aratamomohara*andkiyohidemizuno**:habitat0fplants...

14
Jpn.』・Histor・Bot、Vol6.N().2p,49-62 Aprill999 植生史研究 ArataMomohara*andKiyohideMizuno**:Habitat0fPlants intheLatePlioceneSedimentaryBasin onAwajilsla、。,CentralJapan 百原新*・水野満秀**:淡路島の後期鮮新世堆積盆地における植物の生育地 AbstractLatel〕lioccneplantmflcrofossilassemblagesonAwajilsland,centl・alJapan,werestudiedtorecon, structthehabitatofplantsandtocomplementtheupperPliocenebi()stratigraphyoftheOsakaGroup、The lowerpartoflheAtag()Formati()I1yieldsmanyplantsextinctfr()mJapanandincludesCルOeバノ”0"dAzs“"〃応 andRe“ESノasp、thatcharacterizethebasalpartoftheOsakaGroupintheSennanarea・Stratigraphicoccur, renceofplantfossilsintheuI〕pcrPli()ceneonAwajilslandisverysimilart()thatinthcSennanarea・ Sedimentaryfaciesoffossilbearingbedswereclassifiedintothr・eetypes,i、e,,peat,massivesilt,andstratified sand,andoccurrenceofplanttaxawascomparedbetweenthesetypestoreconstructlheirhabitat.G卵/OSlIノり6ノイs Pe"s"is,7>”αandNymphaeaceaeincludedinpeatweregrowinginbackmarshesinanalluvialplain・Plants commoninmassivesilt,suchas/W““"0mg6”s/ノり60”es,S花z”ノ油〃ノ0ノMIdt〃加,鈍り"Omsp.,andR;e"dbAzlな α"za6〃た,composedf()restsinwetlandsandonsandbanksinanalluvialI〕lain・Plantscommoninstratifiedsand, suchasノ先easect.〃(.α7,βノぱ"sノ"/cノ”ノりノノノロI,andRZgjYs〃z九・ノひcαノカ",grewonfansandmountainslopesarounda sedimentarybasin、 KeyWords:Biostratigraphy,LatcPliocene,OsakaGroup,1,1antmacr()f()ssilasscmblages,Sedimentaryfacies 要旨これまでの大阪僻群の上部鮮新統植物化石層序を補足するとともに,当時の植物の生育地を復元するため に,淡路島の大型植物化石群を検討した”淡路烏の大阪層群の基底部を橘成する愛宕累#1下部からは,日本からの絶 滅極が多産する。この''1には,泉南地域の大阪層群の基底部を特徴づけるチャンチンモドキやチャセンギリ属が含ま れる。淡路島の上部鮮新統での植物化石の牌位的産出状況は,泉南地域とよく類似している。楠物の生育地を復元す るために,植物化石群含有牌の堆積机を泥炭層,塊状シルト,葉理の発達した砂ハの3つに区分し,化石の産出状況 を比較した。泥炭層に含まれるスイショウ,ヒシ属,スイレン科は沖積平野の後背湿地に生育し,塊状シルトに多く 含まれるメタセコイア,ヒメシャラ,セコイア属,イヌカラマツは沖積平野の自然堤防とその周辺の湿地に分布して いた。一方、葉理の発達した砂I曽内に多く含まれるトウヒ属パラモミ節,ツゲ,ヒメプナは堆積盆周辺の扇状地や山 地斜面に生育していたと考えられる。 キーワード:大型植物化石群,大阪胴群、後期鮮新世,生層序、堆積相 Intr⑪duCtion Changesinthegeographicaldistributionofplantsin ,daroundasedimentarybasininducedbyenviron. andaroundasedimentarybasininducedbyenviron‐ mentalchangescausedextinctionandevolutionof plants・Momohara(1994)sh()wedthatthehabitat()f respectiveplantspeciesinandaroundasedimentary basinwasaffectedbytheupliftofsurroundingmoun‐ tainsandtherelativesealevelchangesthatcontinued sincethelatePliocene.’)lantslivinRinthel()wland, suchasル化"se9"。〃g卯/0s/ノ℃60/此s,becameextinct fromJapanprobablyduetothegeomorphological changesinthclastphaseoftheearlyPleistocenein additiontotheclimaticdeterioration(Momoharag/ "/、,1990).Rec()nstructi()nofpastReomorphologyand habitatofplantsinandal・()undasedimentarybasinis, thus,c()mplementaryt()discussfl()ralchangessince thelatePliocene・ Reconstruction()fpastplants’habitatisusually 籾DepartmeI1tofEI1vir()nmenmlScience,FacultyoIH()rticulture,ChibaUniversity,Matsudo648,Chiba271-8510, Japan. 〒271-8510千葉県松戸市松戸648千葉大学園芸学部緑地・環境学科 寧拳OsakaCenterforRegi()nalGeology,GeologicalSurveyofJapan,Govt・Bldg.N().2Bekkan,Otemae4-l-67,Chuo-ku, Osaka54()-0008,Japan. 〒540-0008大阪市中央区大手前4-167大阪合同庁舎第2号別館地質調査所大阪地域地質センター

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Page 1: ArataMomohara*andKiyohideMizuno**:Habitat0fPlants ...hisbot.jp/journalfiles/0602/0602_049-062.pdf11111111111111111111111111 2222212 1 131411 112 11111111111111111111111111 1211

Jpn.』・Histor・Bot、Vol6.N().2p,49-62 Aprill999 植生史研究

ArataMomohara*andKiyohideMizuno**:Habitat0fPlants

intheLatePlioceneSedimentaryBasin

onAwajilsla、。,CentralJapan

百原新*・水野満秀**:淡路島の後期鮮新世堆積盆地における植物の生育地

AbstractLatel〕lioccneplantmflcrofossilassemblagesonAwajilsland,centl・alJapan,werestudiedtorecon,

structthehabitatofplantsandtocomplementtheupperPliocenebi()stratigraphyoftheOsakaGroup、The

lowerpartoflheAtag()Formati()I1yieldsmanyplantsextinctfr()mJapanandincludesCルOeバノ”0"dAzs“"〃応

andRe“ESノasp、thatcharacterizethebasalpartoftheOsakaGroupintheSennanarea・Stratigraphicoccur,

renceofplantfossilsintheuI〕pcrPli()ceneonAwajilslandisverysimilart()thatinthcSennanarea・

Sedimentaryfaciesoffossilbearingbedswereclassifiedintothr・eetypes,i、e,,peat,massivesilt,andstratified

sand,andoccurrenceofplanttaxawascomparedbetweenthesetypestoreconstructlheirhabitat.G卵/OSlIノり6ノイs

Pe"s"is,7>”αandNymphaeaceaeincludedinpeatweregrowinginbackmarshesinanalluvialplain・Plants

commoninmassivesilt,suchas/W““"0mg6”s/ノり60”es,S花z”ノ油〃ノ0ノMIdt〃加,鈍り"Omsp.,andR;e"dbAzlな

α"za6〃た,composedf()restsinwetlandsandonsandbanksinanalluvialI〕lain・Plantscommoninstratifiedsand,

suchasノ先easect.〃(.α7,βノぱ"sノ"/cノ”ノりノノノロI,andRZgjYs〃z九・ノひcαノカ",grewonfansandmountainslopesarounda

sedimentarybasin、

KeyWords:Biostratigraphy,LatcPliocene,OsakaGroup,1,1antmacr()f()ssilasscmblages,Sedimentaryfacies

要旨これまでの大阪僻群の上部鮮新統植物化石層序を補足するとともに,当時の植物の生育地を復元するため

に,淡路島の大型植物化石群を検討した”淡路烏の大阪層群の基底部を橘成する愛宕累#1下部からは,日本からの絶

滅極が多産する。この''1には,泉南地域の大阪層群の基底部を特徴づけるチャンチンモドキやチャセンギリ属が含ま

れる。淡路島の上部鮮新統での植物化石の牌位的産出状況は,泉南地域とよく類似している。楠物の生育地を復元す

るために,植物化石群含有牌の堆積机を泥炭層,塊状シルト,葉理の発達した砂ハの3つに区分し,化石の産出状況

を比較した。泥炭層に含まれるスイショウ,ヒシ属,スイレン科は沖積平野の後背湿地に生育し,塊状シルトに多く

含まれるメタセコイア,ヒメシャラ,セコイア属,イヌカラマツは沖積平野の自然堤防とその周辺の湿地に分布して

いた。一方、葉理の発達した砂I曽内に多く含まれるトウヒ属パラモミ節,ツゲ,ヒメプナは堆積盆周辺の扇状地や山

地斜面に生育していたと考えられる。

キーワード:大型植物化石群,大阪胴群、後期鮮新世,生層序、堆積相

Intr⑪duCtion

Changesinthegeographicaldistributionofplantsin

,daroundasedimentarybasininducedbyenviron.andaroundasedimentarybasininducedbyenviron‐

mentalchangescausedextinctionandevolutionof

plants・Momohara(1994)sh()wedthatthehabitat()f

respectiveplantspeciesinandaroundasedimentary

basinwasaffectedbytheupliftofsurroundingmoun‐

tainsandtherelativesealevelchangesthatcontinued

sincethelatePliocene.’)lantslivinRinthel()wland,

suchasル化"se9"。〃g卯/0s/ノ℃60/此s,becameextinct

fromJapanprobablyduetothegeomorphological

changesinthclastphaseoftheearlyPleistocenein

additiontotheclimaticdeterioration(Momoharag/

"/、,1990).Rec()nstructi()nofpastReomorphologyand

habitatofplantsinandal・()undasedimentarybasinis,

thus,c()mplementaryt()discussfl()ralchangessince

thelatePliocene・

Reconstruction()fpastplants’habitatisusually

籾DepartmeI1tofEI1vir()nmenmlScience,FacultyoIH()rticulture,ChibaUniversity,Matsudo648,Chiba271-8510,

Japan.

