenten Amphibienud Reptilien. Gekkonidae, Pygopodidae, Xan-

husiidae. Das Tierreich 80. Walter de Gruyter &o.

APPENDIXComparative material examinednemaspis affinis (Stoliczka,1870): ZSI 5964(holotype), ZRC 2.1098, "Penang" (=PulauPinang, West Malaysia); ZMA 11987, Pinang,West Malaysia; ZRC 2.4858, Moon Gate,Pulau Pinang, West Malaysia; Maxwell's Hill,Perak, West Malaysia (ZRC 2.1100; 2.1099).nemaspis boulengerii Strauch,1887: CAS73745 (paratype of Gonatodes glaucus Smith,1920), MCZ 39014-23, 'Pulo Condore" (=ConDao), Vietnam.nemaspis chanthaburiensis Bauer & Das,1998: FMNH 215979 (holotype), "Khao SoiDaouw (Dao) Wildlife Sanctuary, Pongnomron(Pong Nam Ron), Chantaburi (Chanthaburi)Provi nce, Thai l and (approxi matel y 13 00' N102 05' E)" ; BMNH 1917. 5. 14. 4(paratype) ,"Chantaburi (Chanthaburi Province) , Siam(Thailand)"; FMNH 191479 (paratype), "KhaoSoi Dao Tai, Pong Nam Ron, Chantaburi(Chanthaburi) Province, Thailand (approxi-matel y 13 00' N 102 05' E), 850m" ; FMNH215978 (paratype), "Khao Khiew (Khieo) Wild-life Sanctuary, Chon Buri Province, Thailand( approximatel y 13 14' N 101 08' E) " ; FMNH215980 (paratype), "Amphoe Muang, Suan

Kaset, Chantaburi (Chanthaburi) Province,Thai l and (approximatel y 123 6' N 102 09' E) " .

Cnemaspis gordongekkoi Das, 1993: ZRC2.3380 and ZRC 2.3381 (holotype andparatype), "..vicinity of Sendanggila Falls, circa0.5km south of Senaru village, Lombok, NusaTenggara District, Republic of Indonesia(8 45' S, 116 30' E)" .Cnemaspis dringi Das & Bauer, 1998:FMNH 148588 (holotype), "Labang Camp(03 20' N 113 29' E), B ntulu D stri ct, FourthDivision, Sarawak, East Malaysia, Borneo";FMNH 221478 (paratype), "Sungai Segaham(02 44' N 113 53' E), Bel aga D stri ct, Sevent hDivision, Sarawak, East Malaysia".

nemaspis kandiana (Kelaart,1852): BMNH60.3.17.1066, 80.2.2.119, 53.4.1.1 (three syn-types), "Kandian hills, Ceylon" (=hills of Kandy[ or Mhanuwara], 07 15' N 80 40' E Cent ralProvince, Sri Lanka); MCZ 4138, 26719,"Ceylon" (=Sri Lanka); ZSI 5971 (holotypeof Gymnodactylus Humei Theobald, 1876),"Kandy" (see comments above); MSNG 8764(four specimens), "Ceylon" (=Sri Lanka).

nemaspis kendallii (Gray, 1845): BMNHXXII.92a (lectotype, designated by Dring,1979), "Borneo"; FMNH 223201, MCZ157158-59. Bako National Park, Sarawak, EastMalaysia (Borneo); FMNH 223201; MCZ157158-59. Bidi, Sarawak, East Malaysia(Borneo); FMNH 184424. Bukit Lanjan,Selangor, West Malaysia; BMNH 1902.12.12.12. Bidi, Sarawak, East Malaysia (Borneo);Bau, Sarawak, East Malaysia (Borneo); BMNH1911.1.20.7-9. Bau, Sarawak, East Malaysia(Borneo); BPBM 7494, Alag Sungei Ayer,Pulau Tioman, Pahang, West Malaysia; ZRC2.1101. Jerantut, Pahang, West Malaysia; ZRC2.1102, Gunung Rokan, Pulau Tioman, Pah-ang, West Malaysia; ZRC 2.1103. Sedagong,Pulau Tioman, Pahang, West Malaysia; ZRC2.1109-10. Pulau Siantan, Anamba, RiaoArchipelago, Indonesia; ZRC 2.1112-13. SungeiUlu, Great Natuna, Riao Archipelago, Indo-nesia; ZRC 2.3014. Bukit Timah, Singapore;ZRC 2.3015. Gunung Ladang, Melaka, WestMalaysia; USNM 26573. Pulau Bunoa, Tam-belan Islands, Indonesia; USNM 26555. St.

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Barbe Island, at present Pulau Pedjantan,Indonesia; USNM 26547-49. Bunguran, Natu-nas, Riao Archipelago, Indonesia; USNM28145. Pulau Lingung, Natuna, Riao Archi-pelago, Indonesia; USNM 28149. Sirhassen,Natuna, Riao Archipelago, Indonesia; alsoU78463 and ZSI 14767 and 19637, fromBorneo".Cnemaspis nigridia (Smith, 1925): BMNH1946.8.22.90 (formerly BMNH 1925.9.1.8;holotype), MCZ 39024 and ZRC 2.1114-115," M . Gadi n" (=Gunung Gadi ng, 01 44' N109 50' E Sarawak, East Ml aysi a; Borneo) ;MCZ 15250, Lundu, Sarawak, East Malaysia;BMNH 1925.9.1.9-10, Gunung Pueh, Sarawak,East Malaysia.

nemaspis siamensis (Smith, 1925): MCZ39025, Maprit, Patiyu, peninsular Thailand;

MCZ 39694, Klong Bang Lai, peninsularThailand.

Cnemaspis limi Das & Grismer, 2003: ZRC2.5289 (holotype); ZRC 2.5290 (paratype)." Gua Tengkuk A r , Gunung Kaj ang (02 50' N104 09' E) , Pul au Ti oman, Pahang, WestMlaysia, al ti tude 980m; ZRC 2. 3504- 06(t hree paratypes). Tekek- Juara t rai l (02 52' N104 12' E) , Pul au Ti oman, Pahang, WestMl aysi a.

Cnemaspis baueri Das & Gri smer, 2003:ZRC 2. 5291 (hol otype); ZRC 2. 5292- 99 (eightparat opotypes) . Kampung Berhal a (02 27' N104 30E) , Pul au Aur, Johor, West Ml aysia.

Accepted: 23 November 2004

das & leong (2004)

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