a systematic checklist of the hemerobiidae of the world

49
-- -- - Advance\ in Neuropterology Proceedings ol the rhird International Sympos~unl on Neuropterology Berg en Dal, Kruger National Park, R S A , !9g Mansell, M W & h k , H (E&) Pretoria, R S A 1990 Pp 215-262 - - A systematic checklist of the Hemerobiidae of the world (Insecta: Neuroptera) Victor J. MONSERRAT Ilniversidad de Complutense, Madrid, Spain ABSTRACT This paper alphabetically catalogues the extant taxa of HEMERORIIDAE. All genera and species described since Linnaeus (1758) are listed to facilitate easy reference, including the original author, data and country for each. Type species of all genera are indicated. The common misidentifications and established synonyms are chronologically listed. More than 50 new combinations are proposed and some homonymies have been detect- ed. Fifteen nominn dubin and 28 rtomim nuda species are also included to make the catalogue more comprehensive. The family Hemerobiidae presently comprises 546 valid extant species belonging to 42 valid genera. The 61 genera and 222 species previously synonymized by various authors are excluded from the above total. Five subspc- cics, 25 varieties and two forn~s are listed as well. Key words: Hemerohiidae. checklist, world c;~talogue,\ysternallcs INTRODUCTION With a nearly world-wide distribution, the family HEMEROBIIDAE can be considered one of the most common families of Neuroptera, both as regards abundance of individu- als and the number of species. Only Myrmeleontidac and Chrysopidae surpass them in terms of described species. This complicates a general systematic study, which is almost irrq~ossible bccause of the large number of species which have been inadequately described on the basis of irrelevant characters. Genital structures, which are definitive in the deter- mination of most species, have been practically ignored. Deficient typification of many species and taxonomic problems emanating from the inadequate delimitation of genera, combined with poor attempts at general classifications, makes a review of the Hemerobii- dae one of the most urgent priorities in the study of Neuroptera. Since Linnaeus (1758) erected the indefinite genus Hrrnerobius the number of taxa has ~ncreased to the point where the volume of data is difficult to handle. Despite this, no general revisions have hitherto been attempted. Numerous papers and monographs of importance have been published, and they have gradu- ally addressed many general questions: Banks (1905a), Nakahara (1960a): or certain genera, Kimmins (1937, 1943): or certain geographic areas, Carpenter (1940), Tjeder (1961), Aspiick & Aspiick (1964b), Aspiick. Aspock & Hiilzel(1980): or countries, Esben-Petersen (1 929a), Killington (1936, 1937), Zimmermann (1940), Meinander (1962), Kuwayama (1 962), Zeleny (1963), Alayo (1968), Kis, Nagler & M2ndru (1970), Penny & Monserrat (1985) etc.

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Page 1: A systematic checklist of the Hemerobiidae of the world

- - -- --p pp -

Advance\ in Neuropterology Proceedings ol the rhird International Sympos~unl on Neuropterology Berg en Dal, Kruger National Park, R S A , !9g Mansell, M W & h k , H (E&) Pretoria, R S A 1990 Pp 215-262

- -

A systematic checklist of the Hemerobiidae of the world (Insecta: Neuroptera)

Victor J. MONSERRAT Ilniversidad de Complutense, Madrid, Spain

ABSTRACT

This paper alphabetically catalogues the extant taxa of HEMERORIIDAE. All genera and species described since Linnaeus (1758) are listed to facilitate easy reference, including the original author, data and country for each. Type species of all genera are indicated. The common misidentifications and established synonyms are chronologically listed. More than 50 new combinations are proposed and some homonymies have been detect- ed. Fifteen nominn dubin and 28 rtomim nuda species are also included to make the catalogue more comprehensive.

The family Hemerobiidae presently comprises 546 valid extant species belonging to 42 valid genera. The 61 genera and 222 species previously synonymized by various authors are excluded from the above total. Five subspc- cics, 25 varieties and two forn~s are listed as well.

Key words: Hemerohiidae. checklist, world c;~talogue, \ysternallcs

INTRODUCTION

With a nearly world-wide distribution, the family HEMEROBIIDAE can be considered one of the most common families of Neuroptera, both as regards abundance of individu- als and the number of species. Only Myrmeleontidac and Chrysopidae surpass them in terms of described species. This complicates a general systematic study, which is almost irrq~ossible bccause of the large number of species which have been inadequately described on the basis of irrelevant characters. Genital structures, which are definitive in the deter- mination of most species, have been practically ignored. Deficient typification of many species and taxonomic problems emanating from the inadequate delimitation of genera, combined with poor attempts at general classifications, makes a review of the Hemerobii- dae one of the most urgent priorities in the study of Neuroptera.

Since Linnaeus (1758) erected the indefinite genus Hrrnerobius the number of taxa has ~ncreased to the point where the volume of data is difficult to handle. Despite this, no general revisions have hitherto been attempted.

Numerous papers and monographs of importance have been published, and they have gradu- ally addressed many general questions: Banks (1905a), Nakahara (1960a): or certain genera, Kimmins (1937, 1943): or certain geographic areas, Carpenter (1940), Tjeder (1961), Aspiick & Aspiick (1964b), Aspiick. Aspock & Hiilzel(1980): or countries, Esben-Petersen ( 1 929a), Killington (1936, 1937), Zimmermann (1940), Meinander (1962), Kuwayama ( 1 962), Zeleny (1963), Alayo (1968), Kis, Nagler & M2ndru (1970), Penny & Monserrat (1985) etc.

Page 2: A systematic checklist of the Hemerobiidae of the world

However, only the Nearctic, European and South African faunas represent a "satisfacto- ry" level of knowledge, although data on biology and larval stages of many species are still woefully inadequate. In general, what is known about the remainder of the brown lacewings is not satisfactory. Although there are some good descriptions, most are based on irrelavant data and need to be adequately revised. New synonymies and combinations should appear after the remaining original material has been properly studied and redescribed, and many species whose type series have been lost will also be designated as nomina dubia.

Sorne attempts at systematic and general classification of the family have been made. Us- ing characters of wing venation, Cornstock (1918) and Kriiger (1922) created five subfa- milies: DREPANEPTERYGINAE, MEGALOMINAE, HEMEROBIINAE, SYMPHEROBIINAE and MICROMINAE. Some were used simultaneously at a tribal level (NavBs 1933a) or were elevated to family category (Brues & Melander 1932; Kuwaya- ma 1962). Lastly, Nakahara (1960a) divided the family into two subfamilies. NOTIO- BIELLINAE and HEMEROBIINAE, based upon the presence or absence of the phallolingual membrane in males. Since the validity of these classifications has been ques- tioned and thcy have not been widely accepted, a general classification on the basis of adequate characters is still pending (Tjeder 1961; Aspock, Aspock & Hiilzel 1980).

The first prerequisite in undertaking a general study of this family is the compilation of a world catalogue of the known taxa. Besides the above-mentioned works, the following papers contain some catalogues that list the local fauna of specific regions or countries (Nakahara 1919; Stange 1967; Ghosh & Sen 1977; Penny 1977). However, since Hagen (1866) listed the species and known synonymies and transferred many species that were originally described in Hemerobius to other genera, there has been no attempt to compile a general list. Some species with poor original descriptions, which are still included in this genus, will probably be transferred to other genera, families or even orders when the original material is located and studied.

This paper catalogues as completely as possible, all taxa of extant Hemerobiidae, listing the genera and species, as well as subspecies, varieties and forms described since 1758 and not subsequently transferred to other families or orders. I wish to note that although every endeavour has been made to consult the most important bibliographical sources, it is possible that some existing taxa are not included here.

The list of genera and species is alphabetically arranged to facilitate easy access, and in- cludes a brief bibliographical reference with the author and date of publication, as well as the country or geographical region where first collected. For all genera, each type- species is marked with an asterisk and the common misidentifications and established syn- omyms are listed chronologically in italics.

All species are listed according to their original designation or follow the actual renomi- nation if the species was transferred to another genus. More than 50 new con~binations are also proposed, and it should be mcntioncd that somc species names will require modifi- cation pending future studies. Some homonymies have been detected, but replacement names will not be designated until the relevant taxa have been adequately studied.

Finally, there is also mention of some early species which lack references, as original descriptions could not be traced or were perhaps not published. In these cases, the first reference citing the name is simply noted. Fifteen nomina dubia and 28 nornina riuda spe- cies are also mentioned in the list to make it more complete.

The existing family HEMEROBIIDAE comprises 546 valid species belonging to 42 valid genera, excluding the 61 genera and 222 species previously proposed as synonyms. Without doubt, a thorough study of this family will reduce the number of presently valid taxa even

Page 3: A systematic checklist of the Hemerobiidae of the world

World catalogue of Hernerobiidae

further, and new descriptions will be added in future. But, whatever the case, 1 hope that this list will provide an initial basis for future general studies of the family.

Family HEMEROBIIDAE Latreille, 1 803.