〒271-8510千葉県松戸市松戸648千葉大学園芸学部緑地・環境学科

寧拳OsakaCenterforRegi()nalGeology,GeologicalSurveyofJapan,Govt・Bldg.N().2Bekkan,Otemae4-l-67,Chuo-ku,

Osaka54()-0008,Japan.

〒540-0008大阪市中央区大手前4-167大阪合同庁舎第2号別館地質調査所大阪地域地質センター

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50 Jpn..'・Histo1。.B()1. Vol、6.N0.2

basedonthepresentccologyofextantplants,This

analogyis,however,problematicforplantsextinct

fromJapan・ThefloraofthelateP1i()ceneincludes

abundanttaxaextinctfromJapan(Mikil948;Itihara,

1961:Momohara,1992).Mostofthemarenowcon‐

finedtolimitedareasinChina,andtheirrelictstatus

hardlyrepresenttheirpastecologicalpreferenceThe

naturalvegetationinChinahasmostlybeendamaged

throughthelonghumanhistory,andananalogical

argumentbasedonplantsinpresentlowlandareasis

impracticableduetotheextensivealldcontinued

humanactivities,Besides,thelatePlioceneflora

includesmanyf()ssilspecies,andtheecologyoftheir

nearestlivingrelativesisnotnecessarilythesameas

thatofthefossilspecies・

Thoughecologyofpresentplantspeciesisnot

applicableasallellvironmentalindicator()ffossil

taxathespacialdistributionoffossilsinasedimen‐

tarybasinandastudyofthedepositionalenvil-on‐

mentsoffossilassemblagesserveaskeysforthe

reconstruction()fpasthabitatoffossilplants・Suzuki

(1976)comparedthespeciesc()mposition()ffossil

assemblageswiththeirdistributioninthelate

MioceneandPliocenesedimentarybasininAizu

Prefecture,northernJapan,andrec()nstructedthe

specialandtemporalchangeofplantcommunitiesin

thesedimentarybasin.M()mohara(1994)recognized

twokindsofsedimentaryfaciesinfluvialdepositsof

theOsakaGroupthatincludcplantmacrofossil

assemblages,unstratifiedpeatysiltswithmoreorless

autochthonousplantsandwell-stratifiedsandysedi‐

mentswithplantstransportedfl・()mwideareas.Dif‐

ferenceofthespeciescompositi()nbetweenthesetwo

typesofsedimentaryfaciespartlyreflectsthedis‐

tributionalpattemofplantspeciesinthesedimelltary

basin、Fossilassemblagesinunstratifiedsiltsh()uld

consistofplantsthathavegrowninandaroundthe

backmarshinanalluvialplain,andthoseinsandy

sedimentshouldincludeplantsgrowingonmountains

farfromtheareaofdeposition(Momoharal994).

Inthisstudy,wewilldescribeplalltmacrof()ssil

assemblagesinthelatePlioceneoftheOsakaGroup

onAwajilslandandwillreconstl-ucthabitatofplants

fromthesedimentaryfacies・TheOsakaGroupdis‐

tributedincentralJapanconsistsmainlyoffluvial,

lacustrine,andmarinesedimentsthatweredeposited

continuouslysincethelatePliocenetothemiddle

Pleistocene(Itihal-a,1961).Themaglletostratigraphy

andthetephrostratigraphyofthisgrouphavebeen

examinedindetail,andthemostdetailedplantma‐

crofossilbiostratigraphyhasbeenestablishedamong

thel〕lio-Pleistocenestrataintheworld(Itihara,1961:

Momohara,1994).ThelowermostpartoftheOsaka

Group,i、e、,upperPli()cene,isdistributedwidelyon

Awajilsland(Mizuno,1993),westoftheOsakaBay,

andintheSennanarea,eastofthebay(Fig.1,Itihara

e/α/、,1975).Miki(19481950,1952,l957andl960)

reportedplantmacrofossillocalitiesonAwajilsland,

butdidnotdescribelocationsorstratigraphyindetaiL

Ourstudyaimstocomplementtheplantbiostratigl-a、

phy()ftheOsakaGroupwithmaterialsfromAwaji

lsland.

Litho-andtephrostratigraphyoftheOsakaGroup

onAwajilsland

O11AwajilslandtheOsakaGroupiscomposedof

fluvialsedimelltsincludinggravel,sandandsilt

(Mizuno,1993).TheOsakaGr()upontheislandis

dividedintonorthem(areasA-CinFig.’),middie

(areasD-O),andsouthem(areasPandQ)parts・

Becausethisgroupisdistributedinisolatedareas,

stratigraphicsubdivisionsandvolcanicashlayersin

eachareaarenamedseparately(Fig.2,Mizuno,1993).

TheOsakaGroupinthenorthernpartoftheisland

(areasA-C)reachesabout600mthickandisdivided

intotheToshimaandKariyaFormationsinthe

ascelldingordel・(Fig.2,Mizuno,1993).TheToshima

Formationisdividedintoalowerpart,mainlycom‐

posedofalternatingbedsofsandandsiltwithocca‐

sionalgravellysedimentsandanupperpart,consist‐

ingofoverwhelmin貝sandandgraveLTheKariya

Formationisdividedintoalowerpartofalternating

bedsofsandandsiltandanupperpartofsandand

graveL

TheOsakaGroupiswidelydistributedinthemiddle

pa1.t()ftheisland(areasD-O).Itisabout450mthick

andisdividedintotheAtagoandGoshikihamaF()r‐

mationsintheascendingorder・TheAtagoFormation

isdividedint()alowel-partmainlyofgravel,amiddle

partofalternatingbedsofsandandsilt,andanupper

parlmainly()fsandandgravel、TheGoshikihama

Formationcollsistsofwell-sortedpebble-cobble

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4.45IE

HabimtofPlalltsinlhcLatel〕liocencSedimcntaryBasin(Momohara&Mizun())

51

"'135.0‘E"1

Harima-NadaHarima-NadaKobiwako

GroupKobiwako

GroupKobiwako

GroupOsakaG

n~へ<‐:.,夢

OsakaG

n~へ<‐:.,夢

OsakaG

n~へ<‐:.,夢 聯聯

「Cup'?丙「Cup'?丙「Cup'?丙

N零“

~一忽Awaiilsland~一忽Awaiilsland~一忽Awaiilsland

へ…′Sennan

area

へ…′Sennan

area

へ…′Sennan

area

。30ON

4.45IE

ToshimaFormationtotheupperpartoftheAtago

Formati()n.TheKariyaF()rmaitoncanbecorrelated

withtheGoshikihamaFormati()nbasedontheexis,

tence()Icrystallineschistillthcgravellaycrs

(Mizun(),1993).

TheOsakaGroupinthes()uthernpartoftheisland

(arcasl)flndQ),namedYudaniFormation,ismore

thanlOOmthickandmainlycollsistsofsandand

gravel(Fig.2;Mizuno,1993).ThestratiRraphicposi‐

ti()nofthisformationisuncertainbecauseofthelack

ofv()lcanicashlayersillthisf()rmati()n.

Somev()lcanicashlayersintheOsakaGroupon

Awajilslandcanbec()rrelatcdwiththoseinthe

layerswithprevalentsiltandclaylayers,andits

gravellayerscharacteristicallyincludeabundantcrys、

tallineschist・

Basedonvolcanicashlayersandc()mp()sitionoI

gravellayers,theOsakaGroupofthemiddleflndthe

northernpartsoftheislandcallbec()rrelatedwith

eachother(Mizuno,1993).TheAsan()lVolcanicAsh

LayerintheToshimaFormati()ncorrespondstothe

SakoVolcanicAshLayeroftheAtag()F()rmation,

andtheAsano2VolcanicAshLayert()theShitoori

VolcanicAshLaycl.(Fig.2).Thelowcrl〕a1.t()fthe

ToshimaFormati()ncorrespondstothemiddleI)artof

theAtagoF()rmati()、,andtheupperpart()fthe

ayKK

、受塁&盤;、受塁&盤; N

I:2'

lN34.23

卿淑67

Fig.1F()ssill()calilies(1-72)andthedislributi()、()fthcOsaka

Gr()up(a1℃asA-Qcnclosedbyl)I・()keIllille)oI1Awajilsland.(Ithiharae/α/,,1975)

f、

111

,イ

54

唖蹄

10km利く,a70

聯z〕0

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JpnJ,Histor、Bol Vol,6.No.2

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画gravel

ⅢbasemenI「ocks

Hyvolcanicashlayer国医匡匡

macrofossilassemblagewiththeigraphicpositioninsectionsA-QI|雌

boundaryofformation三三一一一

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亡、。』の。。.ごO雲何E」oLmE三⑳o」|姫

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52

Fig.2Standardc()lumnarsecli()ns()flhcOsakaGroupinareasA-Q(f()11()wiI1gMizuno,1993)andcolumnarseclions()fthef()ssill()calilies(1-72).ThearrowatthelefthaI1d()fc()lumnarsccti()、indicatesthe

stratigraphicposition()(1)lantmaCl・ofossilassemblagesatl()calilieswith()utv()lcanickeybeds・

VolcanicashlayersKm:KamaguchiVolcanicAsh,Al:AsaI1olV()lcanicAsh,A2:Asano2Volcanic

Ash,Y():YokoyamaV()lcaI1icAsh,Kr:KurumaV()lcanicAsh,My:My()jinVolcanicAsh,Oz:Ouji

VolcanicAsh,Sk:Sak()V()1canicAsh,St:ShitooriVolcanicAsh,K1:Kenj()gaokalVolcanicAsh,K2:

Kenjogaoka2VolcanicAsh,K3:Kenjogaoka3VolcanicAsh,K4:KenjoRaoka4VolcanicAsh.

