reptile family tree - peters 2018 1321 taxa, 231 characters · buriolestes ulbra-pvt280...
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SilvanerpetonGephyrostegeus bohemicus
TijubinaBavarisaurus (= Homoeosaurus) macrodactylusHuehuecuetzpalli
Macrocnemus BES SC 111
LitorosuchusLPV 30280 w/embryo
Macrocnemus T2472Dinocephalosaurus
Langobardisaurus
TanytrachelosTriopticus
Tanystropheus k and q
CosesaurusKyrgyzsaurusSharovipteryxLongisquama
Schoenesmahl (formerly Bavarisaurus)FraxinisauraLacertulus
Daohugou lizard IVPP V13747
Meyasaurus
Carusia
Hoyalacerta
Homoeosaurus solnhofensisIPB R 535 Homoeosaurus tail
IVPP V14386Dalinghosaurus
Eldeceeon 1990.7.1Eldeceeon holotype
Tulerpeton
WestlothianaCasineriaBrouffia
PaleothyrisCoelostegus
Hylonomus
Protorothyris MCZ1532
EudibamusAphelosaurusPetrolacosaurusAraeoscelisSpinoaequalisGalesphyrusYoungina BPI 3859AcerosodontosaurusThadeosaurusTangasaurusHovasaurusClaudiosaurusAdelosaurusAtopodentatus
LargocephalosaurusOmphalosaurus
Sinosaurosphargis
Anarosaurus
Youngina SAM K7710
Youngina TM 3603Youngopsis rubidgei RC90Youngopsis kitchingi TM1490Youngoides RC91Youngina AMNH 5561Youngoides UC1528
Proterosuchus BPI 1 4016
Chasmatosaurus NMQR 1484Teyujagua
Proterosuchus RC96
Euparkeria SAM PK K5867Euparkeria SAM PK K6047A
OsmolskinaTasmaniosaurusRevueltosaurusFugusuchusGarjainiaErythrosuchusLuperosuchusShansisuchus
RiojasuchusOrnithosuchus
VjushkoviaPrestosuchusBatrachotomusSaurosuchus
SmokTeratosaurusSikannisuchusPostosuchus
DecuriasuchusPagosvenatorTurfanosuchusPoposaurusSacisaurusSilesaurusLotosaurusShuvosaurusEffigia
Protorothyris CM 8617Protorothyris MCZ 2149Vaughnictis
MFSN 19235Scandensia
ArchaeovenatorAnningia
OrovenatorPyoziaBroomiaMilleropsis BPI-720Erpetonyx
Ascendonanus MNC-TA0924
Rauisuchus
PVL 4597
Lewisuchus
TarjadiaParringtonia
GracilisuchusMCZ4116SaltopusScleromochlus
TeleocraterArizonasaurus
YarasuchusQianosuchusNundasuchusTicinosuchusAeotosauroidesAetosaurusStagonolepis
Wumengosaurus
ThaisaurusXinminosaurusUtatsusaurusShastasaurus alexandraeShastasaurus pacificus UCMP 9017GrippiaMacgowania
MikadocephalusYGMIRSPCV03108ParvinatatorChaohusaurusBesanosaurusQianichthyosaurus CMNH V1412Qianichthyosaurus IVPP V11839Phalarodon fraasiContectopalatus'Phalarodon' nordenskioeldiiMixosaurusCymbospondylus petrinusShonisaurus sikanniensisGuanlingsaurus GNG dq-50Guanlingsaurus YGMR SPC V03107
WimaniusPlatypterygiusGuizhouichthyosaurus tangae IVPP V 11853'Cymbospondylus' buchseriShonisaurus popularisIchthyosaurus communisOphthalmosaurusLeptonectes tenuirostrisHauffiopteryxBRLSI M1399
Suevoleviathan GPIT 328/4/5Suevoleviathan SMNS15390
Eurhinosaurus
Chasmatosaurus QR 880C.500Chasmatosaurus SAM PK 11208Chasmatosaurus TM 201
Elaphrosuchus RC59
DiandongosuchusTeraterpetonMesorhinosuchusPaleorhinusParasuchusChanaresuchusProterochampsa PVSJ 77Proterochampsa PVSJ606Pseudochampsa PVSJ 567GualosuchusCerritosaurusTropidosuchus PVL 4601Tropidosuchus PVL 4602Lagerpeton