Afiomicromus Nakahara, 1960a Allemerobius Banks, 1940

*Allemerobius flaveolus Banks, 1940. Allotomyia Banks, 1930c. Ameromicromus Nakahara, 1960a Anapsectra Tjeder, 1975

*Anapsectra medleri Tjeder, 1975 Annandulia Needham, 1909 Anomicromu.~ Nakahara, 1960a Anotiobiella Kimmins, 1928 Archaeornicromus Kriiger, 1922 Austromegalomus Esben-Petersen, 1935b

*Austromegalomus brunneus Esben-Petersen, 1935b Austromegalomus insulanus Oswald, 1988

Austromicromus Nakahara, 1960a Bestreta NavBs, 1924b Boriomyia Banks, 1904b nec 1905a

= Allotomyia Banks, 1930c *Borio~-ny ia fidelis (Banks, 1897)

= Hemerobius fidelis Banks, 1897 Boriomyia speciosa (Banks, 1904)

=Hemerobius speciosus Banks, 1904a Boriomyia Banks, 1905a nec 1904b Brauerobius Kriiger, 1922 Buxtonia Esben-Petersen, 192% Canisius Navris, 1913d Carobius Banks, 1909

Carobius angustus Banks, 1909 *Carobius pulchellus Banks, 1909 Carobius subfasciatus Tillyard, 1916 Carobius trifurcatus Kimmins, 1940

Coloma NavBs, 1915c Conchopterella Handschin, 1955

*Conchopterella kuscheli Handschin, 1955 Conchoptcrella maculata Handschin, 1955

Drcpariacra Tillyard, 1916 = Menopteryx Kriiger, 1922

Drepanacra binocula (Newman, 1838) = Drepanepteryx binoculus New man, 1838 = *Drt.punc.pteryx humilis McLachlan, 1866 = Drepanepteryx itzstabilis McLachlan, 1866 = Mega1omu.s lanceolatus Gerstaecker, 1885 = Drepanepteryx humilior Hare, 191 0 = Drepanepteryx maori Hare, 19 10 = Drepanacra hardyi Tillyard, 19 16 = Drepanacra froggatti Till yard, 19 16 =Drr~patzacrci norfolkensis Till yard, 19 17

see Micromus Rambur, 1842

CHINA see Boriomyia Banks, 1904 see Micromus Rambur, 1842

NIGERIA see Psectra Hagen, 1866 see Micromus Rambur, 1842 see Hemerobius Linnaeus, 1758 see Micromus Rambur, 1842

TAHITI RAPA ISLAND

see Micromus Rambur, 1842 see Drepanepteryx Leach, 1815

USA

USA see Wesmaelius Kriiger, 1922 see Hemerobius Linnaeus, 1758 see Notiobiella Banks, 1909 see Drepanepteryx Leach, 1815

AUSTRALIA AUSTRALIA AUSTRALIA AUSTRALIA

see Sympherobius Banks, 1904

CHILE CHILE

AUSTRALIA AUSTRALIA

NEW ZEALAND AUSTRALIA

NEW ZEALAND NEW ZEALAND

TASMANIA AUSTRALIA

NORFOLK ISLANDS

Page 4: A systematic checklist of the Hemerobiidae of the world

Drcpanacra binocula v. longitudinalis Till yard, 1916

=Drepanacra humilis v. longitudinalis Tillyard, 1916

Drepanacra binocula v. tasmanica Tillyard, 191 6 = Drepanacra humilis v. tasmanica Tillyard,

1916 Drepanacra binocula v. pallida Tillyard, 19 16

=Drepanacra instabilis v. pallida Tillyard, 1916

Drepanacra binocula v. rubrinervis Tillyard, 1916

=Drepanacra instabilis v. rubrinervis Tillyard, 1916

Drepanacra binacula ssp. insularis Tillyard, 1917 =Drepanacra instabilis ssp. insularis Tillyard,

1917 Drepanacra binocula v. bilineata Tillyard, 1923 Drepanacra binocula v. divisa Till yard, 1923 Drepanacra binocula v. excisa Tillyard, 1923 Drepanacra binocula v. suboculata Tillyard,

1923 Drepanacra plaga Banks, 1939 Drepanacra yunnanica Y ang, 1986

Drepanacrella Kimmins, 1940 *Drepanacrella khasiana Kimmins, 1940

Drepaneptery X Leach, 18 15 = Drepanopteryx Burmeister, 1839 = Cunisius Navas, 1913d = Oeclobius Nakahara, 19 1 5 =Phlebonema Kriiger, 1922 =Bestreto Navas, 1924b

Drepanepteryx algida (Erichson, 185 1) = Hemerobius algidus Erichson, 1 85 1 = Hemerobius kollari Giiszy , 1852 = Megalonzus conspersus Schoch, 1885

Drepanepteryx berothoides McLachlan, 1869 Drepanej>teryx calida (Kriiger, 1922)

=Phlehnnema calida Kriiger, 1922 nom.nud. Drepanepteryx falculoides Walker, 1860 Drepanepteryx fuscata Nakahara, 1960a *Drepanepteryx phalaenoides (Linnaeus, 1758)

=Hemerobius phalaenoides Linnaeus, 1758 Drepanepteryx pleshanovi Makarkin, 1985a Drepanepteryx punctata (Okamoto, 1905)

= Megalonlus punctatus Okamoto, 1905 nec Kimmins, 1935a

= Oeclobius infulcutus Nakahara, 1915 = Bestreta iaponica Navas, 1924b

Drepanepteryx resinata (K riiger, 1922) = Phlebonema resinatu Kriiger, 1922 nom.nud.

Drepanomima Tillyard, 1916

TASMANIA

TASMANIA

TASMANIA

TASMANIA

NORFOLK ISLANDS NEW ZEALAND NEW ZEALAND NEW ZEALAND

NEW ZEALAND FORMOSA

CHINA

INDIA

USSR AUSTRIA

SWITZERLAND A USTRALIA

HINDOSTAN JAPAN

EUROPE USSR

JAPAN JAPAN JAPAN

Page 5: A systematic checklist of the Hemerobiidae of the world

World catalogue of Hemerobiidae

*Drepanomima gibbosa Tillyard, 1916 AUSTRALIA Drepanopteryx Burmeister, 1839 see Drepanepteryx Leach, 1815 Dyshemerobius Tjeder, 1961

Dyshemerobius falciger Tjeder, 1963 RHODESIA *Dyshemerobius productus Tjeder, 196 1 RHODESIA

Egnyonyx Wesmael, 1836 see Hemerobius Linnaeus, 1758 Eucurobius Esben-Petersen, 192% see Psectra Hagen, 1866 Eurnicromus Nakahara, 19 15 see Micromus Rambur, 1842 Eurobius Kriiger, 1922 see Sympherobius Banks, 1904 Ganchetus Navas, 1929a see Notiobiella Banks, 1909 Gayomyia Banks, 19 13

=Porter Navas, 1924c Guyomyia cinerea Kriiger, 1922 nom.nud. *Gayomyia falcata (Blanchard, 185 1)

= Megalomus falcatus Blanchard, 185 1 CHILE =Porter discolor Navis, 1924c CHILE

Gayomyia stictica (Blanchard, 185 1) =Megalomus sticticus Blanchard, 185 l CHILE

Haarupiella Esben-Petersen, 19 14 see Nusalala Navas, 1913 Hagenobius Kriiger, 1922 see Hemerobius Linnaeus, 1758 Hemerobiella Kimmins, 1940

"Hemerobiella sinuata Kimmins, 1940 ECUADOR Hemerobius Linnaeus, 1758

= Egnyonyx Wesmael, 1 836 = Mucropulpus Rambur , 1842 =Stenolomus Navas, 1906 = Hemerodomia Navas, 1909b = Hagenobius Kriiger, 1922 = Reuterobius Kriiger, 1922 = Brauerobius Kriiger, 1922 = Schneiderobius Kriiger , 1922 = Anotiobiella Kimmins, 1928 =Monorobius Makarkin, 198513

Hemerobius abditus Tjeder, 1961 Hemerobius abyssinicus Esben-Petersen, 1928a Hemerobius adelgivorus Kimmins, 1960 Hemerobius albipennis Banks, 19 10 Hemerobius alpestris Banks, 1908 Hemerobius amurensis Navas, 1929c Hemerobius antigonus Banks, 194 1 Hemerobius aper Tjeder, 1961 Hemerobius atrifrons McLachlan, 1868

= Hemerobius fusciatus Goszy, 1852 nec Stephens, 1836

Hemerobius australis Walker, 1853 Hemerobius azoricus Tjeder, 1948 Hemerobius baguiensis NavBs, 1923 Hemerobius binigripunctatus Fraser, 1957 Hemerobius bispinus Banks, 1940 Hemerobius bistrigatus Currie, 1904a Hemerobius bolivari Banks, 19 10 Hemerobius holiviunus (Kriiger, 1922)

NATAL ABYSSINIA PAKISTAN

COLUMBIA USA

AMUR DOMINICAN REP.

NATAL ENGLAND

AUSTRIA AUSTRALIA

AZORES ISLANDS PHILIPPINES

REUNION ISLAND CHINA

USA COLUMBIA

Page 6: A systematic checklist of the Hemerobiidae of the world

V. J . Monserrat

= Schneiderobius bolivianus Kruger , 1922 nom.nud. Hemerobius californicus Banks. c.f. Banks, 1905b Hemerobius canadai Navris, 1924a

= Hemerobius cazadai Navis, 1924a Hemerobius centralis Navas, 1913b Hemerobius ceraticus Navis, 1924d Hemerobius cercodes Navas, 19 17e Hemerobius chiangi Banks, 1940 Hemerobius chilensis Nakahara, 1965a Hemerobius chomiacensis Dziedzielewicz, 1907 Hemerobius cixiiformis Gerstaecker, 1888 Hemerobius claggi Banks, 1937a Hemerobius colombianus Kriiger, 1 922 nom.nud. Hemerobius comorensis Kruger, 1922 nom.nud. Hemerobius conjunctus Fitch, 1855

= Hemerobius citrinus Hagen, 186 1 = Hemerobius venustus Banks, 1897 = Hemerobius cockereli Banks, 190 1 = Hemerobius caudelli Currie, 1904b =Hemerobius glacialis Currie, 1904b

Hemerobius conspurcatus McLachlan, 1875 Hemerobius contumax Tjeder, 1932

= Hemerobius limbatellus (Zetterstedt, 1840): McLachlan, 1899, nec auct.