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53

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一一一』‐一{一一一一卜』一一一一一一豆

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栂P

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冒弓匡

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AtagoFormation,middlepart

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Plantmacrofossilassemblages

Ji〕''.』・Histor.B()t,54

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121

32

AshLaycr,theShit()()riVolcanicAshLayerwiththe

AsashiroVolcanicAshLayer,theYokoyamaVol‐

canicAshLayerwiththeShinnoikeVolcanicAsh

Layer,andtheKurumaVulcanicAshLayerwiththe

OsakaGroupoftheSeImanareaandwiththosein

theKobiwakoGroup(Fig.3;MizuI1o,1992).Between

AwajilslandandtheSennanarea,theSakoVolcanic

AshLaveriscorrelatedwiththeIIabutakillVolcanic

12

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411

12‐9a・b

2j:。''鴨.45113

114

1卿

糾糾型‐‐‐I‐‐‐‐‐‐‐‐‐‐‐‐洲‐Ⅷ‐‐‐‐‐兼Ⅷ

V()1.6.No.2

状乃/Wa刀店c、/pro/77e"b/dbs

ルたゐazeda'石ch2

≠Cノラoemspo"oウ白Sax"Za"S

≠Rどe”s/asp

*Fbm"7ea砲S"7e"S/S

虻々 U雄/77barSp

羊ルク汐ノリリbdbノフoウ℃"xWbcaノpリノ77

秋〆e店e砲sp

Que”ussubgen.CγC/bbaAa"OPSノs

*F韮us/77/brocaノpa

掌Que”ussect,Ceノァ7ぷ1

*Sどquoおsp

Srewam白mo"a咋ノリ、ノウa3

*GルpオCs”buspe"sノルS

掌Pseudbわノ1スra/77abノルS3

BI"wノs/77/bmp方j"Za

Rbeasect・P7bea 2

*雌rasequoわg〃pオCs”bo姥s3

*Q77"aノ770/77L"77/77aCmpOdt"77

AcerO治bo化Um

≠ノリbmaノ77e応pamo加/pbs

FけうUssubgenhbP/bXWbノフ

Aル'7usノョpo"ノca

Rheノリb咋"0ケ℃ノ7ヨノ77α”"se

状CjノC/bcan'apaノル”s

Cbn"ヒノs力etemphj'Ⅵルヨ

0111

(エ⑪N四ョ山)

工凹ロ匡庁口重一一

Ⅱ1日18ⅡⅡ

(エONO一コ一一)

12

131

11

11111111111111111111111111

2222212

131411

112

11111111111111111111111111

1211

111

2213

23341123

44

3232

22

21132

1.91.7

アm画の三3

11日10一日Ⅱ

StratiR1・aphicoccurrenceofselectedplanttaxaandcorrelatiollofvoIcanicasheswiththose

inthe()sakfl(}r()upinSenI1anAreaandtheKobiwak(〕Gr()up・’’1anlmacr()fossilassemblagesinthel()werandupperpartsoftheToshimaF()rmationareiI1cludedinthemiddleandupperpartsoftheAtaIE〔()Formati()、,respectivelV、TheassemblagesinthelowerpartoftheKariva

F()rmationareincludediI1the(;oshikihamaF()rmation・Taxawilhallasterisk2lreplantsextinctfromJapan・NumbeI・sineachtaxonindicatethenumberoff()ssilassemblageswithineachh()riz()n.Mflgnet()stratigraphyfollowsMizuno(1992).

②ゴーココ◎完①

ⅡⅡ■1日一Ⅱ■

11

VolcanicashlayersintheSennanarea

and(KobiwakoGroup)

Fig.3

Gauss Matuyama OlduvaiMagnetostratigraphy

Age(Ma) ←30→ 25

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IIabitatofP1antsintheLatel)lioceneSedimentarvBasin(Momohara&Mizuno)

TablelSystematiclistofplantfossilsfromtheOsakaGrouponAwajilslandPartsofI)lantfossilsarcsh()wnintheparentheses:b:bud,c:cone,Cs:conescale,Cu:cupule,e:endocarp,f:fruit,fs:fruitingIscale,i:infructescence,l:leaf,s:seed,sh:shoot・

Asterisksindicatelaxaextinctor、olongerpresentinJapan.

CephalotaxaceaeCephaノoIax“sp.(s)

CupressaceacaIamaeCyp”心”旅rα(Sieb・etZucc.)Endlicher(sh,s)

Taxodiaceac

車MEIaSe9"ojagjyPIos"obojdeSHuCICheng(c,s,1,sh)*αypIos"ob“pe"sj"s(Staunl.)Koch.(c,s,sh,l)*C"""i"ghα"mjasp.(c,l)*Se9"ojasp.(c,s,sh)蝋71αiwα"jac”jomerjoid“Hayata(c)

Pinaceae

*Kを舵ノeeriasp.(c,l)*PiceaAorjbaiMiki(c,l)Pjceasect.Piceα(c,l)Pi""ssubgen、Djpノoxyノo〃(c)Pj""ssubgen、比〆oxyノo〃(1)*応e"。b/αrixamab"jS(Nclso、)Rehd.(Cs,s,sh,l)聯RseadoIs"gα皿bra""dαMiki(c)

Juglandaceae*Cycノocα〃apa""r"s(Batal.)Iljinsk.(f)*Jagノα"s腕egac‘"ereaMiki(f)噸P花rOcα'γαS舵"QPZeraCDC.(f)

Salicaceae

SaノixsP.(f)Bctulaceae

Aノ"us〃po"jca(nlunb.)Steud.(i,fS,f)Ca'pj""sIscho"osAijMaxim.(f)CO〃ノasノiaerQpAy"αFisch.(f)

Fagaceae漁fhg“mjCrOcamaMiki(Cu,f)Q"erc"sα"e"αBlume(Cu)Q"erc“(b,f)Querc"ssubgen、Lepidobaノα"“(Cu,f)噸Q"erc"ssect,Cerris(Cu)Q脚erc“subgen,Cycわbam"Opsjs(Cu)

Ulmaceae

ZeノACvasp.(f)Polygonaceae

Pりりgo"""'sp.(f)Magnoliaceae

ノVag"oノiapraecoassjmaKoidz.(s)Schisandraceae

Schisα"drarepα"。α(Sieb、etZucc.)Radlk.(s)Lauraceae

噸α""α碗o腕"腕'"αcrOPOd"腕Miki(S)Menispermaceae

Sj"ome"i"胴α“畑胴(Thunb.)Rehd・etWilson(s)Nymphaeaceae

Brase"jascAreberjJ.F、Gmel.(s)*E"〃αノesp.(s)M4pAersp.(s)

CeratophyllaceaeCemIopAy""、。e腕ers"mL.(f)

Ac1inidiaceae

Aα”djar‘4/tz(Sieb、etZucc.)Planch.(s)AC""jdjasp.(s)

Theaceae

E"〃αsp.(s)Sjewar"α、o"αdeゆノ'αSicb・etZucc.(f)

Hamamelidaceae

車Fbrrα"eariasj"e"s応Rehd、etWils.(fs)噸比加α腕eノなPα〃O"o〃eSMiki(f,s)噸L々"dambersp.(f)

SaxifragaceaeSchizQpAra877zαノjydrα"geojdesSicb、etZucc.(f)

Rosaceae

Cra1aeg図ssp.(e)

Leguminosac

恥Ieriαβorめ""。α(Will。.)DC.(f,b)Lcguminosae(f)

EuphorbiaceaeWe”"escordaIaR,Br.(s)a4pAorbjasp.(s)*SaPmmseb縦ノw"1Roxb.(s)

Rulaceae

Phe"ode"dro〃α'""re"seRupr.(s)Zα"的oxyル腕α"α"IAojdesSieb,eiZucc.(s)Zα"rhoxyノ"胴”er"以腕(L、)DC.(s)