Bergamodactylus MPUM 6009EudimorphodonPreondactyluisDimorphodon
Nyctiphruretus holotypeNycteroleter
Nyctiphruretus PIN 4659/1
BarasaurusKitchingnathusSantaisaurusSauropareionColettaPaliguanaSophineta
PsilotrachelosaurusPIMUZ AIII 0192SMF R 4710StereosternumBrazilosaurus holotypeMesosaurusSerpianosaurusConcavispinaXinpusaurus kohiXinpusaurus suniThalattosaurusWachtlerosaurusEndennasaurusClarazia
AskeptosaurusAnshunsaurusMiodentosaurus
Eusaurosphargis holotypeHelveticosaurus
Vancleavea
DiandongosaurusPalatodontaPappochelysPalacrodonMajiashanosaurusParaplacodus
Placodus gigasHenodusPlacochelysCyamodusPachypleurosaurus
CartorhynchusSclerocormus
Qianxisaurus
KeichousaurusDianmeisaurusDawazisaurusHanosaurusLariosaurusCorosaurusNothosaurusWangosaurus
SimosaurusAnningsauraPistosaurusYunguisaurus
AlbertonectesPlesiosaurus
TrinacromerumBrachaucheniusKronosaurus
Proterosuchus SAMPKK10603DoswelliaYoungina BPI 2871ElachistosuchusTchoiriaIkechosaurusChampsosaurusSimoedosaurusCteniogenysPhilydrosaurusLazarussuchusHyphalosaurusMonjurosuchus GMV216Monjurosuchus NMM3671
SMNS12591skullTerrestrisuchus
LitargosuchusErpetosuchus
HesperosuchusSMNS12352 partial skull
PedeticosaurusYonghesuchusDromicosuchusSichuanosuchusProtosuchusSphenosuchusSaltoposuchusDibothrosuchus
DakosaurusTeleosaurusPelagosaurusDyoplax
MetriorhynchusSebecusIsisfordiaCrocodylus niloticusAraripesuchusSimosuchus
MariliasuchusArmadillosuchus
CaipirasuchusBaurusuchus
Herrerasaurus
TchingisaurusChometokadmonGekko smithiiLialisGekko geckoEublepharisNorelliusArdeosaurusEichstaettisaurus schroederiEichstaettisaurus gouldiJucaraseps
AphanizocnemusAdriosaurusPontosaurusPrimitivus
TetrapodophisDinilysiaPachyrhachisBoaCrotalusLoxocemus
Yabeinosaurus IVPP V13284
XenopeltisAniliusCylindrophisUropeltisAnomochilusLeptotyphlops
Bahndwivici FMNH PR2260
AigialosaurusTethysaurusTylosaurusPlotosaurus
Not Ascendonanus MNC TA1045
Saniwa ensidens FMNHEstesiaVaranus komodensisVaranus griseus
ShinisaurusHelodermaLanthanotusAnniellaGobiderma
OphisaurusKuroyuriella
MyrmecodaptriaElgaria
Cryptolacerta
Chalcides ocellatusChalcides guentheriGymnophthalmusVanzosauraSirenoscincusSlavoia MGR-I-112SineoamphisbaenaCrythiosaurusSpathorhynchus
DibamusTamaulipasaurus
BipesAmphisbaena
BarberenasuchusEodromaeus
Sakurasaurus
LimnoscelisOrobatesTseajaiaTetraceratopsMilleretta RC14Milleretta RC70
BashkyroleterMacroleterEmeroleterLanthanosuchus
UrumqiaBruktererpetonThuringothyris MNG 7729Thuringothyris MNG 10183
Reiszorhinus
CephalerpetonOpisthodontosaurus
Concordia KUVP 8702aConcordia KUVP 96/95Romeria primus
Saurorictus
Romeria texanaProtocaptorhinusParacaptorhinusEocaptorhinusCaptorhinusLabidosaurus
Calanguban
Acanthodactylus
EolacertaLacerta
TupinambisDorsetisaurus
Slavoia holotype ZPAL MgR-I/8Slavoia ZPAL MgR-III/77MacrocephalosaurusTianyusaurus
Diadectes CM25471Silvadectes absitus MNG 8853PhonodusBolosaurusBelebeyOradectes sanmiguelensisDiadectes UC706Diadectes sideropelicusProcolophonKapesScolaparia NSM 99 GF82.1LeptopleuronPentaedrusaurusHypsognathus
