Research Interests

Biological oceanography with emphasis on primary production as a function of nutrient supply, light and variations in marine climate. Pioneered integration of marine laboratory and field investigations in Norway, including studies of chemical composition of phytoplankton using unialgal cultures. Also specialised in the photobiology and bio-optics of marine phytoplankton, with emphasis on photo-acclimation and P vs E curves. The field activity has, besides local waters been concentrated on the ecology of the Barents Sea and the Southern Ocean.

Forskningsprofil

Biologisk oseanografi med vekt på primærproduksjon som funksjon av tilførselen av næringssalter, lys og klimavariasjoner i havet. Var pioner for å integrere laboratorie- og feltarbeid. Kjemisk sammensetning av planktonalger som funksjon av nærings- og lysbegrensning (på artsnivå). Bruk av enalgekulturer i laboratoriet for å oppnå målet, inkludert tolkning av fotsyntese-mot-lys data for anvendelse på feltdata. Initierte fotobiologi som en spesialitet ved TBS. Feltinnsatsen har, foruten lokale farvann vært konsentrert om polarstrøk (Barentshavet, Sørishavet).

Dr. students: Torunn Beate Hancke (fluorescens-kinetikk og fotobiologi hos giftige algerErlend Kjeldsberg Hovland:Bio-optikk hos marine kalkflagellater under

blomstringerM.Sc.student: Raluca Rogin (Romania; akvakultur av stør for kaviarproduksjon, tilhørende miljøproblemer)

Teaching in 2010:

BI 2060: Marine EcologyBI 2061: Physical Oceanography BI 2098: Experts in Team (Underwater Robotics)BI 3061: Biological Oceanography (masters level)

C.V.

Name/Position:

Egil Sakshaug, professor in marine ecology at NTNU (The Norwegian University of Science and Technology), Trondheim, Norway

Education:

1978: Dr. Philos., University of Trondheim1967 Cand. real., University of Oslo

Positions:

1982- Professor, tenured at the Biological station, Department of Biology, NTNU

1970-1982 Associate professor, tenured at the Biological Station, NTNU1969-1970: Grantee, International Biological Programme (IBP), NTNU 1996- Courtesy professor, University of South Florida, College of Marine Sciences, St.

Petersburg, Florida

Sabbaticals:

1973 Graduate School of Oceanography, University of Rhode Island 1973-74, 1986 Food Chain Research Group, Scripps Institution of Oceanography, La

Jolla, California, USA 1985 Bigelow Laboratory for Ocean Sciences, West Boothbay Harbor, Maine, USA 1996 Department of Marine Science, University of South Florida, St. Petersburg2005 Institut de Recherche pour Développement, Nouméa, Nouvelle Calédonie2007 University of Nagoya, Japan, visiting professor

Awards and memberships:

2006- Honorary member, Norwegian Association of Oceanographers, member since 19701996- Norwegian Academy of Science and Letters, Oslo (elected member)1994- Norwegian Academy of Technological Sciences, Trondheim (elected member)1994 Nansen-Prize of the "Fram" committee for merits in polar research1994- American Geophysical Union (AGU)1993- Royal Norwegian Society of Science and Letters, Trondheim (elected member)1993 Antarctic Service Medal; US Coast Guard 1990- Norwegian Society of Photobiology and Photomedicine, 1976- American Society of Limnology and Oceanography (ASLO)

Cruises and field work: Trondheimsfjord, Gulf of California, Norwegian Coastal Current, Lindåspollene (Norway), Scotia Sea (Southern Ocean), North Sea, Litoralstation Sylt (Germany), Antarctic circumnavigation cruise, Barents Sea, section from Capte-Town to Dronning ;aud Land (Antarctica)

Teaching and tutoring:

Since 1971, lectures on algal systematics (seaweeds), physical oceanography for undergraduates, courses in methods and phytoplankton taxonomy, lectures on marine ecology and biological oceanography (master level); 2 summer courses (Nordic Council and EU); tutoring of 22 master students and 8 Ph.D students; 2 summer courses (Nordic Council and EU)

Grants:

Norwegian Research Council (18), EU (7); New in 2010: HYDRALAB (EU). Research infrastructure, initiator in 1998, manager 1998-2004; activity still running, now under HYDRALAB

Administrative (partial):

2004-06 Norwegian Research Council, Section of Environment, member2004 IPCC, Chapter on Polar Climate, referee2001-04 Arctic Systems Inventory Analysis (ACIA), contributing author2000-04 Scientific advisory board, Inst. Meereskunde, Univ. Kiel 2000-04 Arctic Climate Impact Assessment (ACIA) marine, 2000-20042002-09 VISTA programme (Statoil/Academy of Science and Letters, Oslo), steering

board 2002 Working group on overgrazing of kelp forests (Ministry of Fisheries), chairman1999-2003 Marine Science Board, European Science Foundation 1998 Conference on Marine Environment in North West of Russia (MAREN), St.

Petersburg, Russia, deputy chairman of Science Committee; NTNU delegate in 1999 and 2000:

1996-2004 Scientific Committee of Antarctic Research (SCAR) Working Group on Biology

1995 1st Nordic Marine Sciences Meeting, Gothenburg. Initiator and chairman of planning committee Gothenburg, March 1995; 2nd Nordic Sciences Meeting, Hirtshals, 1999; planning committee

1994-2000 Joint Global Ocean Flux Study, Scientific Steering Committee, executive member 1995-2000; Task Team on Photosynthetic Measurements 1993-1999, chairman

1994 International Committee for Evaluation of Finnish Antarctic Research, chairman1993-2003 Strategic Programme for Marine Science and Technology (MODTEQ, at NTNU,

funded by NFR); chairman1993-2001 University Studies at Svalbard (UNIS), science committee; chairman of the

Biology Committee 1998-20011991-92 International Committee for Evaluation of Swedish Polar Research (Forsknings-

rådsnämden, Stockholm) 1989-92 Research Program for North-Norwegian Coastal Ecology (Mare Nor) 1988-90 Research programmes Mare Nor, Causes of the Occurrence of Harmful

Phytoplankton Blooms, and Marine Pollution", member of planning committees 1987- Albert Einstein prize (Consejo Cultural Mundial, Mexico), Kyoto Prize, Japan

Prize, Water Prize (Stockholm), Volvo Prize, nominator1987-99 The Museum of Natural History and Archaeology, NTNU, member of the

executive board 1987-94 Scientific Committee for Antarctic Research (SCAR), Group of Specialists on the

Ecology of the Southern Ocean, member 1985-89 APPRISE (field programme in Auke Bay, Alaska; Univ. Alaska, Oceanic Inst.