Hemerobius costalis Carpenter, 1940 Hemerobius cubanus Banks, 1930a Hemerobius darlingtoni Banks, 1938 Hemerobius deceptor Navds, 19 14b Hemerobius disparilis Navds, 1936b Hemerobius divisus Nava's, 193 1 d Hemerobius domingensis Banks, 194 1 Hemerobius dorsatus Banks, 1904a Hemerobius eatoni Morton, 1906

= Hemerobius cornutus Navds, 1906 nec Linnaeus, 1758

= Hemerobius sciopterus Navis, 1906 = Stenolomus cabrerai Navas, 1906 =Stenolomus scalaris Navas, 1906

Hemerobius elatus Navds, 1914b Hemerobius elongatus Monserrat, 1990 Hemerobius exceptatus Nakahara, 1965a Hemerobius exoterus Navh , 1936d Hemerobius fenestratus Tjeder, 1932

=Hemerobius limbatellus (Zetterstedt, 1840): Wallengren, 1871 part. nec auct.

Hemerobius ferox Tjeder, 1937 Hemerobius frontalis Hagen, 1858b nec Navas,

1932c Hemerobius frontalis Navds, 1932c nec Hagen,

1858b Hemerobius fujimotoi Nakahara, 1960a Hemerobius gilvus Stein, 1863

SPAIN ECUADOR BURINGA

HINDOCHINA CHINA CHILE

CZECHOSLOVAKIA BRAZIL

PHILIPPINES

USA NORTH AMERICA

USA USA

CANADA CANADA

USSR FRANCE

USA CUBA

JAMAICA KENYA CONGO CHINA?

DOMINICAN REP. USA

CANARY ISLANDS

CANARY ISLANDS CANARY ISLANDS CANARY ISLANDS CANARY ISLANDS

KENYA MEXICO

COLUMBIA CHINA

SWEDEN

CHINA

CEYLON

BRAZIL JAPAN

GREECE

Page 7: A systematic checklist of the Hemerobiidae of the world

World cutulogue of Hemerobiidue

= Hemerobius occiduus Navas, 19 15d = Hemerobius burmunni Aspock, 1963

Hemerobius grahami Banks, 1940 Hemerobius greeni Banks, 1913 Hemerobius hageni Navas, 19 18c

= Megalomus pallidus Blanchard, 185 1 = Hernerobius pinnatus Navis, 19 18a nec 19 14f = Hemerobius brethesi Navas, 1929g = Hemerobius piunatulus Navas, 1935a = Hemerobius blanchardi Nakahara, 1960a

Hemerobius hageni var. distincta Navas, 1920 Hemerobius handschini Tjeder, 1957 Hemerobius harmandinus Navas, 1910a

= Hemcrobius nitidulus (Fabricius, 1777): Nakahara 19 15

= Hemerobius nakahurinus Navris, 19 16 Hemerobius hedini Tjeder, 1937 Hemerobius hengduanus Yang, 198 1 b Hemerobius hespericus Navas, 193 1c Hemerobius hopii Dale cf. Curtis, 1854 *Hemerobius humulinus Linnaeus, 1758

= Hemerobius hurnuli Linnaeus, 176 1 = Hemerobius subfasciutus Stephens, 1836 = Hemerobius crispus Stephens, 1836

nec Walker, 1853 = Hemerohius apicalis Stephens, 1836 = Hernerobius obscurus Stephens, 1836 = Hemerobius afinis Wesmael, 1840 nec Stephens, 1836 = Hemerobius muculutus Wesmael, 1840 = Mucrt~pulpu.~ lutescens (Fabricius, 1793):

Rambur, 1842 = Hemerobiu.~ obliterutus Walker, 1853 = Hemerobius castanear Fitch, 1855 = Hemerobius tutatrix Fitch, 1855 = Hemerobius gossypii Ashmead, 1895 =Hemerobius algonquinus Banks, 1924a = Hemerobius obtusus Nakahara, 1954

Hcmerobius hyalinus Nakahara, 1966 Hernerobius immuculatus Olivier, 1792 nom.dub. Hemerobius incursus Banks, 1930b Hemerobius indicus Kimmins, 1938 Hemerobius inversus Navfis, 1927b Hemerobius jamaicensis Banks, 1938 Hemerobius japonicus Nakahara, 1915

= Hemerobius spinigerus Banks, 1937b Hemerobius javanus Kriiger, 1922 nom.nud. Hemerobius jucundus Navis, 1927c Hemerobius kobayashii Nakarara, 1 956 Hemerobius kokaneeanus Currie, 1904b

= Hemerobius hesperus Banks, 1 9 2 4 Hemerobius lacunaris Navas, 1936d

SPAIN ITALY CHINA

CEYLON CHILE CHILE

ARGENTINA

ARGENTINA SWITZERLAND

JAPAN

CHINA CHINA

CANARY ISLANDS ENGLAND

EUROPE EUROPE

ENGLAND

ENGLAND ENGLAND ENGLAND

BELGIUM BELGIUM

EUROPE USA USA USA USA USA

JAPAN FORMOSA

EUROPE MALAYSIA

INDIA PHILIPPINES

JAMAICA JAPAN

FORMOSA

MEXICO JAPAN

USA USA

CHINA

Page 8: A systematic checklist of the Hemerobiidae of the world

Hemerobius lautus Navris, 1909c Hemerobius lentus Hagen, 186 1 nom.nud. Hemerobius lii Yang, 198 1 b Hemerobius limbatellus Stitz, 1927 nec auct. nom.dub. Hernerobius lutescens Fabricius, 1793 nec auct.

= Hemerobius afinis Stephens, 1836 part. nec Wesmael, 1840

= Hemerobius paganus (Linnaeus, 1767): Stephens, 1836

Hemerobius madeirae Tjeder, 1939 Hemerobius mangkamanus Yang, 198 1 b Hemerobius marginatus Stephens, 1836

= Hemerobius lutescens (Fabricius, 1793): Zetterstedt, 1840 part. nec auct.

= Hemerobius j7exuo.s~~ Hagen, 185 8c = Hemerobius humuli (Linnaeus, 176 1): Hagen,

1858a part. nec auct. = Hemerobius irregularis Nakahara, 19 15

Hemerobius marginatus ssp. lapponicus Meinander, 1962 Hemerobius melanostictos cf. Hagen, 1866 Hemerobius mexicanus Navris, 192 1 b Hemerobius micans Olivier, 1792

= Hemerobius punctatus Stephens , 1836 nec Goszy, 1852

= Hemerobius pallidus Stephcns, 1836 =Hemerobius lutescens (Fabricius, 1793):

Burmeister, 1839 nec auct. = Mucropalpus fuscinervis Schneider, 1845 = Mucropu1pu.s irroratus Costa, 1855

Henierobius micans var. conspicua Navris, 19 15d Hemerobius moestus Hagen nec Banks, 1897 cf.

Hagen, 1854 Hemerobius mononeurus Say cf. Hagen, 1873 Hemerobius montanus Kimmins, 1960 Hemerobius monticola Alayo, 1968 Hemerobius nairobicus Navas, 19 lob Hemerobius natalensis Tjeder, 1961 Hemerobius neadelphus Gurney, 1948 Hemerobius nemorensis Kimmins, 1952 Hemerobius nigrans Carpenter, 1940 Hemerobius nigricornis Nakahara, 19 15

= Hemerobius griseus Nakahara, 1956 Hemerobius nigrostigma Monserrat, 1990 Hemerobius nitidulus Fabricius, 1777

= Hemerobius humuli (Linnaeus, 176 I): Zetterstedt, 1840 nec auct.

= Hemerobius ochraceus Wesmael , 1840 = Mucropalpus obscurus Rambur, 1842 = Hemerobius haematicus Navas, 1908b

Hemerobius nitidulus var. fusca Dziedzielewicz, 1920 Hemerobius ovalis Carpenter, 1940 Hemerobius pacificus Banks, 1897

SYRIA BRAZIL CHINA

EUROPE KIIAAE

ENGLAND

ENGLAND MADEIRA ISLAND

CHINA ENGLAND

ENGLAND USSR

JAPAN FINLAND EUROPE MEXICO

ENGLAND ENGLAND

ITALY ITALY SPAIN

PAKISTAN CUBA

KENYA NATAL

CANADA KENYA

CANADA JAPAN JAPAN

VENEZUELA GERMANY

BELGIUM FRANCE

SPAIN POLAND

USA USA

Page 9: A systematic checklist of the Hemerobiidae of the world

World catulogue of Hemerobiidae

= Hemerobius pallescens Currie, 1904a USA = Hemerobius discretus Navas, 1917b USA

Hemerobius pallidulus Kimmins, 1928 = Hemerohius neglectus Hagen, 1861 nec Costa,

1855 (Chrysopidae) MEXICO Hemerobius pallipes Olivier, 1792 nom.dub. EUROPE Hemerobius parvulus (Rambur, 1842)

= Mucropalpus parvulus Rambur , 1842 nom.dub. CERDANIA = Mucropalpus parvulus var. pallidus Costa, 1855 ITALY

Hemerobius pehlkeanus (Kriiger, 1922) =Schneiderobius pehlkeanus Kriiger, 1922 nom.nud.