Youngina BPI 375Prolacerta AMNH 9502IxalerpetonCzatkowiellaMalerisaurusProlacerta BPI 1/471OzimekProtorosaurus NHMW 1943I4*BoreopriceaJaxtasuchusProtorosaurus NMK S 180Protorosaurus WMsN P47361Pamelaria
ZupaysaurusMegapnosaurus
StephanospondylusProvelosaurusDeltavjatia KPM232ScutosaurusBradysaurus NHMWAnthodonPareiasuchusPumiliopareia
JZC Bu1803 - amberLiushusaurusPurbicella
EuposaurusIguana
TriocerosPhrynosoma
BasiliscusAnolis
Pristidactylus
Chlamydosaurus
JKZ Bu267 -amber
Tawa
Staurikosaurus
SegisaurusZuolong
MarasuchusProcompsognathus
Dracoraptor
CoelophysisDilophosaurus
Sinocalliopteryx
Buriolestes CAPPA/UFSM 0035
VelociraptorBalaurHaplocheirusShuvuuiaHalszkaraptorSinornithoidesSauronithoides
Jinfengopteryx Anchiornis LPM B00169 Anchiornis PKUP V1068Anchiornis IVPP V14378
AurornisCaihongBuitreraptorSinovenator
Eosinopteryx
Xiaotingia
ElliotsmithiaApsisaurusAerosaurusVaranopsVaranodon
Varanosaurus FMNH PR 1760Varanosaurus BSPHM 1901 XV20ArchaeothyrisOphiacodon
HaptodusPantelosaurus
SecodontosaurusDimetrodonSphenacodonIanthodonEdaphosaurusIanthasaurusGlaucosaurus
CutleriaStenocybusIVPP V18117KenyasaurusGalechirusSuminiaVenjukoviaEodicynodonDicynodon
HeleosaurusMycterosaurus FMNH UC169NikkasaurusMesenosaurus PIN 158/1Niaftasuchus
Beipiaosaurus
RahonavisJianchangosaurusLimusaurusCaudipteryxKhaan
JuravenatorSinosauropteryx
Compsognathus holotypeAorunStruthiomimusGallimimus
OrnitholestesScriurumimusMicroraptor IVPP V 13352Microraptor M V1002Sinornithosaurus holotypeSinornithosaurus NGMC 91
FukuivenatorMasiakasaurusTianyuraptorHuaxiagnathusZhenyuanlongAlioramus altai
GorgosaurusRaptorex
Tyrannosaurus
Reptile Family Tree - Peters 2018 1326 taxa, 231 characters
EoraptorPampadromaeus
LeyesaurusSaturnaliaPantydracoEfraasiaMassospondylus adultMassospondylus embryoYizhousaurus
ChilesaurusDaemonosaurusJeholosaurus
JesairosaurusHypuronector
VallesaurusAvicranium
Megalancosaurus
TridentinosaurusCoelurosauravusMecistotrachelosLanthanolaniaIcarosaurusKuehneosaurusXianglong
Owenetta rubidgeiOwenetta kitchingorumCandelaria
FeeserpetonAustralothyrisEocaseaDelorhynchusMicroleterAcleistorhinusEunotosaurusEorhynchochelys
OedaleopsEothyrisColobomycter vaughniColobomycter pholeterDatheosaurusEnnatosaurusCaseaCotylorhynchus
Placodus inexpectatus
HipposaurusBiarmosuchusTitanophoneusProcynosuchusThrinaxodonProzostrodonProbainognathusChiniquodonCastorocaudaHaldanodon
Liaoconodon
Pachygenelus
RiograndiaChaliminia
OligokyphusKayentatheriumTritylodonSpinolestesJeholodens
GobiconodonRepenomamus robustusRepenomamus giganticus
Yanoconodon
SinoconodonBrasilodonHaramiyavia
Therioherpeton
Megazostrodon
Hadrocodium
Dromiciops
Asioryctes holotype
Monodelphis domestica
Marmosa
Chironectes
Docofossor
Volaticotherium
Alcidedorbignya
Maotherium
Maelestes
AnagaleLeptictisRhynchocyonHarpagolestes macrocephalusSinonyx