Hawaii), science advisory group1984-2010: Editorial board, J. Plankton Research, European J. of Phycology (1993-) 1984 Norwegian Antarctic Expedition of 1985-1986, member of steering committee

1983-1991 Pro Mare (Norwegian Research Programme for Marine Arctic Ecology, chairman

1982 Writing of the comprehensive plan for Pro Mare (Norwegian Research Programme for Marine Arctic Ecology), with Bjørn Gulliksen,

1978-83 Working group on biological decomposition and effects on micro-organisms, FOH (Research Programme on Oil Pollution), chairman

1976-1980 Steering committee, Norwegian Coastal Current Programme (NFR)1974-85 Budget and Planning Committee, the Museum, member 1974-1985; chairman

1979-19851971-85 Norwegian Oceanographic Committee1971-81 Central Budget and Planning Committee of the University of Trondheim,

chairman 1980-19811971-72 Plankton committee, ICES

Consultant/committee member for promotions and appointments (tenure) at research institutions and universities: Norway (16), USA (16), Germany (2), Sweden (1) Opponent, Ph.D (15; Norway, Sweden, Germany, France)Numerous evaluations of research proposals to research proposals: Norway, USA, EuropeConvener of 3 symposia (University of Alaska, Pro Mare, Trondheim, JGOFS Photosynthesis

Task Team, Svalbard)

Publications: (citations until 2010: H/I Index 26)

Publications are listed in three categories: refereed papers and review articles (series A), non-refereed and popularized science (series B), and books (papers) I have edited (series C).

Science Citation Index (1968-2010): 2900; H/I Index 26

Short version, chronological (2005-2009:

Sakshaug, E. and Johnsen, G. 2005. Absorption, fluorescence excitation and photoacclimation. Pp. 685-714 in Subba Rao, D.D. (ed) Algal cultures – Analogues of blooms and applications. Science Publishers, Plymouth U.K., 971 p (2 vols)

Chauton, M. S, Johnsen, G., Sakshaug, E. 2005. Nuclear Magnetic Resonance spectroscopy of marine microalgae: Metabolic profiling and species discrimination from High-resolution Magic Angle Spinning NMR analysis of whole-cell samples. Ph.D thesis, 113 p, NTNU Trondheim

Sakshaug, Egil; Johnsen, Geir. 2005. Algal cultures, Analogues of Blooms and Applications. Multi-Science Publishing (ISBN 1-57808-393-1), 27 s.

Carmack, E., Barber, D., Christensen, J., Macdonald, R., Rudels, B., Sakshaug, E. 2006.Climate variability and physical forcing of the food webs and the carbon budget on panarctic shelves. Prog Oceanogr; 71: 145-181.

Olsen, L.M., Öztürk, M., Sakshaug, Egi., Johnsen G. 2006. Photosynthesis-induced phosphate precipitation in seawater: ecological implications for phytoplankton. Marine Ecology Progress Series 2006 ;Volum 319:103-110.

Hancke, K., Johnsen, G., Sakshaug, E. 2007. Bio-optical properties and spectral attenuation of photosynthetically available radiation: relation to primary productivity and

phytoplankton composition. In Proc.Arctic Frontiers: balancing human use and ecosystem protection. University of Tromsø

Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2007. Rate of O2 production from PAM fluorescence: Testing three bio-optical approaches against measured O2 production rate. AquaFluo Conference, Tromsø, 28 May-June 2007

Johnsen, G., Sakshaug, E 2007. Application of in vivo fluorescence excitation spectra from different pigment-groups of phytoplankton. AquaFluo meeting; 2007-05-31,NTNU

Johnsen, G., Sakshaug, E. 2007. Biooptical characteristics of PSII and PSI in 33 species (13 pigment groups) of marine phytoplankton, and the relevance for pulse-amplitude-modulated and fast-repetition-rate fluorometry. J. Phycol. 43: 1236-1251

Johnsen, G., Volent, Z., Sigernes, F., Berge, J., Hovland, E.K., Sakshaug, E. 2007Local hyperspectral imaging of ocean colour from tethered balloons in the Arctic. Proc. Arctic Frontiers Meeting: Balancing human use and ecosystem protection. : University of Tromsø

Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2008. Rate of O2 production derived from pulse-amplitude-modulated fluorescence: Testing three biooptical approaches against measured O2-production rate. J. Phycol. 44: 803-813.

Fox, C., Harris, R., Sundby, S., Achterberg, E., Allen, J.I., Allen, J., Baker, A, Brussaard, C.P.D; Buckley, P; Cook, E.J; Dye, S.R.; Edwards, M; Fernand, L; Kershaw, P; Metcalfe, J; Osterhus, S; Potter, T; Sakshaug, Egil; Speirs, D.; Stenevik, E; John, M.S.; Thingstad, F., Wilson, B. 2009. Transregional linkages in the north-eastern Atlantic – an ‘end-to-end’ analysis of pelagic ecosystems. Oceanogr. Mar. Biol. Ann. Rev. 47:1-75.

Sakshaug, E. 2009. Egil Sakshaugs Saga. In: Å forstå livet. Institutt for biologi NTNU - 50 år. De biologiske fagenes historie i Trondheim og ved NTNU. Trondheim: Institutt for biologi, NTNU 2009 ISBN 82-471-6062-5. p. 88-101.

Sakshaug, E., Kovacs, K., Johnsen, G. (eds) 2009. Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0, 587 p.

Sakshaug, E., Johnsen, G., Kristiansen, S., von Quillfeldt, C., Rey, F., Slagstad, D., Thingstad, F. 2009. Phytoplankton and primary production. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 167-208 

Sakshaug, E., Johnsen, G., Volent, Z. 2009. Light. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 117-138

Johnsen, G., Volent, Z., Sakshaug, E., Sigernes, F., Pettersson, Lasse H. 2009. Remote sensing in the Barents Sea. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 139-166

Sakshaug, E., Kovacs, K.M. 2009. Introduction. I: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0.p. 9-32  Sakshaug, E., Rey, F., Ardelan, M.V. 2009. Seawater, its constituents and chemistry. In:

Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. s. 83-116  

von Quillfeldt, C., Hegseth, E.N., Johnsen, G. Sakshaug, E. Syvertsen, E.E. 2009. Ice algae.In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. s. 285-302

FULL LIST of LITERATURE

A. Refereed Papers and Review Articles

1A. Sakshaug, E. 1970. Quantitative phytoplankton investigations in near-shore water masses. K. Norske Vidensk. Selsk. Skr. 1970 (3), 8pp.2A. Jensen, A. and Sakshaug, E. 1970. Producer-consumer relationships in the sea. I. Prelimi-

nary studies on phytoplankton density and Mytilus pigmentation. J. Exp. Mar. Biol. Ecol. 5, 180-186.

3A. Jensen, A. Sakshaug, E. 1970. Producer-consumer relationships in the sea. I. Correlation be-tween Mytilus pigmentation and the density and composition of phytoplankton populations in inshore waters. J. Exp. Mar. Biol. Ecol. 5, 246-253.

4A. Sakshaug, E. and Jensen, A. 1971. Gonyaulax tamarensis and paralytic mussel toxicity in Trondheimsfjorden, 1963-1969. Kgl. Norske Vidensk. Selsk. Skr. 1972 (15), 15 pp.

5A. Sakshaug, E., 1972. Phytoplankton investigations in Trondheimsfjord, 1963-1966. K. Nor-ske Vidensk. Selsk. Skr. 1972 (1), 56 pp.

6A. Haug, A., Myklestad, S. and Sakshaug, E. 1973. Studies on the phytoplankton ecology of the Trondheimsfjord. I. The chemical composition of phytoplankton populations. J. Exp. Mar. Biol. Ecol. 11, 15-26.

7A. Jensen, A. and Sakshaug, E. 1973. Studies on the phytoplankton ecology of the Trond-heimsfjord. II. Chloroplast pigments in relation to abundance and physiological state of the phytoplankton. J. Exp. Mar. Biol. Ecol. 11, 137-155.