Meliaceae

Me"ααze‘jaracAL.(e)Anacardiaceae

*CAoerosPo"djas“"αrな(Roxb.)BurttetHill(e)Aceraceae

,4cerdjabo"“"zBlumeexK・Koch(f),4cercfmo"oMaxim.(f)Acersp.(f)

Sabiaceae

Me"osmasp.(e)Stzbjaノapo"jcaMaxim.(e)

StaphyleaceaeSIapノiy/eab"mαノdα(Thunb.)DC.(s)

Buxaceae

B"x“"'jcrQphy"αSieb、eIZucc.(1)Rhamnaceae

Berche胴jasp.(e)*”""r"s〃”o"jc"sMiki(f)

Vitaceae

IノY1jssp.(s)Sterculiaceae

*Reevesjasp.(f)Lythraceae

Lagers"oemjasp.(f)Trapaceae

乃叩αj"cisaSieb、etZucc.(f)*乃apasP.(f)

AlangiaceaeA/α"gj"msp.(c)

Comaceae

Be"的α、城αノapo"jcaHara(e)Cor"usco"rroversaHemsley(e)Cor""s胴acrQpノ!y"αWall.(c)

Araliaceae

AraノmeノαIα(Miq.)Seemann(e)Ericaceae

〃"α"jA"ssp.(f)Styracaceae

We"わ。e"dro"xyノocα'p"、Hand.-Mazz.(e)SlWaxノ叩o"jcaSieb・etZucc.(s)SlWaxsP.(s)

SymplocaceaeSympノocossp.(f)

Verbenaceae

Qm"icamasp.(e)CノerOde"。r"mrriChOrom”Thunb.(e)

CaprifbliaceaeIノYb””msp.(e)

Alismataceae

Sbgj"αriasP.(f)Sparganiaceae

Spargα"地mSP.(e)Cyperaceae

Qzrexscct・Qzrex(f)CnrexspP.(f)Sと”"scf、ル"cojdesRoxb.(f)

00

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9a34・27‘41.N,134.50‘14.E10

9b34.2741.N,134.50‘14・E10

10a34・27'41。N,134・50‘17.E10

10b34、2741.N‘134.50117.E10

1134.2736.N,134.4957・E20

1234.26,52口N,134。49l30oE45

1334.27I26pN,134.50I40qE25

Jpn.』・Ilistor,Bot.56

SSMMMSS

ListofplaI11macr()f()ssilswilhthel()cationandsedimentary(aciestypes()flheplanlfossilbearingbedAbbreviationsforplantpartsareinTablel・Abundanceinplantmacrof()ssilassemblagesareindicatedaftertheparentheses;A:abundant,C:c()mmon・Sedimcnmryfacies;S:stratifiedsalld,M:massivesilt,I〕:pcat.

Table2

34.27.03qN‘134.50126。E32

34.27049.N,134.54145.E90

34.2745nN'134.54148,E85

34.27‘19.N,134.55‘44口E30

34.25'35.N,134.52‘56・E30

1bssillaⅢudeIlon9iludeal1i-sedi-

assem‐IudemenIa「y

bla9e(、)faciesnumbe「

1axa

MMPSS

34.30‘49.N,134.59‘O5QE30M

34・31‘32W,134.59016qE30

34.32'48側N,134.56127冊E25

34.32‘26.N,134.56132・E30M

34.3228.N,134・56143.E40S

EuノyaIesp.(s)

Me伯sequoja9IyPIosIrobojdes(c);Quemusa#ena(Cu)

ⅧSSP(s);SIy78xノaponjca(s)

GIyPIosImbusPensj#s(Cs‘sh);Sequojasp.(c,s,sh);Mel1Osm8sp.(e)

PSeudojarixamabiノis(Cs),C;P1emca『yasfenqpIela(f);Quercusa#ena(Cu),A;Lj9ujd8mbarsp(f)jSChjZQph旧9ma

hydran9eojdeS(f);ACerd伯boI1bum(f);Me胸smasP(e)iBuxuSmicmPhyノ伯(|);Bemhemiasp.(e);MeIlbdendmnxyjocaノpum(f)i

Sympmcossp.(f);CaMCa巾asp(e);C劇exsp.(f)

G1しP(os1mbuspensjjs(c,s);Sequojasp.(c,s,sh),A

Quefcus8#ena(Cu,f),A;S1ewanYamonade価a(f);ZanfhoxylUmaj伯nIhojdes(s);Buxusmjc和phyノノa(|)

Cham8ecyPa応PjsjIbra(sh,s),C;Cumin9hamjasp.(1);Me伯sequoia91ypiostroboides(1,sh)iSequoj8sp.(sh);Pjceasect・

Pにea(|);Pseudo畑jX8mabjIis(|);Pinussub9en・HapIoxylon(|);Sa伽sp,(f);Wisferj8/brjbunda(b);BuxusmjCmphy"a(|);

Cal#CaPesP.(e);CayBxsp.(f)

MeIasequoi89”osfmbofdbs(c,s);Ju9ノansme9acjheYG8(f);Quef℃ussp.(f);Ma9no脂”ecocjssjma(s)iCjmamomum

m8aoPodum(s);PheModendmnamu旧nse(s),C;Z8nihoXyルma脂nIhojdes(s);ZanIhoxy〃mpjpe伽、(s);S1yraxノ8ponjca(s),C

Cjm8momumm8cmpOdum(s)

Mef8sequoia9b/PfosImbades(c)

C研a1oIphyノlUmdemefsum(s)iSpaf9aniumsp(e),A;Sc〃puscf〃ncoides(f)

MeI8sequoi891i/PIosIroboides(c),C;W1sienaHmbunda(b)iAIeu耐escord8Ia(s)ISfy伯xj8ponjca(s)

G”osImbusPensjノis(c);Euノya伯sp,(s);NUphersp.(s)i71rapesp.(1);Sp副9an心msp.(1)

AInUsj8ponにa(0

Me伯sequoja9IyPIosImbojdes(c),A;PseudoI師x8mabjIS(sh);Quefcus8ljena(Cu)iQu研cussp(1)iMa9no脂sp.(s);W1S1e"a

I1ov↑bund8(b);EUPhovbjasP(s);Pheノリodendのnamu尼nse(s);BenIh8mjd18japonjc8(e)!C;SiWaxjaponjCa(s)

Zelkov8sp.(f),C

Pseudo/8伽8,8b"s(Cs,sh)!C;CycノocaノyaPa伽「us(f);Alhus/aponica(fs);CamjnusIschonosk"(f);CO『yルsheI劇ophyノ伯(f)!C;

H8m8melISparmIioides(f,s);Siyraxノaponjc8(s)

MeIasequoja9Iypios"obojdes(c);Pseudo1a"xamab"s(sh);Covylusheferophyノ13(0;Comusmacrophyノノa(e)

Sequoi8sp,(c,s,sh),C;C副BxsectCarex(O

GlypiosImbuspens"is(c1sh),A;W8pasp.(f)!ClEu『y8伯sp‘(s);SMaxsp,(s)

MeIasequoj89Iypfosfrobojdes(c);Sequioasp.(sh);Pjce8kmb8i(|);Quercussp.(f)I8uxusmiaOphyjIa(s)iVjbumumsp(e)

Sequoi8sp.(c);Pseudbla『jxam8bj#s(Cs);PjCeasectPjcea(1);Qu研cusaIjena(Cu),A;Qu研Cussp(1);Sねw師旧monadeIpha(1),C;Wisfevia伽banda(b);EnkjanIhussp(1);SIy7axjaponjca(s)

Pa伽rusnjPponjcuS(f)

Pjce8sec1Pjbea(c)

万ap8sp.(f),C

Fa9ussp.(f)

Pjce8sec1、PjCea(c)

Quef℃usalena(Cu)

Sequojasp.(s,sh);QuefCussub9en・Lepjdoba伯nus(Cu)iS1ew師jamonadeIpha(f);S版axノaponjCa(s)MeIaqseqUoia91MDIosImbojdes(c);Pseudola「jxamabjlIS(Cs,sh),CIPにe8k”b8i(1)!C;Pjceasp(sh);Cycjoca0/apa伽rus(f);C”inusIschonoskij(f)iCOvy川shefemPMa(f);Queにus8#en8(Cu),A;QueCussp.(f);Hamame応p6MmWojdes(f,s);Acersp(f);Arajaej8fa(e)

GIypfosfmbuspensil1S(c,s,sh)iAlhusノaponjca(i);77Bpajncisa(1)

Me伯sequoia9卯IosIrobojdes(c);Ca"camasp.(e)

MeI8sequoia9Iypfoslmboides(|)iPseudbIarjXamab"s(Cs,sh);CaノpjnusIschonosk〃(f);Queにusa#en8(Cu);Quefcussp.(f);SIew師j8monade伽a(1);Sty畑xノaponj℃a(s)

Mei8sequoia9IypIosiroboides(|);PjceasectPjcea(1)iAcWnidiaru的(s)iW1sieri8Ib"bunda(b)iSIyraxjaponjca(s)Ceph8Ioi8xussp,(s)