AndrewsarchusHemicentetesTenrecLeptictidiumIndohyus
Onychodectes
EctoconusPantolambda
BarylambdaGlyptodonParamylodonBradypusPeltephilusFruitafossor
Orycteropus
Myrmecophaga
HolmesinaChlamyphorus
DasypusTamanduaCyclopesMeniscotherium
OrthaspidotheriumTitanoides
AstrapotheriumPhenacodusThomashuxleya
GobiatheriumArsinoitheriumCoryphodonUintatherium
MesonyxHarpagolestes uintensisOcepeia
HomalodotheriumMiocochiliusProtypotherium
Litolophus
EmbolotheriumEotitanops
MetamynodonAceratherium
Hyracotherium
HyracodonUintaceras
Hyrachyus
Mesohippus
Pappoceras
Equus
JuxiaParaceratherium
Ceratotherium
Anisodon grande
PakicetusMaiacetus
Mammalodon
CoronodonAetiocetus
Zygorhiza
Mystacodon
Physeter
MonodonDelphinapterusTursiopsOrcinus
Hippopotamus
AnthracobuneCambaytheriumCornwallius
Janjucetus
PaleoparadoxiaNeoparadoxia
Desmostylus
Behemotops RBCM sp.
EschrichtiusIsanacetusBalaenoptera
TokarahiaYamatocetus
CetiotheriumSitsqwayk
CapereaEubalaena
ArganacerasSphodrosaurusSclerosaurus
Odontochelys
TrionyxPerochelys
Ocepechelon
Bunostegos
Elginia mirabilisElginia wuyongae
NiolamiaMeiolaniaProterochersisProganochelys
KayentachelysBaena
Kallokibotion_UBB ToK-2Arvinachelys
GlyptopsMongolochelysChubutemysCheloniaSantanachelysDermochelys
Macrochelys temminckiiTerrapene carolinaFoxemys
Carettochelys
PelomedusaAraripemysChelus
SaichangurvelMagnuviator
PleuraspidotheriumPeriptychus
Nambaroo
?Hapalodectes IVPP V2385
70
Anthracodromeus
Reptilia
UtegeniaBalanerpetonCaerorhachisAmphibamusPlatyrhinops
DendrerpetonCochleosaurus
TersomiusAchelomaPlatyhistrix
StegopsDissorophus
BroiliellusCacops
EcolsoniaFedexiaTambachia
GeorgenthaliaMicromelerpetonDoleserpetonApateonGerobatrachusAndriasCeltedensRana Utaherpeton
TuditanusKirktonecta
SauropleuraPtyonius
BatropetesRhynchonkosAsaphesteraMicraroterEuryodus primusPantylus
ScincosaurusBatrachiderpetonKeraterpetonDiceratosaurusDiplocaulusDiploceraspis Acherontiscus
MicrobrachisAdelospondylusAdelogyrinusChinlestegophisBrachydectesLysorophusEocaeciliaDermophis
CheirolepisEusthenopteronGogonasusOsteolepis
PholidogasterColosteus
PanderichthysTiktaalik
GreererpetonKoilopsSpathicephalusTrimerorhachis
DvinosaurusBatrachosuchusGerrothoraxNeldasaurusTrematosaurusMetoposaurusRhineceps
WantzosaurusTrematosaurus long rostrum
OssinodusAcanthostegaMGUH-VP-8160
DeltaherpetonVentastega
Crassigyrinus
PederpesWhatcheeria
BaphetesMegalocephalusSclerocephalus smns90055IberospondylusEryopsNigerpetonSaharastegaIchthyostegaProterogyrinusEoherpetonNeopteroplaxAnthracosaurus
Eucritta
DiscosauriscusSeymouria
MakowskiaSeymouria.tadpole
Guaibasaurus
Miohippus
Pectodens
AncodusLambdotherium
SusCainotherium
AnthracotheriumArchaeotherium
Danjiangia
Ectocion ralstonensis
NotostylopsProcavia
DusisirenProrastomus
PaleomastodonGomphotheriumElephas
Moeritherium
Amynodon
Note: This tree does not support DNA topologies over long phylogenetic distances.Certain dental traits are convergent and do not define clades.