Hemerobius perelegans Stephens, 1 836 ENGLAND Hemerobius phaleratus Hoffmansegg cf. Hagen, 1866 Hemerobius pini Stephens, 1836 nec Leach ENGLAND

= Hemerobius fasciatus Stephens , 1836 nec Coszy, 1852 ENGLAND

= Hemerobius suericus Mjoberg, 1909 SWEDEN = Hemerohius phaleratus Hagen, 1858a part. ENGLAND

Hemerobius pini Leach nec Stephens, 1836 cf. auct. Hemerobius pinidumus Fitch, 1855 USA

= Hernerobius hyalinatus Fitch, 1855 USA = Hemerobius canadensis Banks, 1897 CANADA = Hemerobius kootenayensis Currie, 1904 CANADA

Hemerobius pinnatus Navas, 19 14f nec 19 18a COLUMBIA Hemerobius popii Esben-Petersen, 1921 UST VILUI Hemerobius punctutus Goszy, 1852 nondub. AUSTRIA Hemerobius quettanus Navis, 1930c INDIA Hemerobius radialis Nakahara, 1956 JAPAN Hemerobius reconditus Navas, 19 14b KENYA

= Hemerobius errcrns Navas, 19 14b KENYA = Hemerobius ellenbergeri Navas, 1 933d BASUTOLAND

Hemerobius ricarti Navas, 192% PHILIPPINES Hemerobius rizali Banks, 1920 PHILIPPINES Hemerobius rudebecki Tjeder, 1961 SOUTH AFRICA Hemerobius rufescens Goszy, 1852 nom.dub. AUSTRIA Hemerobius schedli Hiilzel, 1970 TIROL Hemerobius semireticulatus Bremi cf. Hagen, 1866 Hemerobius shibakawae Nakahara, 191 5 JAPAN Hemerobius shikotanus Kuwayama, 1956a KURIL ISLANDS Hemerobius .signatus Hagen, 1861 nec Kruger,

1922 nom. nud. CHILE Hemerobius signatus Kriiger, 1922 nec Hagen, 1961

nom.nud. Hemerobius simulans Walker, 1853 CANADA

= Hemerohi~ds crispus Walker, 1853 part. nec Stephens, 1836 CANADA = Hemerobius stigmuterus Fitch, 1855 USA = Hemerobius orotypus Wallengren, 1870 SWEDEN = Hemerobius nevadensis Banks, 1904a USA = Hemerobills placidus Banks, 1908 USA = Henterobius piceus Navlis, 192% USSR

Hemerobius sjostedti Navas, 192 1 a JUAN FERNANDEZ ISL.

Page 10: A systematic checklist of the Hemerobiidae of the world

= Hemerobius nigrinus Esben-Petersen, 1924 = Hemerohius furnosus Esben-Petersen, 1924

Hemerobius skottsbergi Navas, 192 1a Hemerobius solanensis Ghosh, 1976 Hemerobius stigma Stephens, 1836

= Hemerobius irroratus Stephens, 1836 = Hemerobius strigosus Zetterstedt, 1840 = Hemerohius limbatellz~s Zetterstedt, 1840 = Hernerobius limbatus Wcsmael, 1840 = Hemerobius crispus Walker, 1853 part. nec

Stephens, 1836 =Hernerobius stigmaterus Fitch, 1855 part. = Hemerobius stephensii Fitch, 1 855 = Hemerobius phalerutus Hagen, 1858a part. = Hemerobius limbatellus (Zetterstedt, 1840):

Wallengren, 187 1 part. = Hemerobius rnoestus Banks, 1897 nec

Hagen, 1854 = Hemerohiu.~ dyari Currie, 1904b = Hemerobius simplex Banks, 1905a = Hernerociomiu buyssoni Navas, 1909b = Hemerobius perij2ricu.s Navas, 19 13a

Hemerobius striatus Nakahara, 1915 Hemerobius subfalcatus Nakahara, 1960a Hemerobius sumatranus Navas, 1926b Hemerobius tagalicus Banks, 1920 Hemerobius tateyamai Nakahara, 1960a Hemerobius tibialis NavBs, 1917d Helnerobius tolimensis Banks, 19 10 Hemerobius topali Steinmann, 1965b Hemerobius triangularis McLachlan, 1875 Hemerobius trifasciatus Navas, 1932e Hemerobius tristriatus Kuwayama, 1954 Hemerobius tunkunensis Navas, 1933e Hemerobius vagans Banks, 1937a Hemerobius vaillanti Navas, 1927d Hemerobius withycombei (Kimmins, 1928)

=Anotiobiella withycombei Kimmins, 1928 Hemerobius xizangensis Yang, 1981b Hemerobius zernyi Esben-Petersen, 1935a

= Hemerobius friedeli Aspock & Aspock, 1966 Hemerodomia Navas, 1909b

Heteromicromus Kriiger , 1922

Idiomicromus Nakahara, 1955 *Idiomicromus kanoi Nakahara, 1955 Idiomicromus zanganus Yang, 1981 b

Indomicromus Kriiger, 1922

Kimminsiu Killington, 1937

JUAN FERNANDEZ ISI,. JUAN FERNANDEZ ISL. JUAN FERNANDEZ ISL.

INDIA ENGLAND ENGLAND LAPONIA LAPONIA BELGIUM

NORTH AMERICA USA USA

ENGLAND

SWEDEN

USA CANADA

USA FRANCE

SPAIN JAPAN JAPAN

SUMATRA PHILIPPINES

JAPAN COLUMBIA COLUMBIA

ARGENTINA USSR

ITALY JAPAN

TUNKUN PHILIPPINES

CHINA

MEXICO CHINA

LIB A N 0 ASIA MINOR

see Hemerobius Linnaeus, 1758 see Micromus Rambur, 1842

FORMOSA CHINA

see Micromus Rambur, 1842 see Wesmaelius Kriiger, 1922

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World catalogue c$ Hemerobiidue

AUSTRALIA see Neuronema McLachlan, 1869 see S ympherobius Banks, 1904

AUSTRALIA

Kimminsiclla Nakahara, 1960a *Kimminsiella tillyardi (Kimmins, 1940)

= Annandalia tillyardi Kimmins, 1940 Kulinga Navas, 1936d Luchlanius Kriiger, 1922 Megalomina Banks, 1909

*Megalomina acuminata Banks, 1909 Megalomus Rambur, 1842

= Pleomegalomus Kriiger, 1922 = Pirionus Navas, 1928b =Spinomegalomus Nakahara, 1965a

Megalomus acuminatus Kriiger, 1922 nom.nud. Megalomus acunai Alayo, 1968

=Megalomus acub i Alayo, 1968 CUBA Megalomus andreinii Navas, 1928c nom.dub. ITALY Megalomus angulatus Carpenter, 1940 USA Megalomus angustatus Navas, 1932e PANAMA Megalomus atomarius Navhs, 1935b MOROCCO Megalomus axillatus Navis, 1927b PHILIPPINES Megalomus balachowskyi Lestage, 1928 ALGERIA Megalomus cubanus Banks, 1930a CUBA Megalomus darwini Banks, 1924b GALAPAGOS ISLANDS Megalomus decoratus Kriiger, 1922 nom.nud. Megalomus exterior Navas, 1929f COSTA RICA Megalomus flinti (Nakahara, 1965)

= Spinomegalomus flinti Nakahara, 1965a CHILE Megalomus formosanus Banks, 1937b FORMOSA Megalomus hirtus (Linnaeus, 1761)

= Hemerobius hirtus Linnaeus, 1761 SWEDEN = Hemerobius 3mbriatus Curtis, 1828 ENGLAND = Hemerobius decussatus Leach, cf. Hagen, 1866

Megalomus hirtus var. oropaea Navas, 19291.1 ITALY Megalomus insignis Kimmins, 1935a GUATEMALA Megalomus kimminsi Nakahara, 1960a

=Megalomus punctatus Kimmins, 1935a nec Okamoto, 1905 GUATEMALA

Megalomus linguatus Navas, 19 lob ARGENTINA = Megalornus intricatus Navas, 19 14f PARAGUAY = Megalomus infuscatus Navas, 19 14f PARAGUAY = Megalomus princeps Navis, 1932b ARGENTINA = Megalomus nanus Navis, 1932d ARGENTINA = Megalomus bridarollius Navas, 1932d ARGENTINA

Megalomus linguatus var. pigm. Gonzalez, 1981 ARGENTINA? Megalomus lioni Navas, 1927d HAITI Megalomus luigionii Navis, 1928a nom.dub. ITALY Megalomus marginatus Banks, 19 10 COLUMBIA Megalomus minor Banks, 1905b NICARAGUA Megalomus moestus Banks, 1895 USA

=Megalomus latus Banks, 1903 USA Megulomus morlticcllii Navas, 1928a nom.dub. ITALY Megalomus navasi Lacroix, 1912 ALGERIA Megalomus nebulosus Navas, 1926a PERU

Page 12: A systematic checklist of the Hemerobiidae of the world

Megalomus nigratus (Navas, 1928) = Pirionus nigratus Navas, 1928b

Megalomus obscurus Steinmann, 1965a Megalomus parvulus Kimmins, 193% Megulomus purvus Kriiger, 1922 nom.nud. Megalomus pictus Hagen, 186 l Megalomus pyraloides Rambur, 1842

= Megalomus montserruticus Navas, 1915b Megalomus rafaeli Penny & Monserrat, 1985 Megalomus serrinus Navas, 1936a Megalomus setosulus (Walker, 1860)

=Hemerobius setosulus Walker, 1860 = Megulomus noualhieri Navas, 19 lob

Megalomus sinuutus Kriiger, 1922 nom.nud. Megalomus stangei Gonzalez, 198 1 Megalomus tineoides Rambur , 1842 *Megalomus tortricoides Rambur , 1842

= Megabmus tener Navis, 191 5b Megalomus uniformis Banks, 193 5 Megalomus yunnanus Yang, 1986

Menopteryx Kriiger, 1922

Menutus Navas, 1932a *Menutus haitiensis (Smith, 193 1)