8A. Sakshaug, E. and Myklestad, S. 1973: Studies on the phytoplankton ecology of the Trondh-eimsfjord. III. Dynamics of phytoplankton blooms in relation to environmental factors, bioassay experiments and parameters for the physiological state of the populations. J. Exp. Mar. Biol. Ecol. 11, 157-188.

9A. Sakshaug, E., Myklestad, S., Krogh, T. and Westin, G. 1973: Production of protein and carbohydrate in the dinoflagellate Amphidinium carteri. Some preliminary results. Norw. J. Bot. 20, 211-218.

10A. Sakshaug, E. 1976. Dynamics of phytoplankton blooms in Norwegian fjords and coastal waters. Pp. 139-143 in Skreslet, S., Leinebø, R. and Sakshaug, E. (eds), Freshwater on the sea. Norwegian Association of Oceanographers, Oslo.

11A. Sakshaug, E. 1977. Limiting nutrients and maximum growth rates for diatoms in Narra- gansett Bay. J. Exp. Mar. Biol. Ecol. 28, 109-123.

12A. Sakshaug, E. and Holm-Hansen, O. 1977. Chemical composition of Skeletonema costatum (Grev.) Cleve and Pavlova lutheri (Droop)Green as a function of nitrate-, phosphate-, and iron-limited growth. J. Exp. Mar. Biol. Ecol. 29, 1-34.

13A. Sakshaug, E. 1978. The influence of environmental factors on the chemical composition of cultivated and natural populations of marine phytoplankton. Dr. philos. thesis, University of Trondheim, 83 pp.

14A. Sakshaug, E. and Jensen, A. 1978: The use of cage cultures in studies of the biochemistry and ecology of marine phytoplankton. Oceanogr. Mar. Biol. Ann. Rev. 16, 81-106.

15A. Sakshaug, E. 1980: Problems in the methodology of studying phytoplankton. Pp. 57-91 in Morris, I. (ed.): The physiological ecology of phytoplankton. Blackwell Scientific Publications, 625 pp.

16A. Bell, W.H. and Sakshaug, E. 1980: Bacterial utilization of algal extracellular products. II. A kinetics study on natural populations. Limnol. Oceanogr. 25, 1021-1033.

17A. Sakshaug, E., Myklestad, S., Andresen, K., Hegseth, E.N. and Jørgensen, L. 1981 Phytoplankton off the Møre Coast in 1975-1976: distribution, species composition, chemical composition and conditions for growth. Pp. 681-711 (vol. 2) in Sætre, R. and

Mork, M. (eds), The Norwegian Coastal Current, University of Bergen, 795 pp (vols 1 & 2).

18A. Sakshaug, E., Andresen, K., Myklestad, S. and Olsen, Y. 1983: Nutrient status of phytoplankton communities in Norwegian waters (marine, brackish, and fresh) as revealed by their chemical composition. J. Plankton Res. 5, 175-196.

19A. Hegseth, E. and Sakshaug, E. 1983. Seasonal variation in light- and temperature-dependent growth of marine planktonic diatoms in in situ dialysis cultures in the Trondheimsfjord, Norway (63N). J. Exp. Mar. Biol. Ecol. 67, 199-220.

20A. Myklestad, S. and Sakshaug, E. 1983: Alkaline phosphatase activity of Skeletonema costatum populations in the Trondheimsfjord. J. Plankton Res. 5, 557-564.

21A. Sakshaug, E., Granéli, E., Elbrächter, M. and Kayser, H. 1984. Chemical composition and alkaline phosphatase activity of nutrient-saturated and P-deficient cells of four marine dinoflagellates. J. Exp. Mar. Biol. Ecol. 84, 241-254.

22A. Sakshaug, E. and Holm-Hansen, O. 1984. Factors governing pelagic production in polar oceans. Pp. 1-18 in Holm-Hansen, O., Bolis, L. and Gilles, R. (eds), Marine phytoplankton and productivity. Lecture notes on coastal and estuarine studies, vol. 8. Springer Verlag.

23A. Hewes, C.D., Holm-Hansen, O. and Sakshaug, E. 1985. Alternate carbon pathways at lower trophic levels in the Antarctic food web. Pp. 277-283 in Siegfried, W.R., Condy, P.R. and Laws, R.M. (eds), Antarctic nutrient cycles and food webs, Springer Verlag, 700 pp.

24A. Sakshaug, E. and Olsen, Y. 1986: Nutrient status of phytoplankton blooms in Norwegian waters and algal strategies for nutrient competition. Can. J. Fish. Aq. Sci. 43, 389-396.

25A. Sakshaug, E. and Holm-Hansen, O. 1986. Photoadaptation in Antarctic phytoplankton: variations in growth rate, chemical composition and P versus I curves. J. Plankton Res. 8, 459-473.

26A. Sakshaug, E. and Andresen, K. 1986. Effect of light regime upon growth rate and chemical composition of a clone of Skeletonema costatum from the Trondheimsfjord, Norway. J. Plankton Res. 8, 619-638.

27A. Sakshaug, E., Demers, S. and Yentsch, C.M. 1987. Thalassiosira oceanica and T. seudonana: two different photoadaptational responses. Mar. Ecol. Prog. Ser. 41, 275-282.

28A. Sakshaug, E., Andresen, K. and Kiefer, D.A. 1989. A steady-state model of growth and light absorption in the marine plankton diatom Skeletonema costatum. Limnol. Oceanogr. 34, 198-205.

29A. Sakshaug, E. and Skjoldal, H.R. 1989. Life at the ice edge. Ambio 18, 60-67.30A. Sakshaug, E. 1989. The physiological ecology of polar phytoplankton. Pp. 61-89 in Rey, L.

and Alexander, V. (eds): Marine living systems of the far North (6th Conf. Comité Arctique Int., Fairbanks, Alaska), E.J. Brill, Leiden, The Netherlands. 637 pp.

31A. Hewes, C.D., Sakshaug, E.H., Reid, F.M.H. and Holm-Hansen, O. 1990. Microbial autotrophic and heterotrophic eucaryotes in Antarctic waters: Relationships between biomass and chlorophyll, adenosine triphosphate and particulate organic carbon. Mar. Ecol. Prog. Ser. 63, 27-35.

32A. Thingstad, T.F. and Sakshaug, E. 1990. Control of phytoplankton growth in nutrient recycling ecosystems. Theory and terminology. Mar. Ecol. Prog. Ser. 63, 261-272.

33A. Gilstad, M. and Sakshaug, E. 1990. Growth rates of ten diatom species from the Barents Sea at different irradiances and day lengths. Mar. Ecol. Prog. Ser. 64, 169-173.

34A. Sakshaug, E. 1990. Arctic planktonic ecosystems. Pp. 47-51 in Priddle, J. and Medlin, L. (eds) Polar diatoms - a manual, British Antarctic Survey, Cambridge.

35A. Smith, W.O. and Sakshaug, E. 1990. Primary production in Polar Oceans. Pp. 477-525 in Smith, W.O. (ed.) Polar Oceanography, part B. Academic Press, 760 pp (2 vols).

36A. Sakshaug, E., Johnsen, G., Andresen, K. and Vernet, M. 1991. Modeling of light-dependent algal photosynthesis and growth: Experiments with the Barents Sea diatoms Thalassiosira nordenskioeldii and Chaetoceros furcellatus. Deep-Sea Res. 38, 415-430.