HeptodonLophiodonTapirusProtapirus
Parotosuchus
OestocephalusLethiscus
RileymillerusPhlegethontia CGH129Phlegethontia AMNH6966
Acroplous
Eusaurosphargis juvenile
Fostercooperia
Junggarsuchus
CarnufexPseudhesperosuchusTrialestes
Asioryctes long rostrum
AriekanerpetonKotlassia
PseudocrypturusApteryx
RhynchotusEudromia
StruthioPatagopteryx
CasuariusRhea
CariamaPhoenicopterusSylviornis
SagittariusFalcoPandion
ButeoTorgos
ApusParargornis
TytoAegotheles
Llallawavis
Yanornis
ChangzuiornisJiangchangornis
LongicrusavisJinguofortis
Juehuaornis
RhamphocephalusHongshanornisDingavisCryputurusMegapodius
ApsaravisIchthyornisGansusHesperornis
CoccyzusMonias
Ardea
Ardeotis
Eozygodactylus
Butorides
AnhimaPteroclesFulica
Crex
AepyornisPalaelodus
PasserLonchuraTetrao
GallusPavo
EogranivoraMeleagris
Crax
OpisthocomusMacrocephalon
Dinornis
Ara macaoMelopsittacusNestor
Gastornis
AramusRallus
GaviaAechmophorus
Thalasseus
Charadrius
BalearicaJacanaHimantopus
EurypygaGrus
ChroicocephalusEocypselusArchilochus
MegaceryleJabiru
CinclusTroglodytes
MelanerpesSitta
UriaAptenodytes
Geococcyx
PsophiaMusophaga
MenuraSemiopteraParadisaea
CorvusTurdus
Quiscalus
Ciconia
Hirundo
Cyanocitta
Rhynochetos
PalaechthonCynocephalus
ChriacusOnychonycteris
RousettusMyotisChoeronycteris
ZhangheotheriumMetachcheiromysManis
Archaeopteryx siemensii Thermopolis
Archaeopteryx albersdoerferi Daiting
Archaeopteryx siemensii BerlinLiaoning embryo birdArchaeopteryx LondonChongmingiaCathayornisSulcavisPengornisChiappeavis
Yanornis STM9-52
IberomesornisProtopteryx
SerikornisArchaeopteryx recurva (Eichstätt)JianianhualongLiaoningvenator
Sapeornis IVPP V13276Sapeornis DNHM-F3078
Confuciusornis GMV-2132
Archaeopteryx #11
Confuciusornis MBAv1168Changchengornis
Zhongornis
ArchaeornithuraVegavis
Ostromia - Haarlem
Oedicnemus
Falcarius
DeltatheriumBishops
Caluromys
Ectocion cedrus UM 8615
MicromeryxGiraffa
Camelus
98
Miocaperea
Chonecetus
Archaeopteryx 12 private
Archaeopteryx bavarica (Munich)Jeholornis primaJeholornis curvipesHong Kong specimen
Mei
ScansoriopteryxYi
EpidexipteryxOmnivoropteryx
Archaeovolans (=Yanornis)
Draco volansMoloch
Lyriocephalus
MegachirellaBRSUG 29950-12
Pleurosaurus
MarmorettaGephyrosaurusAnkylosphenodonPaleopleurosaurusPlanocephalosaurusHeleosuchusKallimodonSphenodonBrachyrhinodonClevosaurusSphenotitanLeptosaurusVadasaurusSapheosaurusNoteosuchus
TrilophosaurusAzendohsaurusShringasaurus
EohyosaurusMesosuchusPriosphenodonRhynchosaurus articepsBentonyxHyperodapedon
Colobops
SaurosternonPalaegama
PolychrusJZC Bu154 in amber
Archaeopteryx (Wellnhoferia) grandis
Buriolestes ULBRA-PVT280
PteroglossusBuceros
Septencoarcias