GlypIos1robuspensj#s(sh);Ce「a1OpMUmdeme庵um(1);万8p8jhcisa(c)‘A;Sp8r98n畑msp(1)Sinomenjumacufum(s)

AIan9jumsp(e)

Sequoi8sp.(c1sh)iCjnn8momumma”podum(s);Sfewa〃j8monadeIPha(1);Aノeu"IescordaIa(s);CノerodendrumI"chotomum(e)Kefelbenasp.(C,|)iPjCeasec1・Pjcea(c)iPseudoノafjxamabjIjs(Cs);Fa9usmjcmc8Wpe(Cu);A伯n9jUmsp.(e)iSIyYaxsp.(s)Quefcusal1ena(Cu);Quef℃ussub9en、CyclbbajanOpsjS(Cu);Qu研℃ussp.(0;A1eu耐escofda伯(s);BuxusmjcmPhyIja(s);耐pasp.(1);SIyraxノ8ponjC8(s)

Pseudo伯rjxamab"s(Cs),C;SIew師jamonadeIpha(1)

Me伯sequoja91ツp(osImboIdes(c);PseudolanXamabjlls(Cs),C;Quevcussub9enLepjdobaIanus(Cu);SfewarWamonadeIpha(f)jSI8phyje8bumald8(s);Le9uminosae(f)

Sequoiasp.(c);SIewaノfjamon8deIpha(f);BuxusmjCmPhyIIa(|)

2345

34.31'56.N,134.55,19.E62

34.31'28.N,134.55'15oE100

34o30'30口N,134.53I55pE70

MMS

67髄

8b34.30'33.N,134.53‘56.E88M

Vol6.No.2

25a34。25'10.N1134.5035.E

25b34・25010,N,134.50I35pE

2634.2414アN,134.51'55・E

1434.27‘10.N‘134.50‘54qE50M

1534,27IO4pNl134‘52'O2qE30M

帥帥乃

3534.23‘42.N,134.49'56・E95M

67890

11112

2134.25106.N,134.53‘56・E40

22a34・25‘07.N,134.53'54・E40

22b34・25‘07.N,134.53'54・E40

22C34・25007.N,134.53‘54oE40

22.34.25107.N,134.53‘54・E40

23a34・24159.N,134・52‘25.E95

23b34・24‘59.N,134.52'25。E95

2434.25‘09口N,134。51I16QE85

MMS

34a34.23‘33oN,134.50‘58.E150M

34b34,23f33PN,134.50F58oE150M

MSM

2734.24139nN,134.51121側E75

2834.25124"N,134.50'05"E100

2934.24‘50.N,134.50‘21.E110

3034.24‘47口N,134.50‘17・E110

31a34・24132"N1134.49'34.E130

31b34・24132nN,134.49'34qE130

3234.24・24.N,134.49‘46.E120

3334.24‘31.N,134.49‘48.E130

SSMSSSSS

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5234.2038W,134・45‘28.E50M

5334.20'19。N‘134.46046・E25M

5434.14.45。N,134.46‘16oE75

5534.18‘21.N‘134.49,33oE50

5634.18028.N‘134.5224。E15

5734.18141.N,134.52,O5oE50

llabital()fl)lantsintheLatePlioceneSedimentaryBasin(Mom()hara&Mizuno)

assem・latitude,畑19ihmealI,sed

number(、)facies

IaXa

MMMS

3634.2250W,134。4916.E128M Mefasequoja9”oslmbojdes(|);QuefcusaIena(Cu);QuefCussectCems(Cu),C;Quercussp.(f);SchjSand『aノepanda(s);MJphe「sp.(s);SIewar1jamonadeゆha(f);HamameIspaym↑jbjdbs(f,s);Meルosm8sp.(e);StWaxノaponj℃a(s)

773”jhcjsa(f);刀apasp.(f);Braseniasch旧be〃(s)

Pjce8sectPjcea(c,s)

Quefcusseb9enLePjdobaノanus(Cu)

ViWSsp,(s)

SequojasP(c,s);Fa9usmjcroCaのa(Cu)iQuemusa#ena(Cu);SIWaxjapon/Ca(s)

Sequojasp.(c,s,sh),A;Fa9usmjcmcama(Cu);Aleu"iescada伯(s)iSapiumsebj航um(s);MeノjOsm8sp(e)iComusconImve応8(e);SIyr8xjaponjba(s),A

MeI8Sequoja9卯fosfmbojdes(c),C;Sil"ax/apo、にa(s)

MeI8sequoja9Iypiosimbojdes(c)‘A;Styrax/aponic8(s)!C;H8m8me#Spa汀o#ojdes(f,s)!C;WisIe"aノリov1bunda(b)

MeIaSeqUoia9卯iosIrobojdeS(c)

Mef8sequoja9IyptosImboides(c)

Met8sequoj89IyPioslmbojdes(c)

Me的sequoj891ypfoslmbojdes(sh);PにeasectPにe8(1);Pseudbノ8『jX8m8bjノノs(Cs)iEuphab伯sp.(s);BuxusmjCmphy脂(|)

Cinnamomummacmpodum(s)iPa伽usnjPponjCus(f)

Mefasequoja9卯Ioslmbojdes(c);Pseudo伯亦am8M1S(Cs,sh,s),A;Que℃uSsub9en、Leadobajanus(Cu);COwy畑shefemphyIa(f);Qafaegussp.(e)

Me伯sequoj89卯fos1mbojdes(c),A;PCO/9onumsp(1)iCjmamomummaaopodum(s);H8m8me#spafmWojdes(e,s);W7siefaノリmbunda(b);S1y7axノapmjCa(s),CiSympbcossp.(e)

Sa9jff師asp.(0;Carexsecl・CarGx(f);C"xsp.(f)

朋創asequoja9”ostmbades(c,b),C;Hamamel1sp8ym(畑des(f,s),C;Wリs1evjajWonbunda(1,b)iWjssp.(s);SIy7axノapo、にa(s);Scjpuscf.〃ncojdes(f)

Ljqujdambarsp.(f)

Mef8sequoja9”oslmboldes(c);Sfew師jamonadeIpha(f);Ma9noj8sp,(s);BuxusmjCmphy脂(|)

KeIeleev7asp.(|);EufyaIesp.(s),C;MJph研sp.(s);Fafune誠asjnensjs(s);Sapjumsebねum(s);778pajncjSa(f);刀aPesp.(f)

PjCeasec1、Pjcea(Cs,|);Cafplnusfschonosk#(f);F89usmicmcafp8(Cu,f),C;Queにussecl,CemS(Cu);Siewa"jamonadeIpha(f);Buxusmjbmphyノノa(|);ReeveSiasp(1)

Cunnjn9hamjasp.(c,|);Sequojasp.(c,s,sh),C;KeIeノee"asp(c)iF89ussp(f);Que応ussub9en・CydobaノanOPsjS(Cu);

S伯wa"jamonadeIpha(1),C;Choemspondi8saxj伽s(s);Aノan9iumsp(e);C8rexsecI、Ca伯x(f);C副exsp.(f)

Sequoj8sp.(c),C;Piceasect、Picea(c);Cjmamomummacmpodum(s);Choevospond7asaxj"8応(e);A伯、9ルmsp.(e)

Me伯sequoリ8911/pros"obojdes(c),C;Sequojasp(c);Pice8sectPice8(c);Pjhussub9enpjPbxylon(c);PseudoIsu9a

submfund8(c);F89usmノcmcama(Cu);L剛jdambarsp(f),C;Choerospond1as8xノノノ8"s(e);Acersp.(f);万apasp(1)

GlyPIoSImbusPensj侭(c)

KeIe伯e"asp(c)iFa9uSmjcmcaノpa(Cu);MelリOSmasp(e);CarGxsP(f)

Sequojasp.(s,sh);Pseudola〃x8m8bjl1S(Cs)iSfewa巾amon8deIph8(f)i8uxusmjcmphy"a(|)

PseudoノarjXam8Mis(sh);SIewa"jamonadeIph8(f)

Meiasequoja9卵IosIrobojdes(c);Sequoj8sp.(c);Pjce8kofib8i(c);Pjce8sect,Pjcea(|)iPseudoIa欧amabj脂(Cs,|);Fa9us

mIaocama(Cu),A;Quefcusa〃ena(Cu);Que「cussectCe"js(Cu)iSfew8巾8mon8delpha(f),C;Ljqujdambarsp.(f),C;Buxus

mjcmPM8(|);SfW8xj8Ponjca(s)

Sequoj8sp.(c);BuxusmicmpMa(|)

Pにeasec1、Pにea(|,sh),A;ん9ノansme9acjnerea(f);QueYCussec1,CemS(Cu);BuxusmjcrQPhy"a(|)

Sequoj8sp.(c);Pj℃easec1‘PソCe8(c)iPseudofsu9asubvDIunda(c);F89usmicmcaノp8(Cu);Qu研cussub9en、Cems(Cu);SiewarWamonadelpha(0;Forlunea7j8sjhensjS(s)

Pj℃eakajbaj(C)iPjceasectPjCea(1,Cs);Carpmussp.(f);Fa9ussp.(f);SIewa巾8monadeIPha(f),A;FomJnea"asjnensjs(s);StW8x/aPOnjca(S)