37A. Sakshaug, E. 1991. Food webs and primary production in the Barents Sea. Proc. 12th NIPR Symp. Polar Biol. (Tokyo) 4, 1-8.

38A. Sakshaug, E. and Slagstad, D. 1991. Factors controlling the development of phyto- plankton blooms in the Antarctic Ocean - A mathematical model. Mar. Chem. 35, 259-272.

39A. Sakshaug, E. and Slagstad, D. 1991. Light and productivity of phytoplankton in polar marine ecosystems. Polar Res. 10, 69-85.

40A. Verity, P.G., Smayda, T.J. and Sakshaug, E. 1991. Photosynthesis, excretion, and growth rates of Phaeocystis colonies and solitary cells. Polar Res. 10, 117-128.

41A. Johnsen, G., Sakshaug, E. and Vernet, M. 1992. Pigment composition, spectral characterization and photosynthetic parameters in Chrysochromulina polylepis. Mar. Ecol. Prog. Ser. 83, 241-249.

42A. Sakshaug, E. Slagstad, D. 1992. Sea-ice and wind: Effects on primary production in the Barents Sea. Atmosphere-Ocean 30, 579-571.

43A. Gilstad, M., Johnsen, G. and Sakshaug, E. 1993. Diurnal- and daylength-dependent variations in photosynthetic parameters, pigment composition and the respiration rate of Skeletonema costatum. J. Plank. Res. 15, 939-951.

44A. Sakshaug, E. 1993. The relationship between phytoplankton growth rate and production with emphasis on respiration and excretion. ICES Mar. Symp. Ser. 197, 63-68.

45A. Nielsen, M. and Sakshaug, E. 1993. Adaptation to spectrally different light regimes: Studies of the growth rate and Carbon to Chlorophyll a ratio of the marine diatom Skeletonema costatum. Limnol. Oceanogr. 38, 1576-1581.

46A. Johnsen, G. and Sakshaug, E. 1993. Bio-optical characteristics and photoadaptive responses in the toxic and bloom-forming dinoflagellates Gyrodinium aureolum, Gymnodinium galatheanum, and two strains of Prorocentrum minimum. J. Phycol. 29, 627-642.

47A. Cullen, J.J., Geider, R.J., Ishizaka, J., Kiefer, D.A., Marra, J., Sakshaug, E. and Raven, J.A. 1993. Toward a general model of phytoplankton growth for biogeochemical models. Pp 153-176 in Evans, G.T. and Fasham, M.J.R. (eds): Towards a model of ocean biogeochemical processes. NATO ASI Series I, vol. 10, Springer-Verlag, Berlin, Heidelberg, 350 pp.

48A. Sakshaug, E. 1993. Variability in photosynthetic parameters. Pp. 2-10 in Eilertsen, H.C. (ed.) Proc. Europto Series, SPIE 2048, 190 pp.

49A. Johnsen, G., Samseth, O., Granskog, L. and Sakshaug, E. 1993. In vivo absorption characteristics in 10 classes of phytoplankton: Taxonomic characteristics and responses to photoadaptation by means of discriminant and HPLC analysis. Mar. Ecol. Prog. Ser. 105, 149-157.

50A. Baumann, M.E.M., Lancelot, C., Brandini, F.P., Sakshaug, E. and John, D.M. 1994. The taxonomic identity of the cosmopolitan prymnesiophyte Phaeocystis: a morphological and ecophysiological approach. J. Mar. Systems 5, 5-22.

51A. Sakshaug, E, Bjørge, A., Gulliksen, B., Loeng, H. and Mehlum, F. 1994. Structure, biomass distribution, and energetics of the pelagic ecosystem in the Barents Sea: A synopsis. Polar Biol. 14, 405-411.

52A. Dalløkken, R., Sandvik, R. and Sakshaug, E. 1994. Variations in bio-optical properties in the Greenland/Iceland/Norwegian Seas. Ocean Optics 12. SPIE 2258, 266-276.

53A. Dalløkken, R., Sandvik, R. and Sakshaug, E. 1995. Seasonal variations in the vertical light attenuation coefficient in the Greenland Sea: Effect of phytoplankton light absorption. Pp. 33-43 in Skjoldal, H.R., Hopkins, C.C.E., Erikstad, K.E. and Leinaas, H.P. (eds). Ecology of fjords and coastal waters. Elsevier Science, Amsterdam.

54A. Sakshaug, E., Rey, F. and Slagstad, D. 1995. Wind forcing of marine primary productivity in the atmospheric low-pressure belts. Pp. 15-25 in Skjoldal, H.R., Hopkins, C.C.E., Erikstad, K.E. and Leinaas, H.P. (eds). Ecology of fjords and coastal waters. Elsevier Science, Amsterdam.

55A. Johnsen, G. and Sakshaug, E. 1995. Light harvesting in bloom-forming marine phytoplankton: species-specificity and photoadaptation. Scientia Marina 60 (Suppl.), 47-56.

56A. Vernet, M., Mitchell, B.G., Sakshaug, E., Johnsen, G., Iturriaga, R. and Wassmann, P. 1996. Evidence for a novel pigment with in vivo absorption maximum at 708 nm associated with Phaeocystis cf. pouchetii blooms. Mar. Ecol. Prog. Ser. 133, 253-262.

57A. Sakshaug, E. 1996. Biomass, productivity distributions and their variability in the Barents Sea. ICES mar. Sci. Symp. 54, 341-350.

58A. Grzymski, J., Johnsen, G. and Sakshaug, E. 1997. The significance of intracellular self-shading on the bio-optical properties of brown, red and green macroalgal species. J. Phycol. 33, 408-414.

59A. Sakshaug, E., Bricaud, A., Dandonneau, Y., Falkowski, P.G., Kiefer, D.A., Legendre, L., Morel, A., Parslow, J. and Takahashi, M. 1997. Parameters of photosynthesis: definitions, theory and interpretation of results. J. Plank. Res. 19, 1637-1670.

60A. Johnsen, G., Volent, Z., Tangen, K. and Sakshaug, E. 1997. Time series of harmful and benign phytoplankton blooms in northwest European waters using the Seawatch buoy system. Pp. 115-143 in Kahru, M. and Brown, C.W. (eds). Monitoring algal blooms: New techniques for detecting large-scale environmental change. Landes Bioscience, New York.

61A. Johnsen, G., Samset, O. and Sakshaug, E., 1997. Bio-optical classification of potential harmful chlorophyll c3-containing prymnesiophyte blooms in Scandinavian waters.

62A. Sakshaug, E. 2000. Wind forcing of marine primary production in the northern belt of westerlies. Pp. 19-36 in Hanson, R., Ducklow, H. and Field. J.G. (eds). The changing ocean carbon cycle. IGBP Series 5, Cambridge Univeristy Press, Cambridge, U.K., 514 p.

63A. Sakshaug, E. and Walsh, J. 2000. Marine biology: Biomass, productivity distributions and their variability in the Barents and Bering Seas. Pp. 161-195 in Nuttall, M. and Callaghan, T.V. The Arctic: a guide to research in the natural and social sciences. Harwood, London.

64A. Johnsen, G. and Sakshaug, E. 2000. Monitoring of harmful algal blooms along the Norwegian coast. S. Afr. J. Mar. Sci. 22, 309-321.