Pelecanus
Catharacta
Balaeniceps
Fregata
PelagornisPuffinus
MacronectesMorus
Fraterculus
RynchopsPinguinus
Psilopogon
Cyrilavis
Phaethon
Upupa
Scopus
Phalacrocorax
ThreskiornisPlatalea
AnasAnserMergusPresbyornisPumiliornis
ColumbaCaloenas
PezophapsRaphus
AptornisDidunculus
CoragypsVultur
Goura
Lepidosaurmorpha
Archosauromorpha
JuramaiaChaoyangodensBrasilitheriumKuehneotherium
CifelliodonTachyglossusAkidolestesOrnithorhynchusUkhaatherium
Morganucodon
EomaiaAgilodocodon
AcristatheriumCronopio
DidelphisAmbolestes
Arctocyonnot AmphicyonThylacinus
Thylacosmilus
OxyaenaVincelestes
HyaenodonErnanodonDidelphodonMasrasectorBorhyaena
PeramelesMacrotisAnebodon
Notoryctes
InteratheriumMesotherium
Procoptodon
Dendrolagus
Proargyrolagus
Macropus
EurygeniumToxodon
PyrotheriumVombatusPhascolarctos
PaedotheriumGroeberiaVintanaBalbarooPhalangerPetaurusThylacoleo
Sinodelphys
TheosodonMacrauchenia
Chalicotherium? goldfussi
PaludidracoCymatosaurus
Lactodens
?Hapalodectes IVPP V5235NotharctusIndriRooneyiaDarwiniusTarsiusAegyptopithecusProconsulPanGorillaArdipithecusHomo sapiens
Ptilocercus
Tupaia
MacroscelidesChrysochlorisNecrolestes
ApatemysTrogosusRhyncholestesScutisorexLimnogaleMicrogale cowaniPotamogaleMicropotamogaleZalambdalestesSolenodonHenkelotheriumGomphosOryctolagus
BrachyerixErinaceusEchinopsThomomysHeterocephalusCeratogaulusCoendouMaiopatagiumNeoreomysChinchillaCaviaHydrochoerus
ShenshouVilevolodonRattusMusParamysRatufa
PaulchoffatiaIgnaciusPlesiadapisDaubentoniaCarpolestesMegaconusRugosodonXianshouKryptobaatarPtilodusYubaatarTaeniolabis
Mirischia
ColiusPharomachrus
BrachiosaurusCamarasaurus
DiplodocusRapetosaurus
Pteropus
VulpavusProtictisNandiniaProcyonAilurusProhesperocyonHerpestesTalpaMiacisHyopsodusEnaliarctosDesmatophocaPithanotariaZalophusProtelesCanisAmphicyon majorHesperocyonPantheraCrocutaAmphicyon galushiMustelaAiluropodaPuijilaNeotheriumUrsusGuloArctodusStylinodonAmphicynodonPsittacotheriumPalaeosinopaPhocaOdobenus
DiplovertebronRomeriscusBystrowiellaSolenodonsaurusLaosuchusChroniosaurus PIN3585/124ChroniosaurusChroniosuchus
HayaPisanosaurusScelidosaurusEmausaurusMinmiAgilisaurusStegocerasKulindadromaeusTianyulongHeterodontosaurusLesothosaurusScutellosaurusLiaoningosaurusStegosaurusChangchunsaurusDryosaurusEdmontosaurusLaquintasauraAquilopsHexinlusaurusYinlongPsittacosaurus mongoliensisLiaoceratopsAuroraceratopsLeptoceratopsStenopelix
Triceratops adultTriceratops juvenileChasmosaurus adultChasmosaurusjuvenile
72
86
9796
88
71100
8799
69
9498
82
70
9598
6264
64
79
7496
62
100 100
100<50
Compsognathus CNJ79
ProceratosaurusScipionyx
Deinocheirus
XiongguanlongSuchomimusSpinosaurusDilong