GIypfoslmbuspensjl/s(c,sh);Sequojasp.(s)

Pseudo1am(amabjlVs(Cs,sh),CiQuerCussectCems(Cu);CarpjnusIschonoskjj(f);Sfewa巾amonadelPha(f),A;Fomノneana

sjnensjs(e,s),A;l-jqujdambarsp.(f);BuxusmjcrqpM8(1)iV"ssp.(s)iSympjocossp.(e)

Sequojasp.(c);Fa9usmjcmca巾a(Cu);Quevcussub9enCycloba伯nopsjs(Cu);QuefcussectCemS(Cu);Quef℃ussp.(f);

AC伽djanノ畑(s);EuIyasp(s);S伯w師jamonadeIpha(f);Fof7une師8sjnensjs(f,s);Zan1hoXyルmsp.(s);Vj1jssP(s);StWaxノ8ponjca(s)

MeI8sequoia91I/pIosfmbojdes(c);耐jwanjacJypIom研jOjdes(c);Pseudoノa"xamabj#s(Cs,sh);Pfefoca/yasIenopfeIa(1),C;

M89noIj8〆aecocjss(s);Acercf・mono(f);Aceysp.(f);La9efWoemjasp.(f);ComusconIroversa(e)iSympIocossp.(e)

Me脂azedarach(e);SMaxノョponjca(s)

MeIasequoja91ソpIostmbojdes(c)iMa9no#asp.(s),C;AC伽。旧sp(s)iSIewaノW8mon8delPh8(1)iW7sfe"a"onbunda(b);

S8piumseMbrum(s);Zanfhoxy山majノanIhojdes(s);Me#88zed釘8ch(e)iS8bj8japonにa(e);AmpelbPsjSsp(s);S肋axj8ponjc8(s)

Ma9nO"asp(s)

Sequoj8sp.(s);PjceasecI,P1Cea(c,|);PseudbIam(amabjルs(sh);Quer℃usa"ena(Cu)iSIewamamonadeIpha(f);StyvaxノapOnjca(s)

Pjceasec1PjCea(|);Fa9usmJcmcaノpa(Cu);QueYcussectCef7js(Cu)

MeI8sequoja9”osfrobojdes(c)iPseudob"xamaMjs(sh);QuefcusseclCems(Cu)iSIew銅jamonadelbha(f)‘C;

Ljqujd8mbarsp(0;AIeurjlescovdaIa(s);Me#osmasp.(e)iMelWodendmnxylbc6のum(1)

34.23‘26.N,134.51'49.E60

34.23‘050N,134.5228。E30

34.21.31.N‘134.5227.E60

34.24'21。N,134.46148oE30

34.23009W,134.46159。E50

34.230040N1134.461589E55

789012

333444

PS

SMM

34.23.00口N,134.47111"E40

34.21033口N,134.47'O1nE50

34、22‘33。N,134.48138"E85

34.21‘590N,134.48114。E50

34.22‘53.N,134.45'49.E1

34.2255。N,134.45152口E1

34.21‘12W,134・45130.E50

34.21136.N,134.45151.E25

個糾妬垢岬杷相印

MMSMSSMS

5134.2042.N,134・45028QE50M

7134.11‘49.N,134.44.37・E25S

7234・11‘25.N,134.44'01.E10S

SSS

6234.19‘26qN‘134.54014.E45

6334.19‘250N’134.54‘23.E45

6434.19‘30。N,134.54135・E40

58a34.18122.N,134.52‘08.E100M

58b34.18122W’134.52'O8qE100M

58c34・18'22W,134.52'08.E100S

58.34.18122.N,134.52IO8QE100

58e34・18'22.N,134.52'O8nE100

5934.18'32.N,134.52'09口E63

6034.18'41.N,134.52.20oE80

6134.18'55.N1134.52.33・E40

PSSMS

69a34.12‘13.N,134.48153.E5M

69b34、1213。N,134.48153pE5M

69c34.12.13.N,134.48'53.E5S

7034.12‘16.N‘134.44‘28・E30M

6534.1928nN,134.54'38.E40S

6634.19‘30.N‘134.54‘39.E40M

67a34・1755qN,134.56lO2oE20M

67b34.1755qN,134.56‘02.E20S

6834.12‘35.N’134.4916.E60M

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58 jpn.』、Hisl()r、B()t、 Vol、6.No.2

FukudaVolcanicAshLayer(Mizun(),1992).Withthe

KobiwakoGroup,theKenjogaokalVolcanicAsh

LayeriscorrelatedwiththeHazamaVolcanicAsh

Layer(Mizunoe/α/,,inpress).

Materialsandmethods

Wefound89plantmacrofossilassemblagesat72

1ocalities(Figs、1,2,Tablel).Atlocalitieswithout

volcanickeybeds,wecorrelatedplantmacrofossil

assemblageswiththestandardcolumnarsectionof

eachareabasedonthecontinuityoflithofaciesand

thelocalgeologicalstructure

Mostplantmacrofossilswerecollecteddirectly

fromtheoutcrops・Asformacrofossilbearingbeds

withoutmacroscopicplantparts(fossilassemblaRes

8a24and27),plantmacrofossilswereobtainedby

sievingofsediments、Wedidnotsievesedimentsat

mostotherfossilbeal-ingbeds、becausethisstudy

mainlyaimedtoclarifyarborealplantfossilsthatcan

beobviouslyrecogllizedattheoutcrops,Plantma、

crofossilsarepreservedin70%alcoholandarede‐

positedintheFacultyofHorticulture,ChibaUniver‐

sity.

ResultS

Stratigraphicoccurrenceofplallts

From89fossilassemblages,l4coniferousand70

angiospermoustaxahavebeen()btained(Tablesl,2).

Amongthe84taxaobtained,26taxa(31%)areextinct

fromJapan、Theyarec()mmollillthehorizonlower

thantheSak()V()lcanicAshLayerilltheAtago

FormationandinthcYudaniFormation(Fig.3).

C〃09ノりSjD0"d加s“"伽γisandR“zwasp、areconfined

inthelowerpartoftheAtag()F()rmation.RW""g”‐

〃s伽g"s応,Lj9"/血"z6aノ′sp.,〃e"/0火"。”〃xzyA〕cαγ・

か"ノ2,K碓睦γ〃sp.、andRZg"S〃"c'りcαゆaoccuronly

inthehorizonlowerthantheSakoVolcanicAsh

Layer.Sセ(/"0〃sp.,ハe"do〃γjixaノw6"is,andG“/0s‐

/ノ℃6ノ応Pg"s"jscommoninthemiddlepartofthe

AtagoFormationdisappearedintheupperpartofthe

sameformation、InhorizonsupperthalltheSako

VolcanicAshLayer,ノら〃7"?"ノ"eノis/)"γノ'イノノm/“s,P加"s

subgen・HZZ/)ノ0ノWノ0"、A〃7"sノ”イノ"/“,〃je"0咋""うり〃

α"zI"ぜ"sg,CW/OL.αノyaPα""γ"s,alldCo7ツノ"sルc/eノ”ノIリノノー

〃appear,and〃e/(zW"OmglIy/)/0s/γYノ6(ノノ“s()ccurs

morecommonlythaninthelowerhorizon,Fossil

assemblagesintheGoshikihamaFormationinclude

/W(zse9"0z〃g卯/0sfγ060〃es,C加"α"zo"?""z〃Iα‐

cノ”0ロルノ"z,andHz"2α"ze"Spaγ”"0〃Cs・

FossilassemblageintheYudaniFormationincludes

plantsextinctfromJapan,suchasnIMWi"ノα

c”/0"ze”0雄s,〃e"iO此"めり〃xy〃αzゆ"》泌踊e"〔ノOA7ノ'‐

ぱα"za6"応,and艶9"0〃sp.、anditscompositionis

similartothoseinthemiddleandlowerpartsofthe

AtagoFormation.

Sedimentaryfaciesoffossilbearingbeds

Mostplantmacrofossilassemblagesareincludedin

thesandyandsiltysedimentsofthealternatingbeds

ofsiltandsand,butnotinthegravellayers、The

sedimentaryfaciesof82plantmacrofossilassem‐

blagescanbeclassifiedintothreetypes(Table2).

Thefirsttypeispeatlayerswithhighorganic

contentsandincludesthreefossilassemblages(18,37,

and58d).Thesepeatlayersaremadeofun,

decomposedplantpartsincludingroots,shoots,and

reproductiveorgansofherbsandarbors・Thefossil

bearingbedsofassemblagesl8and58darewo()dy

peatofcompactedleavesandshootsofG卯/0sかり6"s

le”s"fs,andoverliemassivesiltconformably、The

organiccontentsofthesedimentsincreasegradually

fromthemassivesilttothewoodypeat、

Thesecondtypeismassivesiltlayersoftenshowing

horizontalcontinuityandincludes43fossilassem‐

blages・Inthisfaciestype,plantremainsarescattered

uniformlyinthesiltandareverypoorlysorted

accordingtotheirsize・Themassivesiltlayersnear

thebasementrocksaresometimesill-sortedaI1d

includesandparticles、

Thethirdtypeisplantfossillayersincludedin

stratifiedsandysedimentsandincludes36plantma‐

crofossilassemblages・Fossilremainsandsediments

arewell-sortedaccordingtothesize:largerfruitsand

woodsincoarsersands,andleavesandsmallerseeds

infinerones.