65A. Aas E, Højerslev NK, Sandvik R, Sakshaug E 2002. Spectral properties and UV-attenuation in Arctic marine waters. Pp. 23-56 in Hessen, D (ed) Radiation and Arctic ecosystems. Springer, Heidelberg-Berlin.

66A. Özturk, M., Steinnes, E. And Sakshaug, E. 2002. Iron speciation in the Trondheim Fjord from the perspective of iron limitation for phytoplankton. Est. Coastal Shelf Sci 55, 197-212.

67A. Sakshaug, Egil (2003). Primary production in Arctic and subarctic waters. Pp 57-81 in Stein R, Macdonald R (eds) The organic arctic arbon cycle. Springer, Berlin-Heidelberg, 363 p.

68A. Macdonald, RW, Sakshaug, E and Stein, R. 2003: The Arctic Ocean: Modern Status

and climate change. Pp. 6-21 in Stein, R. and Macdonald RW (eds): The organic carbon cycle in the Arctic Ocean. Springer, 363 p. 70A.

69A. Öztürk, M., Vadstein, O. and Sakshaug, E. 2003. The effects of enhanced phytoplankton production on iron speciation and removal in mesocosm experiments in a landlocked basin of Hopavågen, Norway. Mar. Chem. 84, 3-17.

70A. Öztürk, M., Croot, P., Bertilsson, S., Abrahamsson, K., Karlson, B., Davis, R., Fransson, A. and Sakshaug, E. 2004. Iron enrichment and photoreduction of iron under UV and PAR in the presence of hydrocarboxylic acid: Implication for4 phytoplankton growth in the Southern Ocean. Deep-Sea Res. II 51, 2841-2856

71A. Sakshaug, E. and Johnsen, G. 2005. Absorption, fluorescence excitation and photoacclimation. Pp. 685-714 in Subba Rao, D.D. (ed) Algal cultures – Analogues of blooms and applications. Science Publishers, Plymouth U.K., 971 p (2 vols)

72A. Chauton, M. S, Johnsen, G., Sakshaug, E. 2005. Nuclear Magnetic Resonance spectroscopy of marine microalgae: Metabolic profiling and species discrimination from High-resolution Magic Angle Spinning NMR analysis of whole-cell samples. Ph.D thesis, 113 p, NTNU Trondheim

73A. Sakshaug, Egil; Johnsen, Geir. 2005. Algal cultures, Analogues of Blooms and Applications. Multi-Science Publishing (ISBN 1-57808-393-1), 27 s.

74A Carmack, E., Barber, D., Christensen, J., Macdonald, R., Rudels, B., Sakshaug, E. 2006.Climate variability and physical forcing of the food webs and the carbon budget on panarctic shelves. Prog Oceanogr; 71: 145-181.

75A. Olsen, L.M., Öztürk, M., Sakshaug, Egi., Johnsen G. 2006. Photosynthesis-induced phosphate precipitation in seawater: ecological implications for phytoplankton. Marine Ecology Progress Series 2006 ;Volum 319:103-110.

76A. Hancke, K., Johnsen, G., Sakshaug, E. 2007. Bio-optical properties and spectral attenuation of photosynthetically available radiation: relation to primary productivity and phytoplankton composition. In Proc.Arctic Frontiers: balancing human use and ecosystem protection. University of Tromsø

77A. Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2007. Rate of O2 production from PAM fluorescence: Testing three bio-optical approaches against measured O2 production rate. AquaFluo Conference, Tromsø, 28 May-June 2007

78A. Johnsen, G., Sakshaug, E 2007. Application of in vivo fluorescence excitation spectra from different pigment-groups of phytoplankton. AquaFluo meeting; 2007-05-31,NTNU

79A. Johnsen, G., Sakshaug, E. 2007. Biooptical characteristics of PSII and PSI in 33 species (13 pigment groups) of marine phytoplankton, and the relevance for pulse-amplitude-modulated and fast-repetition-rate fluorometry. J. Phycol. 43: 1236-1251

80A. Johnsen, G., Volent, Z., Sigernes, F., Berge, J., Hovland, E.K., Sakshaug, E. 2007Local hyperspectral imaging of ocean colour from tethered balloons in the Arctic. Proc. Arctic Frontiers Meeting: Balancing human use and ecosystem protection. : University of Tromsø

81A. Hancke, T.B., Hancke, K., Johnsen, G., Sakshaug, E. 2008. Rate of O2 production derived from pulse-amplitude-modulated fluorescence: Testing three biooptical approaches against measured O2-production rate. J. Phycol. 44: 803-813.

82A. Fox, C., Harris, R., Sundby, S., Achterberg, E., Allen, J.I., Allen, J., Baker, A, Brussaard, C.P.D; Buckley, P; Cook, E.J; Dye, S.R.; Edwards, M; Fernand, L; Kershaw, P; Metcalfe, J; Osterhus, S; Potter, T; Sakshaug, Egil; Speirs, D.; Stenevik, E; John, M.S.; Thingstad, F., Wilson, B. 2009. Transregional linkages in the north-eastern Atlantic – an ‘end-to-end’ analysis of pelagic ecosystems. Oceanogr. Mar. Biol. Ann. Rev. 47:1-75.

83A. Sakshaug, E. 2009. Egil Sakshaugs Saga. In: Å forstå livet. Institutt for biologi NTNU - 50 år. De biologiske fagenes historie i Trondheim og ved NTNU. Trondheim: Institutt for biologi, NTNU 2009 ISBN 82-471-6062-5. p. 88-101.

84A. Sakshaug, E., Kovacs, K., Johnsen, G. (eds) 2009. Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0, 587 p.

85A. Sakshaug, E., Johnsen, G., Kristiansen, S., von Quillfeldt, C., Rey, F., Slagstad, D., Thingstad, F. 2009. Phytoplankton and primary production. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 167-208 

86A. Sakshaug, E., Johnsen, G., Volent, Z. 2009. Light. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 117-138

87A. Johnsen, G., Volent, Z., Sakshaug, E., Sigernes, F., Pettersson, Lasse H. 2009. Remote sensing in the Barents Sea. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. p. 139-166

88A. Sakshaug, E., Kovacs, K.M. 2009. Introduction. I: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0.p. 9-32  89A. Sakshaug, E., Rey, F., Ardelan, M.V. 2009. Seawater, its constituents and chemistry. In:

Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. s. 83-116  

90A. von Quillfeldt, C., Hegseth, E.N., Johnsen, G. Sakshaug, E. Syvertsen, E.E. 2009. Icealgae. In: Ecosystem Barents Sea. Tapir Academic Publishers, Trondheim. ISBN 978-82-519-2461-0. s. 285-302

B. Popular and non-refereed publications

1B. Sakshaug, E. 1969. Planteplankton (Phytoplankton). Pp. 22-39 in Strømgren, T. (ed.): The Borgenfjord Investigations, Preliminary report 1968. The Royal Norwegian Society of Sciences and Letters, The Museum, Trondheim (in Norwegian).

2B. Sakshaug, E. and Jensen, A. 1971. Gonyaulax tamarensis og paralytisk muslinggift i Trondheimsfjorden, 1970-1971 (Gonyaulax tamarensis and paralytic mussel toxicity in the Trondheimsfjord, 1970-1971). Pp. 14-20 in Strømgren, T. (ed.): The Borgenfjord Investigations, Preliminary report 1970. The Royal Norwegian Society of Sciences and Letters, The Museum, Trondheim (in Norwegian).