GuanlongMegaraptor
Yutyrannus
SinraptorMajungasaurus
AllosaurusSinosaurusMonolophosaurusAcrocanthosaurusEustreptospondylusCeratosaurus
Tanycolagreus
83
92
10067
9676
78
<50
62
9270
97
7180
6267
77
Anchiornis BMNHC PH804
Barlochersaurus
Llanocetus
EohupehsuchusParahupehsuchusHupehsuchus
88
100
74
87
82
89
95
95
87
75
79
98
87
100
97
87
100100 92
72
75
85
72
94
83
85100
100
100
98100
72
84
89
100
94
9860
100
91
85
100
73
6694
<50
99
80
<50 959960
63
82
99
7791 78
87
84
93
94
85
94
82
100
9194
99
9797
99
96
100
95
84
8093
84
60
95
9772
92
90
99
85
98100
89
97
98
99
9390
97
98
72
9991
98
57
65
54
57
80
62
81
88
95
89
89
100100
100
98
89
5592
81
70
90
7577
96
9564
51
100
<5091
93
7856
80
5585
7882
73
6460
95
62
98
62
8079
9897
66
84
8176
8955
7162
6094
99
90
7195
89
91
87
81
9396
91100
83
5792
9088
9495
95
76
7588
84
100
7998
62
57
80
65
89
72
75
78
55
100
95 90
67
93
99
86
92
76
6169
9696
99
83
100
7794
98
87<50
79
8368
100100
100
97
8698
77
9997
9998
83
83
100
92
57
64
79
9059
81<50
<5085
9281
100100
99
96
99
86
93
98
97
88
9497
100
9997
9464100
99
87
96
92
66
7377
99
97
81
100
91
100
97
88
98
100
100
90
90
98
96
97
9991
82
84
84
<50
87
51
93
10095
9688
7167
6569
6791
73
73
79100
100
95
94
100
79
10097
98100
95
86
62
60
86
10098
9889
89
96
78
55
65100
90
75
79
8377
100
6993
86100
9370
9499
10099
9992
99
99
97
7080 98
99
10096
7466
8696
99
88
86
<50
98
95
95
75
67
72
98
7198
9296
5898
5269
9898
98
99
94
91
85
68
8082
7188
<5057
5384
<5096
9582
83
99
6887
81
6691
100
57
65
70
83
55
84
66
80
67 99
78
69
93
67
9575
99
83
7885
77
95
88
10091
96
<50
80
80
5090
76
93
63
<50
91
82
82
61100
91
81
84
84
67
5186
90
84
89
82
76
80
97
97
82100
8599
82
87
97
62
62
52
7182
8394
100
8199
94
ArchaeothyrisHaptodusDimetrodonEdaphosaurusVaranosaurusOphiacodonCutleriaStenocybusIVPP V18117GalechirusGalepusGaleopsPatranomodonSuminiaEodicynodonDicynodonVenjukoviaOtsheriaAnomocephalusTiarajudensHipposaurusIctidorhinusHerpetoskylaxLemurosaurusProburnetiaBurnetiaBiarmosuchusEotitanosuchusCyonosaurusPromoschorhynchusScymnognathusGorgonopsAelurognathusLycosuchusProcynosuchusThrinaxodonChiniquodonPachygenelusMegazostrodonIVPP V18119MicrouraniaIVPP V18120SinophoneusDeuterosaurusEstemmenosuchus uralensisEstemmenosuchus miribilisPhthinosuchusArchaeosyodonAustralosyodonSyodonTitanophoneusAnteosaurus magnificusAnteosaurus abeliBiseridensJonkeriaStruthiocephalusStruthionopsMoschopsUlemosaurus
84
8886
68
79
81
89
7098
85100
89
98
94
51
93
100
68
85100
5560
99
93
100
98
83
92 8297
9781
9590
10099
100
66
6968
77
87100
76
7479
86
89
8992
8881
9895
92
5766
87
9868
86
63
86 86
88
99
99
98
10075
98