=Micromus haitiensis Smith, 1931 Meomicromus Kruger, 1922

Mesohemerobius Nakahara, 1966 nec Ping, 1928

Micromus Rambur, 1842 = Ne.somicromus Perkins, 1899a = Pseudopsectru Perkins, 1899a = Nesothuumu Perkins, 1899a =Nenus Navas, 1912a = Eumicromus Nakahara, 19 15 = Paramicromus Nakahara, 19 19 nec Kriiger, = Parumicromus Kriiger, 1922 nec Nakahara, = Pscudomicromus Kruger , 1922 = Archaeomicromus Kruger, 1922 = Meomicromus Kruger , 1922 = Neomicromus Kriiger , 1922 = Stenomicromus Kriiger, 1922 = Indornicromus Kruger , 1922 = Heteromicromus Kruger, 1922 = Phlebiomus Navas, 1922a = Tunca Navas, 1929e = Anomicromus Nakahara, 1960a = Spilomicromus Nakahara, 1960a =Au.stromicromus Nakahara, 1960a = Ameromicromus Nakahara, 1960a = Afromicromus Nakahara, 1960a

Micromus acutipennis Kimmins, 1956

CHILE MONGOLIA

WEST INDIES

NORTH AMERICA FRANCE

SPAIN BRAZIL

COSTA RICA

HINDOSTAN INDIA

ARGENTINA SPAIN

HUNGARY SPAIN

USA CHINA

see Drepanacra Tillyard, 1916

HAITI see Micromus Rambur, 1842 see Semohemerobius Yang, 1983

BELGIAN CONGO

Page 13: A systematic checklist of the Hemerobiidae of the world

World catalogue of Hemerohiidue

Micromus africanus Weele, 1910b = Micromus ludicrus Navas, 1933d

Micromus alternans Navas, 1922d Micromus angularis (Perkins, 1899) nov.comb.

= Nesomicromus angularis Perkins, 1899b = Nesomicromus phaeostictus Perkins, 1899b

Micromus angulatus (Stephens, 1836) =Hemerobius aphidivorus Schrank, 1781 nom.dub. = Hemerobius angulatus Stephens , 1836 = Hernerobius villo.sus Zetterstedt, 1840 = Hemerobius intricatus Wesmael , 1840 = Micromus tendinosus Rambur, 1842 = Hernerohius lineatus Goszy , 1852 = Micromus jonus Needham, 1905

Micromus angustipennis (Perkins, 1899) nov.comb. = Nesomicromus angustipennis Perkins, 1899a

Micromus atlanticus Tjeder, 1976 Micromus audax (Kriiger, 1922)

= Heteromicromus audax Kriiger, 1922 nom.nud. Micromus australis (Hagen, 1858) nec Froggatt, 1904

=Micrornerus australis Hagen, 1858b Micromus bellulus (Perkins, 1899) nov.comb.

= Nesomicromus bellulus Perkins, 1899a =Nesomicromus rninor Perkins, 1899a = Nesomicromus molokaiensis Perkins, 1899a = Nesomicromus inJumatus Perkins, 1899a

Micromus benardi Navis, 19 12b Micromus bifasciatus Tillyard, 1923 Micromus bifasciatus var. amabilis Tillyard, 1923 Micromus bifasciatus var. imperfectus Tillyard, 1923 Micromus brunnescens (Perkins, 1899) nov-comb.

= Nesomicromus brunnescens Perkins, 1899a Micromus calidus Hagen, 1859 Micromus canariensis Esben-Petersen, 1936 Micromus canariensis var. curvatus Esben-Petersen,

1936 Micromus carpentieri Lestage, 1925 Micromus confusus (Nakahara, 1964)

=Eumicromus confusus Nakahara, 1964 =Eumicromus sauteri Nakahara, 1956 nec

Esben-Petersen, 1912 Micromus cookeorum (Zimmerman, 1946)

= Pseudopsectra cookeorum Zimmerman, 1946 Micromus costulatus (Motschulsky , 1863)

=Micromerus costulatus Motschulsky, 1863 Micromus delamarei (Auber, 1956)

=Eumicromus delamarei Auber, 1956 Micrornus dispar Banks, 1910 Micromus dissimilis (Nakahara, 1915) nov.comb.

= Eumicromus dissimilis Nakahara, 19 15 Micromus distinctus (Perkins, 1899) nov.comb.

= Nesomicromus distinctus Perkins. 1899a

TANZANIA KENYA CHINA

HAW AI1 HAWAII

AUSTRIA ENGLAND LAPONIA BELGIUM SARDINlA AUSTRIA

USA

HAWAII ST. HELENA ISL.

CEY LON

HAWAII HAWAII HAWAII HAWAII

CHINA NEW ZEALAND NEW ZEALAND NEW ZEALAND

HAWAII CEY LON

CANARY ISLANDS

CANARY ISLANDS BELGIAN CONGO

JAPAN

JAPAN

HAWAII

CEYLON

IVORY COAST COLUMBIA

JAPAN

HAWAII

Page 14: A systematic checklist of the Hemerobiidae of the world

V. J . Monserrcit

Micromus drepanoides (Perkjns, 1899) nov.comb. = Nesomicromus drepanoides Perkins, 1899a HAWAII

Micromus duporti Lestage, 1928 INDOCHINA Micromus exiguus NavBs, 1908a CANARY ISLANDS Micromus falcatus Banks, 1910 nec Zimmerman, 1957 COLUMBIA Micromus falcatus (Zimmerman, 1957) nov.comb. nec Banks, 1910

= Nesomicromus falcatus Zimmerman, 1957 HAWAII Micromus forcipatus (Perkins, 1899) nov.comb.

= Nesomicromus forcipatus Perkins, 1899a HAWAII Micromus formosanus (Kruger, 1922)

= Archaeomicromus formosanus Kruger, 1922 nom.nud.

Micromus fractus Navas, 1914c CEYLON Micromus fulvescens (Perkins, 1899) nov.comb.

= Nesomicromus fulvescens Perkins, 1899a HAWAII Micromus fuscatus (Nakahara, 1965) nov.comb.

=Pseudomicromus fuscatus Nakahara, 1965a MEXICO Micromus fuscotriatus (Fraser, 1951)

=Eumicromus fuscotriatus Fraser, 1951 MADAGASCAR Micromus gradatus Navas, 1912c CORSICA Micromus gratus Banks, 1937a PHILIPPINES Micromus haleakalae (Perkins, 1899)

= Nesomicromus haleakalae Perkins, 1899a HAWAII = Nesomicromus stenopteryx Perkins, 1899a HAWAII

Micromus haleakalae (Perkins, 1899) = Nesothauma haleakalae Perkins, 1899a HAWAII

Micromus igorotus Banks, 1920 PHILIPPINES = Eumicromus okinawanus Nakahara, L956 JAPAN

Micromus insulanus Navas, 1925b MAURITIUS Micromus insularis Hagen, 1886 MADAGASCAR Micromus jacobsoni Esben-Petersen, 1926 SUMATRA Micromus japonicus (Kruger, 1922)

= Archaeomicromus japonicus Kriiger , 1922 nom.nud.

Micromus javanus Kruger, 1922 nom.nud. Micromus kapuri (Nakahara, 1971) nov.comb.

= Eumicromus kapuri Nakahara, 197 1 Micromus lanceolatus Navas, 1910b Micromus lanosus (Zeleny, 1962)

= Stenomicromus lanosus Zeleny , 1962 Micromus latipennis (Perkins, 1899) nov.comb.

= Nesomicromus latipennis Perkins, 1899a Micromus laxus Navis, 1930a Micromus linearis (Hagen, 1858)

= Micromerus linearis Hagen, 1858b Micromus lobipennis (Perkins, 1899)

=Pseudopsectra lobipennis Perkins, 1899a Micromus longispinosus (Perkins, 1899) nov.comb.

= Nesomicromus long ispinosus Perkins, 1899a Micromus lorianus (Navas, 1929) nov.comb.

=Tanca loriana NavBs, 1929e

INDIA MADAGASCAR

CZECHOSLOVAKIA

HAWAII CONGO

CEYLON

HAWAII

HAWAII

NEW GUINEA

Page 15: A systematic checklist of the Hemerobiidae of the world

World catalogue of Hemerobiidae

Micromus maculatipes (Nakahara, 191 5) nov.comb. = Eumicromus maculatipes Nakahara, 19 15 = Micromus pictipes Banks, 1920

Micromus maculipes (Fraser , 1957) = Eumicromus maculipes Fraser, 1957

Micromus malgassicus (Fraser, 195 1) = Eumicromus malgassicus Fraser, 195 1

Micromus marquesanus (Kimmins, 1932) nov.comb. = Nesomicromus marquesanus Kimmins, 1932

Micromus meridionalis (Costa, 1855) = Mucropalpus meridionalis Costa, 1855 nom .dub.

Micromus minimus (Perkins, 1899) nov.comb. = Nesomicromus minimus Perkins, 1899a

Micromus montanus Hagen, 1886 Micromus morosus Gerstaecker, 1894 Micromus multinervosus (Fraser, 1955)

=Eumicromus multinervosa Fraser, 1955 Micromus multipunctatus Matsumura, 1907

= Micromus nngustior Weele, 19 1 0a = Micromus novitius Navis, 1910a =Nenus longulus Navis, 19 12a = Nenus luzonensis Navis, 19 18b

Micromus neocaledonicus (Nakahara, 1960) nov.comb.

=Eumicromus neocaledonicus Nakahara, 1960b Micromus nigrifrons Banks, 1937a Micromus nilghiricus Navis, 1910b Micromus numerosus Navris, 1910a

= Eumicromus arakawae Nakahara, 19 15 Micromus obliquus (Kriiger , 1922)

= Meomicromus obliquus Kriiger, 1922 nom.nud. Micromus oblongus Kimmins, 1935b

=Nenus parallelus Navis, 1936e Micromus ombrias (Perkins, 1899) nov.comb.