Sedimentaryfaciesandtheoccurrenceofplallt

taxa

Weselectedl7taxaincludedinmorethansix

assemblagestoclarifycorrelationbetweenthesedi、

mentaryfaciesandtheoccurrenceofplanttaxa

(Table3).Basedonthesedimentaryfacies,l7taxa

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644幅

IIabitatofPlalltsintheLatePli()c(pncSc(Iim()I11aryBasiI1(Momohara&Mizullo)

28

Table30ccurl・eI1ceofselectedplanttaxainthl・eesedimentarvfaciestypes・Nymphaeaceaeincludesヴ

Bノ枢Scノノル7sc"ノゼ6cノ,E"ノフノ"/(′sp.,andノVノゆノノ”sl〕、ノ)/“asect・P7”αiI1cludesf〕ノヒ・”ル0ノ6"/,

alluvialplain(wetland-riverbank)

alluvialplain(wetland-riverbank)

alluvialplain(wetland-riverbank)

alluvialplain(wetland-riverbank)

alluvialplain(wetland-riverbank)

alluvialplain(wetland-riverbank)

alluvialplain(wetland-riverbank)

alluvialplain(wetIand-riverbank)

59

sedimentarvfaciesoffbssilbearin反beds presumedhabitattaxa

eat(3) massivesilt(43)stratifiedsand(36)

Discussion

C⑪mparis()nOfbiostratigraphyin&mdar⑪und

Aw&ljiIslaI1d

Thestratigraphic()ccurrence()fextillctlaxafr()m

Jflpan()nAwajilslandissimilartothatintheSellI1an

area・IntheSennanarea,Reez'ESノ〃sp・andCノノOeノりV)イノル

(ノル/Sax"ル〃八()ccuronlyinthebasalh()rizon()flhc

OsakaGrouplowerthantheTsuchimarulVolcanic

AshLayer(M()mohara,1992).勘γ/""“γ/asノノノcノバノs

andKc/e/egγmsp・becameextinctbelowthcMizuma

llV()lcanicAshLayer,andL〃ノ"”ノノめαノ.sp、,ル化/,

/加咋"めりノ?sp・andSセ9"o血zsp,disappearedataboul

thellabutakillVolcanicAshLayel。.必fw〔ルノ〃ノ"次

alluvialplain(swamp)

alluvialplain(swamp)

alluvialplain(swamp)

alluvialplain(swamp-riverbank)

G{しprostmbUspe"sノルS

Nymphaeaceae

万百pa

Stw召x〃poノフノCa

2221

αノノ?α6"たbecameextinctatab()uttheShilm()ikeVol,

callicAshLayer,InthehorizollsbetweentheShinn()i‐

keVolcanicAshLayerandFukudaV()lcanicAsh

Layer,G帥/0s/ノり6zfs”zsIWS,〃e"w"(ノルzgl1WOs‐

/ノ'イノルィノノ咋s・P/“〃ル0ノ花α/,C加""ノ"0"z""γノ"(/〔・ノ”(ノ。/"",

andノヵノ"αノ"c/jsPαγ”"0,i火αoccurcomm()llly(M()m0.

hara,1992).AccordingtothecorrelationofHabutaki

ll,Asashiro,Shinnoike,andFukudaV()lcanicAsh

LayerswithvolcanicashesonAwajilsland(Fig.3),

KC/e/“ノ'msp・and艶9"oねsp・lingeredoninmore

y()ungeragesonAwajilslandthanintheSennanarea、

TheYudaniFormationcanbecorrelatedwiththe

basalhorizonoftheOsakaGroupintheSennanarea,

Because71fwWノ""”cソayP/0"ノcノ()/火s()ccul・sinf()ssil

asscmblage68intheupperpal・toftheYudaniForma.

ti()ninareal〕(Fig.2).nziz{w"〃cノ”/()ノ"cγ/0/此sis

c()I1Iilledbel()wTsuchimarulV()lcanicAshLaycrill

thebasalh()rizonoftheOsakaGroupilltheSennan

area、The()ccurrence()fL町"/的""6αノ′sp・andルグcノー

ノノp〔/(抑(ノノ℃〃Xy/0(Pαゆ""lintheYudaniF()1・matiollindi‐

catescorrelationwithhorizonsbelowSakoVolcanic

AshLayerintheAtagoFormati()、.

areclassifiedintothreegroups・Thefirstgroupis

c()mp()sedofG“/0s/ノ'イノ伽s”澱"ん,Nymphaeaceae,

7)W/)",andSノ(ymxノ〃イノノ"Ca()ccurringinI)catillld

massivesilt・Thesecondgroupconsistsoftaxac()、.

m()l1inmassivesiltlayers,i、e、,ノWmW"Cmg4の/イノs‐

/γ(ノ6(ノノ〔/9s,s/“〔!αγ"αノ"0"αde伽α,S“"0ノasp.,

除(’"dM7ノな〃ノノノα6"is,Q"gノ℃"sα"g"α,I叩耐/”/MIO"6""‐

(ん,ノノ"ノ"αノ"cノパノウαγノり//0/火s,andCノW"(7ノ"イノノ""ノ〃〃/".

c”Pイノ〔/""z・ThethirdgroupincludesP7r“sect./γ(w,

βノmfsノ"icノ”ノM〃,RZg"s〃"cノり“ノアα,Q"fノで"ssect・

Ce〃八,andLノりノ"”ノノノ6"ノ′sp.,occurringc()mm()nlyin

stratifiedsandbutrarelyinmassivesilt.

SedimentarVenvirOnment0fplantmacrofossil

a昌烏emblages

TheOsakaGrouponAwajilslandhasthesedimen.

/1作曲sequoおg〃pオostmbo雄s

Ste順mbmoノ7adbノID/7a

s唇qUo心sp・

PSごUdbルa/1iAramabノルぢ

QUe”usa〃台"a

州Ste砲ノヲbノ7b4"7Ob

hIamame"Spaノ、加/ぴちS

c/h"aノフ7Omumノ77aCmpOdt"77

43196554

111

29916412

月beasect、月bea

Buwsm/bmpノ7j"Ma

F白gUsmノbmcaノフoaQueノ℃Ussect.CちmS

L々Uノ乱ambarsp.

24222

幅加864

fanandmountainslope

fanandmountainslope

fanandmountainslope

fanandmountainslope

fanandmountainslope

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60 Jpll.』、IIistor.B()!. Vol、6.No.2

taryfaciesofanalluvialplainorafan・Thealterllat‐

ingbedsofsiltandsanddominatingintheOsaka

Groupofthisislandshowsedimcntaryfacies()fan

alluvialplainwheresandymeallderingchalmelsal・e

developing(Walker,1984).Theoften-stratifiedsandy

sedimentsarechalmel-filldepositsorflooddeposits

overbanks・Massivesiltyandclayeysedimentsare

backmarshdep()sitsofrivers,Gravellayerscharac‐

teristicallydepositinafanoralongalargeriver、The

gravellayersintheAtago,Toshima,andYudani

Formationsconsistofpoorly-sortedsubangularor

subroundedgravelwithoccasionaldiscontinuoussilty

layers,andtheircompositionisgreatlyinfluenccdby

thebasementrocksaroundthesedimentarybasin

(Mizuno,1993).Thesecharacteristicsofthegravel

depositscorrespondtothoseofthedistalendofafan

wherebraidedchannelsaredeveloping・Onthecon‐

trary,thegravellayersintheGoshikihamaF()rmati()n

arecomposedofwell-sortedgravelandcommonly

includecrystallineschistnotfoundinthepresent

basementrocksonAwajilsland(Mizuno,1993).This

schistisderivedfromareasoftheSanbagawa

metamorphicbeltnowdistributedalongtheKinok‐

awaRiverinWakayamaPrefectureandtheYoshino

RiverinTokushimaPrefecture・InthelatePli()cene,

Sanbagawaschistwasprobablyexposedonlandto

thesouthofthepresentAwajilsland・Thesedimen.

taryfaciesandtheexistenceoflong-transported

RravelintheGoshikihamaFormationindicatedeposi、

tioninalargeriversystem、Plantmacrofossilassem‐

blagesincludedinthealtematingbedsofsand,siltand

claywerethusdepositedinanalluvialplainoratthe

distalendofafan、

Amongthethreetypesofsedimentaryfaciesof

plantbearingbeds,theundecomposedpeatisusually

formedinwaterloggedenvironmentsuchasfenand

bogs・Thehighcarboncontentsandthelackofcoar‐

sersandparticlestransportedbyfloodingwalerinthe

matrixindicatespeatformatiollfromplantsgr()winR

insituwithoccasionalinclusi()I1offloatingall()chth、

onousfruitsandseeds・Plantmacrofossilassemblages

inthepeatcanthusbeinterpretedasbackmarsh

depositsandautochthonousorigins・

Plantmacrofossilassemblagesinmassivesiltwere

alsodepositedinthebackmarshofariver、Thefiner

matrixwith()utstratificationandtheuniformdistribu.