9899
97
81
98
100
96
98
94
9593
85
96
88100
100
100
93
66
74
94
9870
55
70
93
9363
99
74
85
63
98
85
72
5090
7385
9197
81
8873
91
8174
84
100100
100
9983
86
96
96
100
89
9 4
<50
<50
97
100
71
87
100
100
95
97
67
72
87
91
79
9583
78
72
85 6798
81
90
100
58
98
73
79
7569
87
99
76
80
71
66
80
81 71
100
9986
67
97
95
75
9590
97
100
55
56
94
90
88
95
<50
96
76
66
89100
<5098
87
98
90
100
91
67
97
59
<50
10094
78
85
9953
55
56
92
64
61<50
99<5078
<50
8175
90100
100
9590
9385
9896 90
51
89
95
79
9575
73
73
9496
94100
72
100
92
99
<50
100
7667
5599
7880
88
51
89
8272
91
70100
86
87
90
84
77
9791
98
100
81
83
98
96
88
99
56
65 100
7385
8668
76
<50
58
83
95
778555
7795
7987
94
96
9998
98
97
<50
100
58
93
76
98
95
93
<50
54
50
96
95
98
91
86
86
77
7895
99
72
80
63
58
58
58
8590
77
71
95
69
87
96
82
9879
89
<50
99
9592
96
83
<50
82
8288
98
7399
100
9697
<50
92
9385
78
7671
77
99
100
98
75
9560
64
88
65
65
95
8596
7286
99
81
100
8994
96
69
100
92
99
99
9195
96
66
Basal Therapsida59 taxa75 skull only characters
82
8795
96
97
94
100
76
<50
9597
95
62
78
<50
74
88
99
96
100
96
96
91
85
85
100
97
94
93
89
88
93
87
92
9796
83
82
74
99
100
83
75
100
88
100
91
81
52
98
98
100
83
66
99
100
87
71
6479
100 55
92
69
73
92
89
60
83
7384
80
85
95
85
98100
52
<50
<50
98
88
91
84
92
98
100
100
89
97100
73
51
70
<50
72
93
65
8198
82
95
98
95
82
76
76
79
<50
77
96
81
95
69
8387
87
93
9494
97
8084
94
72
8968
91
96
71
95
9494
<50
<50
71
8688
77
96
9999
80
9870
87
70
89100
96
87
96
6387
83
89
73
65
83
93
66
86
57
80
98
97
78
90
6877
78
62
88
60
57
85
92
99
56
98
92
57
90
82
Bootstrap scores are based on testing overlapping subsetsdue to limitations in computing power.
94
99
94
95
88 99
100
9299
9097
87
89
100
100
8199
99
100
60
75
79
68
64
100
90
62
8087
80
91
88
100
97
88
62
97
9099
91
7910098
89
10099
90
77
96
100
100
82
74
95
<50
79
57
90
79
93
95
60
96
65
74
64
90
75
67
87
95
9398
87
7385
90
87
78
6371
63
95
64
97
100
61
94
61
88
74
64
82
81
100
64
57
95
98
98
67
85
67
80
9395
91
89
<50
88
97
96
98
88
88
75
97
7598
<50
54
56 99
93
80
96
96
78
6489
9293
94
<50
85
<5072
<50
7778
<50
96
<50
51
77
91
83
92
<50
6985
85
6881
100
96
62
88
67
84
81
94
100
99
94
95
<50
85
85
85
99
90
<50
53
72
96
8890
98
97
<50
93
8552
78
78
100
93
78
<50
67
100
97
98
67
100
97
100
97
93
<50
99
<5067
70
77
98
95
<50
76
PriacodonTrioracodon
Tinodon
ShuotheriumAusktribosphenosAmphitheriumDocodon
81 99
81
91
57
88
93
97
100
78
57
10076
96100
67100
96
8377
88
<5055
81
99
8596
95
96
<50
95
57
88
81
<50
61
99
6288
100
100100
82
7778
8594
9799
58
59
76
Milosaurus
83100
7065
80
57
81
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