= Nesomicromus ombrias Perkins, 1899b Micromus paganus (Linnaeus, 1767)

=Hemerobius paganus Linnaeus, 1767 =Hemerobius nemoralis Stephens, 1836 part. = Micromus lineosus Rambur, 1842 = Hemerohius eleguns Goszy , 1852 nec Stephens,

1836 = Eumicromus alpinus Nakahara, 1915

Micromus paganus var. ustulatus Heyden, 1896 Micromus paradoxus (Perkins, 1899) nov.comb.

=Nesomicromus paradoxus Perkins, 1899a Micromus parallelus (Fraser, 1957)

=Eumicromus parallelus Fraser, 1957 Micromus pcrelegans Tjeder, 1937 Micromus placidus Banks, 1937a Micromus plagatus Navas, 1934a Micromus posticus (Walker, 1853)

= Hemerobius posticus Walker, 1853

JAPAN PHILIPPINES

REUNION ISLAND

MADAGASCAR

MARQUESAS ISLANDS

ITALY

HAWAII USA

JAVA

MADAGASCAR JAPAN

JAVA JAPAN CHINA

PHILIPPINES

NEW CALEDONIA PHILIPPINES

INDIA JAPAN JAPAN

SOUTH AFRICA KENYA

HAWAII

ENGLAND ?

AUSTRIA JAPAN

GERMANY

HAWAII

REUNION ISLAND CHINA

PHILIPPINES MADAGASCAR

USA

Page 16: A systematic checklist of the Hemerobiidae of the world

= Micromus insipidus Hagen, 186 1 = Micromus sobrius Hagen, 1861

Micromus ramosus Navas, 1934b Micromus remiformis Oswald, 1987 Micromus rubrinervis (Perkins, 1899) nov.comb.

= Nesomicromus rubrinervis Perkins, 1899a Micromus sabulosus Navas, 1908a Micromus sjostedti Weele, 1910b

= Micromus capensis Esben-Petersen, 1920a = Micromus braunsi Navas, 1929f

Micromus subanticus (Walker, 1853) =Hemerobius subanticus Walker, 1853 = Micromus angustus Hagen, 1 8 86 = Micromus nesoticus Navris, 19 14d

Micromus subochraceus (Perkins, 1899) nov.comb. = Nesomicromus subochraceus Perkins, 1899a

Micromus sumatrunus (Kriiger , 1922) = Archaeomicromus sumatranus Kriiger, 1922 nom.nud.

Micromus swezeyi (Zimmerman, 1940) = Pseudopsectra swezey i Zimmerman, 1940

Micromus tasmaniae (Walker, 1860) = Hemerobius tasmaniae Walker, 1860 = Micromus australis Froggatt, 1904 nec Hagen,

1858b = Micromus froggatti Banks, 1909 = Micromus perkinsi Banks, 1939

Micromus tasmaniae var. manapouriensis Tillyard, 1923 Micromus tasmaniae var. nigroscriptus Tillyard, 1923 Micromus tessellatus Gerstaecker, 1888 Micromus theryanus Navas, 19 lob Micromus timidus Hagen, 1853

= Micromus navigatorum Brauer, 1867 = Micromus vinaceus Gerstaecker , 1885 = Micromus pusillus Gerstaecker, 1894 = Micromus suuteri Esben-Petersen, 19 12 nec

Nakahara, 1956 =,Micromus weryiae Lestage, 1924 =Micromus philippinus Navas, 1926b = Archaeomicromus modestus Navas, 1936b = Eumicromus diminutus Nakahara, 1956

Micromus umbrosus Navis, 1930c Micromus usingeri (Zimmerman, 1940)

= Pseudopsectra usingeri Zimmerman, 1940 Micromus vagus Perkins, 1899a *Micromus variegatus (Fabricius, 1793)

= Hemerobius variegatus' Fabricius, 1793 = Micromus pulchellus Nakahara, 1915

Micromus variolosus Hagen, 1886 Micromus xia Yang, 1981b Micromus yunnanus (Navas, 1922) nov.comb.

= Phlebiomus yunnanus Navas, 1922a Micromus zonarius Navas, 1936c

NORTH AMERICA NORTH AMERICA

CHINA USA

HAWAII INDIA

TANZANIA SOUTH AFRICA SOUTH AFRICA

USA USA USA

HAWAII

HAWAII

TASMANIA

AUSTRALIA

AUSTRALIA NEW ZEALAND NEW ZEALAND

BRAZIL ALGERIA

MOCAMBIQUE SAMOA

AUSTRALIA JAVA

BELGIAN CONGO PHILIPPINES

CONGO JAPAN INDIA

HAWAII HAWAII

FRANCE JAPAN

USA CHINA

CHINA MADAGASCAR

Page 17: A systematic checklist of the Hemerobiidae of the world

WorM catalogue of Hemerobiidue

Mixomicromus Ghosh, 1977 *Mixomicromus lampus Ghosh, 1977

Monorobius Makarkin, 1985b

Mucropalpus Rambur, 1842

Nefasitus Navas, 19 15c

Nenus Navas, 1912a

Neomicromus Kriiger, 1922

Neosympherobius Kimmins, 1929b *Neosympherobius cinereus Kimmins, 1929b

Nesbe Navb , 1929d *Nesbe irrebita Navis, 1929d

Nesobiella Kimmins, 193% *Nesobiella hospes (Perkins, 1899)

= Megalomus hospes Perkins, 1899a Nesomicromus Perkins, 1899a

Nesothauma Perkins, 1899a

Neuronema McLachlan, 1869 =Ninguta Navas, 1912d nec Moore, 1892

(Lepidoptera) =Ningu Navis, 1913e = Kulinga Navas, 1936d

Neuronema albadelta Y ang, 1964 Neuronema albostigma (Matsumura, 1907)

=Hemerobius albostigma Matsumura, 1907 = Megulomus deltoides Navas, 19 10a part.

Neuronema anzobica Makarkin, 1986 Neuronema assamensis Kimmins, 1943

= Hemerobius decisus Walker, 1860 part. Neuronema chungnanshana Yang, 1964 *Neuronema decisum (Walker, 1860)

= Hernerobius decisus Walker, 1860 part. Neuronema jluva Kriiger, 1922 nom.nud. Neuronema huangi Yang, 198 1 b Neuronema indica NavBs, 1927a

= Neuronema indicus Navas, 1927a Neuronema irrorata Kimmins, 1943

= Hemerohius decisus Walker, 1860 part. Neuronema kulinga Yang, 1964 Neuronema kuwayamai Nakahara, 1960a Neuronema kwanshiensis Kimmins, 1943 Neuronema laminata Tjeder, 1937 Neuronema laminata ssp. choui Yang, 1964 Neuronema laminata ssp. jilinensis Yang, 1964 Neuronema laminata ssp. tsinhnga Yang, 1964 Neuronema liana Yang, 1986

INDIA see Hemerobius Linnaeus, 1758 see Hcmerobius Linnaeus, 1758 see Sympherobius Banks, 1904 see Micromus Rambur, 1842 sec Micromus Rambur, 1842

ARGENTINA

COSTA RICA

HAWAII see Micromus Rambur, 1842 see Micromus Rambur, 1842

CHINA

JAPAN JAPAN

USSR ASSAM

CHINA

HINDOSTAN

CHINA

INDIA?

HINDOSTAN CHINA JAPAN CHINA CHINA CHINA CHINA CHINA CHINA

Page 18: A systematic checklist of the Hemerobiidae of the world

V. J . Monserrat

Neuronema navasi Kimmins, 1943 = Megalomus deltoides Navas, 19 10a part.

Neuronema nepalensis Nakahara, 197 1 Neuronema obscuru Kriiger , 1922 nom-nud. Neuronema omeishana Yang, 1964 Neuronema pielina (Navas, 1936)

=Kulinga pielina Navas, 1936d Neuronema tienmushana Yang, 1964 Neuronema tjederi Kimmins, 1943

=Megalomus deltoides Navas, 1910a part. Neuronema unipunctata Yang, 1964

Ninga Navas, 1913e

Ninguta Navas, 19 12d

Niremherge Navis, 1909a

FORMOSA NEPAL

CHINA

CHINA CHINA

CHINA CHINA

see Neuronema McLachlan, 1869 see Neuronema McLachlan, 1869 see Sympherobius Banks, 1904

Noius Navas, 1929c Noius noumeanus Kimmins, 1958 *Noius oceanicus Navas, 1929c

Nomerobius Navis, 191 5c Nomerobius annulicornis Navas, 1929d *Nomerobius psychodoides (Blanchard, 185 1)

= Megalomus psychodoides Blanchard, 185 1 = Sympherobius marinoratus Navas, 19 10b = Sympherobius modestus Banks, 1910

Nomerobius psychodoides var. connexus (Banks, 1914) = Sympherobius modestus var . connexus Banks, 19 14

Notiobiella Banks, 1909 = Vuja Navas, 1925a =Buxtonia Esben-Petersen, 1928b nec Thomas,

19 14 (Brachiopoda) = Gunchetus Navas, 1929a

Notiobiella africana (Navas, 1929) =Ganchetus africanus Navis, 1929a

Notiobiella barnardi Monserrat, 1984a Notiobiella bella Navas, 1930a Notiobiella brasiliensis Monserrat & Penny, 1983 Notiobiella costalis Banks, 1918 Notiobiella decora Kimrnins, 1929a Notiobiella dentata Monserrat, 1990 Notiobiella fulva (Esben-Petersen, 1928)

=Buxtonia fulva Esben-Petersen, 1928b Notiobiella gloriosa Navis, 1933c Notiobiella hargreavesi Kimmins, 1936 Notiobiclla israeli (Alayo, 1968)

=Sympherobius israeli Alayo, 1968 Notiobiella luisae Monserrat, 1990 Notiobiella maculata Monserrat & Penny, 1983 Notiobiella mariliae Monserrat, 1984a Notiobiella mexicana Banks, 1913 Notiobiella multifurcata Tillyard, 191 6

NEW CALEDONIA OCEANIA, VITI ISL.