3B. Sakshaug, E., Jensen, A. and Prakash, A. 1971. Gonyaulax tamarensis - the causative organism of mussel toxicity in Trondheimsfjord. ICES C.M. 1971, Helsinki. Report no. L:14. 13 pp.

4B. Sakshaug, E. and Haug, A. 1972. Forurensinger i marint miljø (Pollution in the marine environment). Pp. 125-142 in Carlsen, K., Celius, H., Halmø, T. and Nestaas, I. (eds) Miljøkunnskap (Environmental Science). Tapir, Trondheim. 308 pp (in Norwegian).

5B. Sakshaug, E., Haug, A., Jensen, A. and Myklestad, S. 1972. Phytoplankton ecology of the Trondheimsfjord. ICES, C.M. 1972, Copenhagen. Report no. L:13, 20 pp.

6B. Sakshaug, E. and Brun, P. 1974. Plantenæringsstoffer og planteplankton i Trond-heimsfjorden (Plant nutrients and phytoplankton in the Trondheimsfjord). Vann 1974 (3) 1-12.

7B. Sakshaug, E. 1975. Noen tanker om modeller for akvatisk produktivitet (Some thoughts on models for aquatic productivity). Pp. 195-211 in Rept 10th Nordic Symp. Water Res., Værløse, Denmark, May 1974. Nordforsk Publication 1975 (1). 559 pp.

8B. Sakshaug, E. 1977. Plantenæringsstoffer og klorofyll a (Plant nutrients and chlorophyll a). Pp. 134-151 in Lande, E. (ed.): Resipientundersøkelsen av Trondheimsfjorden, 1972-75 (The Trondheimsfjord Investigations in 1972-75). The Royal Norwegian Society of Sciences and Letters, The Museum, Rept Zool. Series 1977 (11), in Norwegian.

9B. Sakshaug, E. 1978: Trondhjem Biologiske Stasjon - en god historie (Trondhjem Biological Station - a good story). Akvarie-Tanken, no 3. The Royal Norwegian Society of Sciences and Letters, The Museum, 16 pp (in Norwegian).

10B. Sakshaug, E. and Andersen, B. 1980: Norsk oseanografi - fortid og fremtid (Norwegian Oceanography - past and future), Forskningsnytt 1980 (1), 26-28 (in Norwegian).

11B. Sakshaug, E. 1980. Organiske partikler i sjøvann (Organic particles in seawater). Naturen (2), 29-40 (in Norwegian).

12B. Paden, C.A., Hewes, C.D., Neori, A., Holm-Hansen, O., Weaver, E., Kiefer, D.A. and Sakshaug, E. 1981. Phytoplankton studies in the Scotia Sea. Ant. J. U.S. 16(5), 163.

13B. Sakshaug, E. 1982. Vassdragsregulering og liv i hav (Regulation of rivers and the effect on life in the sea). Trøndersk Natur 1982 (1), 8-12 (in Norwegian).

14B. Sakshaug, E., 1982. Krill - mat i mengder i Sørishavet (Krill - food in abundance in the Southern Ocean). Syn og Segn, 88, 482-489 (in Norwegian).

15B. Hewes, C.D., Holm-Hansen, O. and Sakshaug, E. 1983. Nanoplankton and microplankton studies during the circumnavigation cruise. Ant. J. U.S. 1983, 169-171.

16B. Legendre, L., Collos, Y., Elbrächter, M., Fasham, M.J.R., Gieskes, W.W.C., Herbland, A., Holligan, P.M., Margalef, R., Perry, M.J., Platt, T., Sakshaug, E. and Smith, D.F. 1984. Studies on marine autotrophs: Recommendations for the 1980s. Pp. 629-639 in Fasham, M.J.R. (ed.), Flows of energy and materials in marine ecosystems. Plenum Press.

17B. Sakshaug, E. 1984. Going north - the need for ecological studies in the Barents and Spitsbergen areas. Annual meeting, Norw. Petroleum Society, Bergen, May 1984.

18B. Sakshaug, E. 1986. Pro Mare - The Norwegian research program for marine arctic ecology. Scandinavian Review 74, 44-47.

19B. Sakshaug, E. 1987. Fosfor og nitrogen som begrensende faktorer for algevekst Phospho- rus and nitrogen as limiting factors for algal growth). Pp. 247-255 in: Rept 22nd Nordic Symp. Water Res., Laugarvatn, Iceland, May 1986. Nordforsk Publication, Nature Conservation Series, No. 1987(1), Helsinki.

20B. Wassmann, P. and Sakshaug, E. 1987. Barentshavets økologiske grunntrekk (Short review of the ecology of the Barents Sea), Naturen 1987(4), 151-158 (in Norwegian).

21B. Sakshaug, E. 1988. Pro Mare 1984-1989. UnitNytt 18 (2), 8-10 (in Norwegian).22B. Sakshaug, E. and Tangen, K. 1988. Tilførsel av plantenæringsstoffer: potensielle effekter

på algevekst (Plant nutrient supply: Potential effects on algal growth). Resipient- undersøkelse for Trondheim, Delrapport 9 (Investigations of the Trondheim basin as a recipient for sewage, Report no. 9). Oceanor Trondheim, (in Norwegian) 40 pp.

23B. Sakshaug, E. 1989. Spleiselag i Arktis. UnitNytt 19, 8-13.24B. Sakshaug, E. 1990. Mot en ny istid? (Towards a new glacial age?). UnitNytt 20 (4), 3-4 and

18 (in Norwegian).25B. Sakshaug, E. 1990. Marine ecology and climate. Compendium for Norsk Hydro A/S, 24 pp

(Norwegian and English edition).26B. Sakshaug, E. and Tangen, K. 1990. The Chrysochromulina bloom in Skagerrak and

Kattegat summer 1988. OECD Environmental Report, 1990.27B. Sakshaug, E., Johnsen, G., Samseth, O and Volent, Z. 1991. Identification of phytoplank-

ton blooms by means of remote sensing. Environment Northern Seas Meeting, Stavanger 28 August 1991, 91-100.

28B. Sakshaug, E. 1991. Den biologiske karbonpumpen: Primærproduksjon i havet (The biological carbon pump: Primary production in the sea). Kjemi 52, 27-32 (in Norwegian; English summary).

29B. Sakshaug, E. 1992. The Barents Sea: The physical-biological connection. ("Anton Bruun Lecture" of IOC/UNESCO). Pp 23-26 In Bruun Memorial Lectures, 1991: Modelling and prediction in marine science. IOC/UNESCO, Paris, 33 p.

30B. Sakshaug, E. 1992. Brukes "føre var"-prinsippet på en uforsvarlig måte? Biolog, 1992 (1), 18-26.

31B. Sakshaug, E., Bjørge, A., Gulliksen, B., Loeng, H. and Mehlum, F. 1992. Innledning (Introduction). Pp. 11-19 in Økosystem Barentshavet (7C; in Norwegian).

32B. Sakshaug, E., Bjørge, A., Gulliksen, B., Loeng, H. and Mehlum, F. 1992. Epilog (Epilogue). Pp. 275-285 in Økosystem Barentshavet (7C; in Norwegian).