tion()ff()ssilsinthesedimentsreflectdepositionunder

agentlefl()w()rwith()utawaterflow・Sizeofplant

remaillstransportedinwaterisc()rl・elatedwellwith

thegrainsizeofthcmatrix(M()m()hara&Yoshi‐

kawa,1997).Apool。c()rrelationbetweenfossilsize

andtheRrainsizeofthematrixcomposingthemas‐

sivesiltshowsthatmostplantpartswerenottran.

sl〕()1.1cdinwater,butwerefloateddownoverthe

watel・Surface・Insuchacasemostplantremainsa1・e

derivedfromplantsIg【rowingaroundtheplaceof

depositioninthebackmarsh・Fossilassemblagesin

themassivesiltshouldthusreflectpale()vegetationon

wetlandsandriverbanksinanalluvialplaill、

Onthecontrary,plantfossillayerswithinstratified

sandweredepositedullderahigh-energywaterflow・

Stratifiedsandcomp()seschannel-filldepositsor

flooddepositsoverbanks、Plantfossilsinthesefossil

layc】・sarem()stlytransportedfromtheupstl・eam

watershed・Inchannelfilldeposits,sizeandshapeof

plantremainsal・ecorrelatedwellwiththegrainsize

ofthesedimentmatrix(Momohara&Yoshikawa,

1997).Fossilassemblagesofthistypeshouldthus

reflectregionalpaleovegetationincludingfansand

mountainslopesal()nganalluvialplain.

1,lanthabitatinthelatePli⑪cenesedimentary

ba烏in

Taxafoundinthepeatprobablygrewinthemost

aque()usenvironment(Table3).Theabundantoccur‐

rence()fshootsandconesofG卯/0sノノ℃6"stogether

withn”αandNymphaeaceaeilldicatesthatG帥/(パ.

〃7ノカ"sjbe"s"おwereRrowinginshallowwater.

G卯/()s/ノ℃6"s/)g"s"八isnowdistributedinswampy

areaso[adeltainsouthChinaanddevelopsrespira‐

toryr()()tsabovethewatersurface(Fu,1992).Afew

S(yノ“seedsinapeatlayerwithG卯/0s/、6"swere

pr()bablyderivedfr()mdrierareassuchasriverbanks,

becausetheyfloatwell、

Taxacommoninthemassivesiltmusthavegrown

inandaroundthebackmarshwheretheirseeds,fruits,

orleavescouldeasilybetransportedintothearea()f

dcp()siti()nbyaReI1tlcstreamorwind‘Sandbanksor

m()undsandwetlalldsar()ulldtheminterspersedinan

alluvialplainpl・()bably()fferedhabitatforthese

plants、Fruitsandseeds()fplantsgrowinRinthese

arcas,whichweresusceptiblet()fl()()d,mustalsohave

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IIabitalofPlantsintheLatel)lio“neSedimentaryBasin(Momohara&Mizuno) 61

beentransportedbyfloodingwaterandweredeP()sit‐

edalsoinstratifiedsand(Table3).Amongtheplants

inthemassivesilt(Table3),Slj'うてzx”0"たα,/'ん/砥e‐

り"(ノルzg卯/0s/パノ6(ノ〃ES,鈍り"0”sp.,andWHs/eノかβ()ノ'.

/6""血arenowgrowinginandaroulldthewetland.

Sl(WYzx"0"ノmandWXs/”血ルノ茄""dhIareveryc()m‐

mollaroundthewetlandinalluvialplainsinJapan、

Thchabitat()f/was“"(ノ〃g妙/Cs/灯ノル(ノ〃csins()uth‐

westChinaisc()nfinedtothewetlandinvalleyb()t、

toms(Chu&Cooper,1950:Momoharae/α/、,1993),

andS”"omse"ゆeγzノi""sisgrowinginthewetplaces

alongriversinthewestcoastofNorthAmerica(Li,

1953).Ontheotherhand,S陀肌'α″〃ノ"0"α(1/e〃MIand

Q""℃Jfsα/花ノノaarenowdistributedinmountainsand

hillsinJapan,and庇e"伽ルノγfxα”α6"isisdistributed

inhillyregionsincludinglimestoneareasincentral

andsouthChina(Fu,1992:Momoharae/α/、,1993).

Thehabitat()fthesethreetaxaextendedtoalluvial

plainsinthePliocene,andisdifferentfromtheir

presentdistribution、ExtinctspeciesofC/"〃α"70ノノノ"ノ〃

〃ノαcツ”0α""ZandHZz"zα"Ze/応Pαγ?り"0/〔んsalsogrewin

alluvialplains.

I)lalltsc()mmoninthestratifiedsandbutrareillthe

massivesiltpr()bablygrewonfansandmountain

slopesfarfromtheareaofdepositi()'’inanalluvial

plain・Pi“〃sect.Pice",Bノ“‘s〃成洲ノβノMルz,R"ILJ"s

""C”cαゆα,Q"g〃"ssect・Ceγ》た,andL伽姉ノ"6αγsl).

mainlygrewonfansandmountainslopes・These

plantsmayhavegrownalsoinanalluvialplain,

becausetheyareincludedinthemassivesilt・Their

rareoccurrenceinthemassivesiltmay,however,

indicateoccasionalinclusionoffloatingplantremains

afterfloods・ノ〕/“asect.P/cgaisnowdistril)uted

mainlyinr()ckyareasineastAsiae、質.,Pk〃α)/0ノ′in

centralllonshu(Noteg/α/,,1998).P/“〃sect./¥〔、“is

alsodistributedinwetlandsinanalluvialplain,e、g、,

P/“〃g/eル"〃illH()kkaid()(Note2/αノ.,1998).F()ssils

()fノ)/“〃sect.P〃eaoccurredfrommassivesiltinthe

OsakaGroupalldwasassumedtohavebeenac()mp().

nent()fforestsinanalluvialplainincoolerstaIges()f

thel)Ieistocene(Momohara,1992).円“asect.〃“a0n

Awajilslandpreferredfanandmountainslopesinthe

Plioceneratherthananalluvialplainasitshabitat.

I〕lanthabitatassumedinthispaperisconsistent

withthatintheNeogeneAizusedimentarybasin

clarifiedbySuzuki(1976).Suzuki(1976)examiI1edthe

speciescompositionofplantmacrofossilassemblages

andtheirspacialandtemporaldistributioninalate

MioceneandPliocenesedimentarybasininAizu

PrefecturenorthernJapan,Hec()nsideredcoexis,

tellceofspeciesinfossilassemblagesandthepresent

habitatoffossilspeciesandtheirmodernrelativesas

iml)()rtantcriteriatoreconstructpastplantcommu・

llitics、InthePliocenesedimentarybasin,G帥/(ノs‐

/ノヤノル"sinhabitedthemostaqueousenvironmentwith

A/""sノ“oノ"caandpルパzgノ""ES,〃gい”"o〃、Sb'ノYzx,

βノ“"s,andS彫拠'αγ/”grewinthedrierhabitatinthe

l()wland,andP/Ceαル0γめα/,Rsg"”A1Iγな、andRIg"s

〃"Lwノcαゆaoccupiedmountainslopes(Suzuki,1976).

Amongthem,habitatsofG卯/0s"6"s,ルィ”se9"0ノα,

Sノノ:Wzzx,S/gⅨ'”/〃,piCeaル0γ必α/,andF地ノハ〃zノα℃“ゆa

clarifiedbySuzuki(1976)arecomparablewiththose

assumedinthispaper.

Ackllowledgements

ThisworkwaspartlysupportedbyaGrant-in-Aid

forScienceResearch(No.10680097)fromtheMinistry

()fEducationScienceandCulture,Japan.

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(AcceptedonJanuaryl4,1999)

書評(新刊紹介):日本林業技術協会編.1999.森林の環境100不思議.218pp・日本林業技術協会,東京.本体

1300円.

本書は,森林の環境に関心をもつ人だけでなく,地球

人として森林と隙境のかかわりに関心をもってもらいた

いあらゆる人を対・象に執筆・編集されている。森林総合

研究所を中心にした中曜・若手の研究者が執筆にあたっ

ている。1項112ページで100項目200ページ,それぞ

れの項目を得意とする研究者が担当している。それに加

えて,平易なです。ます体で語られ,項目ごとに親しみ

やすい題目が付けられているので,読者の興味のあると

ころから読みはじめることができる。

そうは言っても,ただ項目の羅列だけなら辞背同然で

あるが,本書はそうではない。環境問題と森林,気候変

動と森林,日本の森林の変遷と現状,森林が守る私たち

の生活環境の4つのおおきな部分に分かれており,ス

トーリー性を失っていない。そのストーリー性とは,今

日の地球蝋境問題を理解するための緑の生態系,’41然・

人類の歴史における緑の生態系,現代社会史における緑

の生態系といった具合である。

編集・発行にあたっている林業技術協会,執筆者の大

半が所属する森林総合研究所とは,いったいどういうこ

とをしているところなのか。それをアピールする目的も

本書にはあるようだ。そういう視点も忘れずに読んでみ

ると,一隅おもしろくなるかも知れない。

(辻誠一郎)