COLUMBIA

CHILE ARGENTINA ARGENTINA

PERU

CAMEROON CAMEROON

CONGO BRAZIL

GOLD COAST UGANDA

INDONESIA

SAMOA CHINA

SIERRA LEONE

CUBA INDONESIA

BRAZIL CAMEROON

MEXICO AUSTRALIA

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World cutulogue c$ Hernerobiidue

Notiobiella nitidula Navas, 1910b Notiobiella ochracea Nakahara, 1966 Notiobiella paddiae Monserrat, 1984b Notiobiella peterseni Banks, 1932 Notiobiella pinarensis (Alayo, 1968)

=Sympherobius pinarensis Alayo, 1968 Notiobiella punctata Tjeder, 1961 Notiobiella rosea Kimmins, 1933 Notiobiella rubrostigma Navas, 19 14f

= Notiobiella cullunguna Kimmins, 1940 Notiobiel la semeriai Monserrat, 1984a Notiobiella spinosa Monserrat & Penny, 1983 Notiobiella stellata Nakahara, 1966 Notiobiella stigmatica Banks, 1909 Notiobiella subolivacea Nakahara, 19 15 Notiobiella turnidu (Navas, 1925)

= Vuju turnidu Navas, 1925a nom.dub. Notiobiella turneri Kimmins, 1933 Notiobiella ugandensis Kimmins, 1939 *Notiobiella unita Banks, 1909 Notiobiella valida Banks, 1920 Notiobiella vicina Kimmins, 1936 Notiobiella viridinervis Banks, 191 3 Notiobiella viridis Tillyard, 1916

Nusalala Navas, 191 3b = Huurupiellu Esben-Petersen, 19 14

Nusalala andina Penny & Sturm, 1984 =Nusalala andinus Penny & Sturm, 1984

Nusalala camposina Navas, 1928b Nusalala championi Kimmins, 1937 Nusalala colombiensis (Banks, 1910)

= Boriomyia colombiensis Banks, 19 10 Nusalala cubana (Hagen, 1886)

=Micromus cubanus Hagen, 1886 Nusalala damiensis Smith, 193 1 *Nusalala erecta Navas, 191 3b Nusalala escomeli Navas, 1922c Nusalala falcata Kimmins, 1940 Nusalala gradata (Navas, 1925) nec 1930b

=Haarupiella gradata Navas, 1925e Nusalala impudica (Gerstaecker, 1888)

= Hemerobius impudicus Gerstaecker, 1888 Nusalala infirma Navas, 19 14e Nusalala krugeri Nakahara, 1965a Nusalala marginata Navas, 1926a Nusalala navasi Kimmins, 1937

=Nusalala gradata Navhs, 1930b nec 1925e Nusalala neotropica (Esben-Petersen, 1914)

= Haarupiella neotropica Esben-Petersen, 19 14 Nusalala pallida (Esben-Petersen, 19 14)

= Haarupiella pallida Esben-Petersen, 19 14 Nusalala reticulata (Navas, 19 10)

KENYA FORMOSA

PARAGUAY MADAGASCAR

CUBA NATAL

SEYCHELLES PARAGUAY

PERU CAMEROON

MEXICO RYUKYU ISLANDS

AUSTRALIA JAPAN

JAVA SOUTH AFRICA

UGANDA AUSTRALIA

BORNEO CONGO

CEYLON AUSTRALIA

COLUMBIA ECUADOR

GUATEMALA

COLUMBIA

CUBA HAITI

ECUADOR PERU

BRAZIL

BRAZIL

BRAZIL JAMAICA

MEXICO COSTA RICA

SIBERIA

BRAZIL

ARGENTINA

Page 20: A systematic checklist of the Hemerobiidae of the world

=Micromus reticulatus Navis, 1910b Nusalala rhegmatica Navas, 19 14f Nusalala uncata Kimmins, 1937 Nusalala uruguaya (Navas, 1922)

= Haarupiella uruguaya Navis, 1922d Nyrma Navis, 1933a

*Nyrma kervillea Navas, 1933a Oedobius Nakahara. 19 15

Oxybiella Tillyard, 1916 *Oxybiella bridwelli Tillyard, 1916

Palaeomicromus Kriiger , 1922 nom.nud. Palaeomicromus schmidti (Kriiger , 1922)

= Hemerobius schmidti Kriiger, 1922 nom.nud. Palmobius Needham, 1905

Paramicromus Nakahara , 19 19 nec K riiger , 1922

Paramicromus Kriiger, 1922 nec Nakahara, 1919

Phlebiomus Navas, 1922a

Phlebonerna Kriiger, 1922

Pirionus Navas, 1928b

Pleomegulomus Kriiger, 1922

Porter Navas, 1924c Psectra Hagen, 1866

=Annandalia Needham, 1909 = Eucurobius Esben-Petersen, 1928b

Psectra affinis (Banks, 1913) nov. comb. = Notiobiella affinis Banks, 19 13

Psectra capensis (Kimmins, 1935) = Annandalia capensis Kimmins, 1935b

Psectra decorata (Nakahara, 1966) nov.comb. = Annandalia decorata Nakahara, 1966

*Psectra diptera (Burmeister, 1839) = Hemerobius dipterus Burmeister , 1839 = Hemerobius delicatulus Fitch, 1855 = Psectra buenoi Navas, 1909b = Notiobiella galloisi Navas, 1924b

Psectra externa (Banks, 1909) = Notiobiella externa Banks, 1909

Psectra fasciata (Esben-Petersen, 1928) =Eucarobius fasciatus Esben-Petersen, 1928b

Psectra franzeni (Kimmins, 1940) = Annandalia franzeni Kimmins, 1940

Psectra graeffei (Brauer , 1867) = Hemerobius graeffei Brauer, 1867

Psectra hageni (Banks, 1932) nov.comb.

BRAZIL PARAGUAY

PANAMA

URUGUAY

ASIA MINOR see Drepanepteryx Leach, 1815

HONOLULU

see Sympherobius Banks, 1904 see Micromus Rambur, 1842 see Micromus Rambur, 1842 see Micromus Rambur, 1842 see Drepancptcryx Leach, 1815 see Megalomus Rambur, 1842 see Megalomus Rambur, 1842 see Gayomyia Banks, l913

PHILIPPINES

SOUTH AFRICA

FORMOSA

GERMANY USA USA

JAPAN

AUSTRALIA

SAMOA

AUSTRALIA

SAMOA

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World catalogue of Hemerobiidcre

= Annandalia hageni Banks, 1932 Psectra iniqua (Hagen, 1859)

= Hemerobius iniquus Hagen, 1859 = Annandalia curta Needham, 1909 = Notiobiella maindronina Navas, 19 lob = Notiobiella khandalensis Navas, 1930d

Psectra irregularis (Carpenter, 1961) nov.comb. =Annandalia irregularis Carpenter, 1961

Psectra jeanneli (Navas, 19 14) = Notiobiella jeanneli Navas, 19 14b = Notiobiella congensis Navas, 193 1a =Notiobiella antennuta NavAs, 193 1b

Psectra maculosa (Carpenter, 196 1) nov.comb. = Annandalia maculosa Carpenter, 1961

Psectra minima (Banks, 1920) = Notiobiella minima Banks, 1920

Psectra mombassina (Navas, 1936) = Notiobiella mombassina Navas , 1936b

Psectra mozambica Tjeder, 1961 Psectra obliqua (Banks, 1909)

= Notiobiella obliqua Banks, 1909 Psectra oblonga (Esben-Petersen, 1928)

=Eucarobius oblongus Esben-Petersen, 192813 Psectra oblonga var. brunnea Esben-Petersen, 1928

=Eucarobius oblongus var. brunneus Esben-Petersen, 1928b

Psectra pretiosa (Banks, 1909) nov.comb. =Notiobiella pretiosa Banks, 1909

Psectra siamica Nakahara & Kuwayama, 1961 Psectra yunu Yang, 1981a

Pseudomicrornus Kruger , 1922

Pseudopsectra Perkins, 1899a

Psychobiella Banks, 1909 Psychobiella fusca Tillyard, 1916 *Psychobiella sordida Banks, 1909

Reuterobius Kruger , 1922

Schneiderobius Kriiger, 1922

Semohemerobius Yang, 1983 = Mesohemerobius Nakahara, 1966 nec Ping,

1928 (+) *Semohemerobius subacutus (Nakahara, 1966)

=Mesohemerobius subacutus Nakahara, 1966 Sineuronerna Yang, 1964

Sineuronerna bomeana Yang, 198 1b Sineuronema gy irongana Yang, 198 1 b Sineuronema Inagmangana Yang, 198 1 b Sineuronen~a ny ingchiana Yang, 1981 b Sineuronema quxamana Yang, 198 1b

CEYLON

CEYLON INDIA INDIA INDIA

MICRONESIA

KENYA CONGO CONGO

MICRONESIA

BORNEO

CONGO MOCAMBIQUE

AUSTRALIA

SAMOA

SAMOA

FIJI ISLANDS THAILAND

CHINA see Micromus Rambur, 1842 see Micromus Rambur, 1842

AUSTRALIA AUSTRALIA

see Hemerobius Linnaeus, 1758 sec Hemerobius Linnaeus, 1758

FORMOSA

CHINA CHINA CHINA CHINA CHINA

Page 22: A systematic checklist of the Hemerobiidae of the world

Sineuronerna shensiensis Yang, 1964 CHINA Sineuronema similis (Banks, 1940)