33B. Sakshaug, E., Volent, Z., Kögeler, J. and Slagstad, D. 1992. Lys i polarstrøk (Light in polar areas). Pp. 65-78 in Økosystem Barentshavet (7C; in Norwegian).

34B. Sakshaug, E., Kristiansen, S., Syvertsen, E., Eilertsen, H.-C., Gilstad, M., Hegseth, E.N., Hewes, C.D., Holm-Hansen, O., Johnsen, G., Mitchell, G.B., Rey, F., Slagstad, D., Støle-Hansen, K., Thingstad, T.F. and Vernet, M. 1992. Planteplankton (Phytoplankton). Pp. 79-108 in Økosystem Barentshavet (7C; in Norwegian).

35B. Rey, F., Sakshaug, E. and Myklestad, S. 1992. Sjøvannskjemi (Seawater chemistry). Pp. 54-63 in Økosystem Barentshavet (7C; in Norwegian).

36B. Slagstad, D., Sakshaug, E., Loeng, H., Tande, K and Ådlandsvik, B. 1992. Matematisk modellering (Mathematical modelling). Pp. 250-262 in Økosystem Barentshavet (7C; in Norwegian).

37B. Vinje, T., Gulliksen, B., Loeng, H., Hegseth, E.N., Sakshaug, E. and Syvertsen, E.S. 1992. Havis (Sea ice). Pp. 125-126 in Økosystem Barentshavet (7C; in Norwegian).

38B. Sakshaug, E. 1993. Discussant's report: primary production in the Antarctic pelagial - a view from the North. Proc. BIOMASS symposium, Bremerhaven, September 1990. p. 125-126.

39B. Sakshaug, E. 1995. A synopsis of the biomass distribution and energetics of the pelagic ecosystem of the Barents Sea. ICES.CM 1995/Mini:3, 11 p.

40B. Sakshaug, E., Olsen, Y. and Hoell, E. 1995. Maricult research programme: Fertilising marine ecosystems to harvest more. ICES.CM 1995/T:1, 16 p.

41B. Sakshaug, E. 1995. Weather and climate: effects on life in the sea. Kgl. Norske Vidensk. Selsk. Forh. 1995, 103-123.

42B. Sakshaug, E. 1995. Biomass and biological production in Arctic seas. Proc. EuroMab Conf., Kangerlussuaq, Greenland, September 1995 (submitted).

43B. Sakshaug, E. 1997. The Nordic region: cycles in climate and marine life. Pp. 40-51 in Andresen, K.H., Arlov, T.B., Hagen, D., Løset, S., Reinertsen, R.E. and Thomassen, J. (eds). Sustainable development in the Arctic. Consequences of industrial development in the European Arctic region. Centre for environment and development, Norwegian University of Science and Technology, Trondheim.

44B. Sakshaug, E. 1997. The nordic region: Cycles in climate and marine life. Forh. norske vidensk.Akad.

45B. Sakshaug, E. 1997. Istid, ikke nå igjen (Ice age, not now again). Universitsavisa, 21 August 1997.

46B. Sakshaug, E. 1998. El Nino er her igjen (El Nino is here again). Universitetsavisa, November 1998.

47B. Sakshaug, E. and Slagstad, D. 2000. Fjordens posisjon og solens gang )The location of the fjord and the solar track). P. 18 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

48B. Sakshaug, E. 2000. Hvorfor istider? (Why ice ages?). Pp. 28-29 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

49B. Sakshaug, E. 2000. Trondheimsfjorden og havet utenfor (The Trondheimsfjord and the sea outside). Pp. 38-41 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

50B. Sakshaug, E. 2000. Om salt, CO2 og organisk stoff i sjøen- Pp. 59-64 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

51B. Sakshaug, E. and Killingtveit, Å. 2000. Elvene: et stort bidrag (The rivers: a large contribution. Pp. 65-75.

52B. Sakshaug, E. and Olsen, Y. 2000. Litt om økologi (A little about ecology). Pp. 76-82 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

53B. Sakshaug, E. and Tangen, K. 2000. Lyset, næringssaltene og planteplanktonet (The light, the nutrients and the phytoplankton). Pp. 83-95 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

54B. Tangen, E. and Sakshaug, E. 2000. Skadelige alger (Harmful algae). Pp. 96-102 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

55B. Tokle, N. and Sakshaug, E. 2000. Dyreplanktonet (The zooplankton). Pp. 103-109 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

56B. Sakshaug, E. 2000. Matalger (Edible seaweeds). P. 164 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

57B. Sakshaug, E. 2000. Søl - sagaens alge (Palmaria palmata - the seaweed of the sagas). P. 167 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

58B. Sakshaug, E. and Sneli, J.-A. 2000. Trondhjem biologiske stasjon: en sprek hundreåring (Trondhjem Biological Station: a vigorous centenarian). Pp. 316-318 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

59B. Sakshaug, E. 2000. Units. Pp. 319-322 in Sakshaug, E. and J.-A. Sneli (eds) Trondheimsfjorden. Tapir, Trondheim, 336 p (in Norwegian).

60B. Sakshaug, E., Christie, H., Dale, T., Fosså, J.H., Fredriksen, S., Hedlund, N. og Sivertsen, K. 2002. Nedbeiting av tareskog i Norge. Fiskeridepartementet, Oslo, 47 p.

61B: Laws, E, Sakshaug, E., Babin, M., Dandonneau, Y., Falkowski, P., Geider, R., Legendre, L., Morel, A., Sondergaard, M., Takahashi, M. And Williams, P. leB. 2002. Photosynthesis and primary productivity in marine ecosystems. Practical aspects and application of techniques. Joint Global Ocean Flux Study, Rapport 36; Univ. Bergen, 88 p.

62B: Sakshaug, E. 2002. Primary Production and marine climate in the Nordic and Arctic Seas. IRISEN II (Integrated regional impact studies in the European North), Advanced Study Courses, Abisko, Sverige, 26. juli 2002, 10 p (manuscript).

63B: Sakshaug, E. 2002. Phytoplankton. Advanced Nordic courses: Sensitivity of the Westnordic Marine Ecosystem to past, present and future climate. Laugarvatn, Island, 19 august, 2 p (manuscript).

64B: Sakshaug, E. Phytoplankton. DKNVS Arctic Symposium, Kongsvoll, 23. august, 55 p (manuscript)

65B: Sakshaug, E. 2002. Primary production and marine climate in Nordic and Arctic Seas. Workshop, Havforskningsinstituttet, Thórshavn, Færøyane, 25. september, 9 p. (manuscript).

66B: Sakshaug E. and Walsh, J.J. 2002. Productivity, distributions and their variability in the Barents Sea and the Bering shelf: a summary. Univ. South Florida, St. Petersburg, Florida, 18. oktober, 9 p (manuscript).

67B: Sakshaug, E. 2002. Variability in primary production in Arctic Seas. Climate workshop, Nordic Arctic Research Programme, Sigulda, Latvia, 4. november, 35 p.

68B: Sakshaug E. 2002. Arctic primary production vs. climate variation. Dept Biology, University of California Santa Barbara, 12. november (set of overhead copies)

69B. Holm-Hansen, O.and Sakshaug, E. 2002. Polar Marine Phytoplankton. Pp. 2554-2565in Bitton, G. (ed.). Encyclopedia of Environmental Microbiology, vol. 5. John Wiley & Sons, New York.

70B Sakshaug, E. 2004. Primary production and marine climate in the arctic seas. Pp. 283-297 in Lange, M.A. and Pöszig, D. (eds) IRISEN II. Proc. Adv. Stud. Course, Abisko,

Sweden. Centre Environ. Res. Univ. Münster, Münster, 484 p.

Official reports:

1976: Norwegian Oceanography ─ Status and Perspectives (2C in the list of publications), the Norwegian Oceanographic Committee (NAVF)

1979: Norwegian Oceanography ─ Recent Development and Perspectives, the Norwegian Oceanographic Committee (NAVF)

1982 Comprehensive Program for the Norwegian Research Program for Marine Arctic Ecology (Pro Mare, published by NFFR and NAVF

C. Books/Editorial work

1C. Skreslet, S., Leinebø, R., Matthews, J.B.L. and Sakshaug, E. 1976. Freshwater on the sea. Proc. Symp. on the influence of fresh-water outflow on biological processes in fjords and coastal waters, Geilo, Norway, April 1974. Norwegian Association of Oceanographers, Oslo. 246 p.

2C. Sakshaug, E., Wedege, N.P. and Dahl, F.E. 1976. Norsk oseanografi - status og per- spektiver (Norwegian Oceanography - status and perspectives). Norwegian Oceano-graphic Committee, Norwegian Research Council for Science and the Humanities, Oslo. 248 p (with English summary).

3C. Sakshaug, E. 1979: Norsk oseanografi - nyere utvikling og perspektiver (Norwegian Oceanography - recent development and perspectives). Norwegian Oceanographic Committee, Norwegian Research Council for Science and the Humanities, Oslo. 100 p.

4C. Sakshaug, E. and Gulliksen, B. 1983. Marinøkologisk forskningsprogram i Arktis: rammeprogram (A comprehensive plan for the Norwegian Research Program for Marine Arctic Ecology, Pro Mare). The Norwegian Fisheries Research Council, the Norwegian Research Council for Science and the Humanities, Trondheim/Oslo. 55 p.

5C. Sakshaug, E. and Hegseth, E.N. 1984. Sluttrapport fra faggruppe IV: Biologisk nedbrytning og effekter på mikroorganismer. II. Studier med planktonalger (Final report from scientific committee IV: Biological degradation and effects on microorganisms. II. Studies on planktonic algae. Pp. 88-111 in Lange, R. (ed.): Faggruppenes sluttrapporter (Final reports from the scientific committees). Forskningsprogram om Havforurensninger (The Norwegian marine pollution research and monitoring programme (FOH). Miljøverndepartementet (The Ministry of the Environment), Oslo (in Norwegian).

6C. Sakshaug, E., Hopkins, C.C.E. and Øritsland, N.A. (eds.), 1991: Proceedings of the Pro Mare Symposium on Polar Marine Ecology. Polar Res. 10, 662 p.

7C. Sakshaug, E., Bjørge, A., Gulliksen, B., Loeng, H. Mehlum, F. (eds) 1992. Økosystem Barentshavet (Ecosystem Barents Sea). NAVF, NFFR, Ministry of the Environment, 320 p. 2nd edition: Universitetsforlaget, Oslo, 1994.

8C. Sakshaug, E. and J.-A. Sneli (eds) 2000. Trondheimsfjorden. Tapir, Norway, 336 p (in Norwegian).

Comments re 2C-8C:

2C: Pp. 5-31, 185-188, and 209-232 have been written by myself. This includes the chapters: "Resources of the sea", "What is oceanography?", "Costs and personnel in Norwegian oceanographical research, 1963-1972", and "A review of the history of Norwegian oceanograp-hical research up to the second world war"; the latter was written in cooperation with Professor H. Mosby, University of Bergen. The final version was approved by the Norwegian Oceanographical Committee

3C: The first 63 pages were written by myself. The final version was approved by The Norwegian Oceanographical Committee

4C: Introductory parts and parts covering pelagic research were written by myself. The document was extended with a chapter covering questions related to oil pollution and approved by the two research councils in question and the Ministry of the Environment in May 1983

7C: This book in Norwegian has been edited and in part written by the editorial committee on basis of input from participants in Pro Mare. For resons mainly of style the final versions of the chapters were in the main written by me. This book is the final report from Pro Mare to Norwegian fisheries and environmental managements as well as stafthe research councils. It also is used by ocean- and fisheries-relevant organizations and as a textbook for students (universities, district colleges etc). 2500 copies were printed; 2000 more were printed in spring 1994.

8C: This book has been written/edited on basis of inputs from NTNU researchers and colleagues at other research institutions in Trondheim. It was presented as a millenium gift from the Museum to the people of the Trondheimsfjord rregion.

Series D. Posters/Abstracts:

1D. Johnsen, G., Volent, Z., Tangen, K. and E. Sakshaug 1996. Time-series of potential toxic phytoplankton blooms in North european waters in 1993 using the SEAWATCH-Europe buoy system. Eos, suppl. 5, 35.

2D. Sakshaug, E. and Walsh, J. 1999. Barents Sea versus Bering Shelf ecology. Gordon Research Conference on Polar Marine Carbon Cycle, Ventura, California, March.

3D. Zielinski, O., Barth, H., Breves, W., Chauton, M., Godoy, J., Figueroa, F., Johnsen, G., Perez-Rodriguez, E., Reuter, R., Sakshaug, E. and M. Villgarcia. 1999. Underwater light field in relation to particulate and dissolved matter north of the Canary Islands. Final CANIGO meeting in Las Palmas. 12-16 September.

4D. Chauton, M., Zielinsky, O., Perez, E., Lorenzo, L. M., Johnsen, G., Egeland, E., Sakshaug, E., Reuter, R., Figueroa, F. and F. Figueiras. 1999. Pigment composition and in vivo fluorescence excitation spectra from the Canary Islands and Azores region. Final CANIGO meeting in Las Palmas. 12-16 September.

5D. Legrand, C. Johnsen, G., Granéli, E., Sakshaug, E. 2000. Effect of polyamines on growth and toxicity of Chrysochromulina leadbeateri and Prymnesium patelliferum (Haptophytae). 9th International conference on Harmful Algal Blooms. Hobart, Tasmania, 7-11 February 2000.P. 32.

6D. Johnsen, G., Legrand, C.C., Granéli, E, Sakshaug, E. 2000. The use of a DIVING-PAM for detection of irradiance, biomass and photosystem II fluorescence yields in harmful algal bloom experiments: Discussion of methodological aspects. 9th International conference on Harmful Algal Blooms. Hobart, Tasmania, 7-11 February 2000. P 144.

7D. Chauton, M., Johnsen, G., Legrand, C., Granéli, E., Sakshaug, E. 2000. Development of a potential harmful algal bloom in mesocosm experiment monitored through pigment analysis and bio-optical measurements. 9th International conference on Harmful Algal Blooms. Hobart, Tasmania, 7-11 February.

8D. Granéli, E., Legrand, C., Carlsson, P., Maestrini, S. Y., Hummert, C., Reichelt, M., Johnsen, G., Pallon, J., Luckas, B. and E. Sakshaug 2000. Nitrogen And phosphorus deficiency favours the dominance and toxin content of harmful species in summer Baltic Sea phytoplankton. 9th International conference on Harmful Algal Blooms. Hobart, Tasmania, 7-11 February 2000.