= Neuronema sirnilis Banks, 1940 CHINA "Sineuronerna sinensis (Tjeder, 1937)

= Neuronerna sinensis Tjeder, 1937 CHINA Sineuronerna yadongana Yang, 1981b CHINA Sineuronerna yunica Yang, 1986 CHINA Sineuronerna zharnana Y ang , 198 1 b CHINA

Spudobius Needham, 1905 see Sympherobius Banks, 1904

Spilomicromus Nakahara, 1960a see Micrornus Rarnbur, 1842

Spinomegalomus Nakahara, 196% see Megalornus Rarnbur, 1842

Stenolomus NavBs, 1906 see Hernerobius Linnaeus, 1758

Stenomicromus Kruger, 1922 see Micromus Rarnbur, 1842

Stenornus Navis, 1922b nec Rafinesque, 18 15 (Coleoptera) *Stenornus nesaeus Navis, 1922b MADAGASCAR

Subboriornyia Steinmann, 1967 *Subboriornyia fusca Steinmann, 1967 HUNGARY

Syrnpherobius Banks, 1904 = Spadobius Needharn, 1905 = Palmobius Needharn, 1 905 = Niremberge Navas, 1909a = Nefasitus Navas, 19 15c = Coloma Navas, 19 15c = Eurobius Kruger, 1922 = Luchlanius Kruger, 1922

Syrnpherobius arnazonicus Penny & Monserrat, 1985 *Syrnpherobius amiculus (Fitch, 1855)

=Hernerobius arniculus Fitch, 1855 =Sympherobius buenoi Navas, 1912e

Syrnpherobius angustus (Banks, 1904) = Hernerobius angus tus Banks, 1904c =Sympherobius tristis Navas, 19 14d

Sympherobius ariasi Penny & Monserrat, 1985 Sympherobius arizonicus Banks, 191 1 Syrnpherobius axillaris Navas, 1927c Sympherobius barberi (Banks, 1903)

= Hemerobius barberi Banks, 1903 Sympherobius bearneri Gurney , 1948 Sympherobius bifasciatus Banks, 191 1 Sympherobius bisignatus (Kruger, 1922)

=Spadobius bisignatus Kruger, 1922 nom.nud. Sympherobius blanchardi (Navis, 1929)

= Colorna blanchardi Navas, 1929b Syrnpherobius brunneus Nakahara, 196% Sympherobius californicus Banks, 19 1 1 Sympherobius dilutus Nakahara, 1960a Syrnpherobius distinctus Carpenter, 1940

BRAZIL

USA USA

USA USA

BRAZIL USA

MEXICO

USA USA USA

CHILE USA USA

JAPAN USA

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World catalogue of Hemerobiidae

Sympherobius doniesticus Nakahara, 1954 JAPAN Sympherobius elegans (Stephens, 1 836)

=Hemerobius elegans Stephens, 1836 nec Goszy, 1852 ENGLAND = Hemerobius murshami Stephens, 1836 ENGLAND = Hemerobius beckwithii Stephens, 1836 ENGLAND = Hemerobius paucinervis Zetterstedt, 1 840 LAPONIA = Hemerobius striatellus Klapilek, 1905 TRANSYLVANIAN ALPS = Sympherobius venustus Navas, 1908b SPAIN = Sympherobius vicenfei Navas, 19 14a SPAIN = Nefasitus catalaunicu.s Navas, 1 930e SPAIN

Sympherobius fallax Navas, 1908a CANARY ISLANDS = Sympherobius schmitzi Navas, 1908a MADEIRA = Sympherobius tenellus Navis, 1908b SPAIN =Sympherobius amicus Navas, 19 15a ERYTREA =Sympherobius sanctus Tjeder, 1939 PALESTINE

Sympherobius fuscescens (Wallengren, 1863) = Hemerobius fuscescens Wallengren, 1863 SWEDEN = Hemerobius pellucidus (Walker, 1853):

Wallengrcn, 1871 = Hemerobius obscurellus Zetterstedt, 1840 LAPONIA = Hemerobius inconspicuus McLachlan, 1868 ENGLAND = Niremberge limpida Navas, 1909a SPAIN

Sympherobius fuscinervis Kozhantshikov, 1956 USSR Sympherobius gayi Navas, 1910d CHILE Sympherobius gracilis Carpenter, 1940 USA Sympherobius gratiosus Navhs, 190% SPAIN Sympherobius humilis Navas, 19 14f PARAGUAY Sympherobius innoceus Steinmann, 196% ARGENTINA Sympherobius insulanus Banks. 1938 CUBA Sympherobius intervenalis Banks, 19 14 COLUMBIA Sympherobius killingtoni Carpenter, 1940 USA Sympherobius klapaleki Zclenf , 1963 CZECHOSLOVAKIA

=Sympherobius hyalinus Monserrat, 1976b SPAIN = Syrnpherobius outereloi Monserrat, 1976a SPAIN

Sympherobius limbus Carpenter, 1940 USA Sympherobius luojiaensis Yang, 1980b CHINA Sympherobius maculipennis Kimmins, 1929b ARGENTINA Sympherobius manchuricus Nakahara, 1960a MANCHURIA Sympherobius marmoratipennis (Blanchard, 185 1)

= Megalomus marmoratipennis Blanchard, 185 1 CHILE Sympherobius matsucocciphagus Yang, 1980b CHINA Sympherobius miranda (Navas, 1920)

=Niremberge miranda NavBs, 1920 ARGENTINA = Sympherobius molinarii Nakahara, 1960a ARGENTINA

Sympherobius notatus Kimmins, 1932 WEST INDIES Sympherobius occidentalis (Fitch, 1855)

=Hemerobius occidentalis Fitch, 1855 USA Sympherobius parvus (Kriiger, 1922)

=Spadobius parvus Kriiger, 1922 nom.nud. Sympherobius pellucidus (Walker, 1853)

= Hemerobius pellucidus Walker, 1853 nec Wallengren, 187 1 ENGLAND

Page 24: A systematic checklist of the Hemerobiidae of the world

=Sympherobius carpaticus Kis, 1965 Sympherobius perparvus (McLachlan, 1869)

= Hemerobius perparvus McLachlan, 1869 = Sympherobius sparsus Banks, 19 1 1

Sympherobius pictus (Banks, 1904) = Hemerobius pictus Banks, 1904a

Sympherobius pupillus Navas, 1915f nondub. Sympherobius pygmaeus (Rambur, 1842)

= Mucropalpus pygmaeus Rambur, 1842 = Hemerobius coccophugus Goszy , 1852 nom .dub. = Micromus pumilio Stein, 1863 =Sympherobius venosus Navis, 1908b =Sympherobius conspersus NavBs, 1908b = Sympherobius lambereti Navas, 19 lob = Sympherobius bellus Navas, 19 1 1 = Sympherobius menendezi Navas, 19 13c = Sympherobius melanogaster Navis, 19 15d =Sympherobius fortini Lacroix, 1924 = Nefasitus italicus Navas, 1932e = Sympherobius laetus Steinmann, 1967

Sympherobius riudori Navas, 1915e Sympherobius scriptus (Navas, 1917)

=Coloma scriptus Navas, 1917c Sympherobius signatus (Kruger , 1922)

=Spadobius signatus Kruger, 1922 nom.nud. Sympherobius similis Carpenter, 1940 Sympherobius smitheri Nakahara, 1960a

= Sympherobius brincki Tjeder, 196 1 = Sympherobius impar Tjeder, 196 1 =Sympherobius nigricornis Tjeder, 1961

Sy mpherobius stangei Nakahara, 1960a Sympherobius subcostalis Monserrat, 1990 Sympherobius tessellatus Nakahara, 19 15

=Sympherobius okinawensis Kuwayama, 1964 Sympherobius texanus Nakahara, 1965b Sympherobius thaumasta Navas, 1915 cf. Penny, 1977 Sympherobius tuomurensis Yang, 1985 Sympherobius umbratus (Banks, 1903)

=Hemerobius umbratus Banks, 1903 Sympherobius weisong Yang, 1980b Sympherobius wuyianus Yang, 1981a Sympherobius yunpinus Yang, 1986 Sympherobius zelenyi Alayo, 1968

Sympheromima Kimmins, 1928 *Sympheromima marginata Kimmins, 1928

Tanca Navas, 1929e Vaja Navis, 1 9 2 5 Wesmaelius Kriiger, 1922

= Boriomyia Banks, 1 9 0 5 nec 1904b = Kimminsia Killington, 1937

Wesmaelius acuminatus (Yang, 1980) nov.comb. = Kimminsia acuminata Yang, 1980a

ROMANIA

USA USA

USA MEXICO

FRANCE AUSTRIA GREECE

SPAIN SPAIN

FRANCE SPAIN SPAIN SPAIN

FRANCE ITALY

HUNGARY SPAIN

ARGENTINA

USA RHODESIA

SOUTH AFRICA RHODESIA RHODESIA

USA MEXICO

JAPAN JAPAN

USA BRAZIL CHINA

USA CHINA CHINA CHINA CUBA

GUATEMALA see Micromus Rambur,1842 see Notiobiella Banks, 1909

CHINA

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World catalogue of Hemerobiidue

Wesmaelius altissimus (Ohm, 1967) =Boriomy ia altissirna Ohm, 1967

Wesmaelius amseli (Aspiick & Aspock, 1966) =Boriomyia amseli Aspock & Aspock, 1966 = Kimminsia pekinensis Yang . 1980a

Wesmaelius asiaticus Yang, 1980a Wesmaelius baikalensis (NavQs, 1929) nov.comb.

=Hemerobius (Boriomyia) baikalensis NavQs, 1929c Wesmaelius balticus (Tjeder, 193 1)

=Boriomyia baltica Tjeder, 193 1 Wesmaelius barnardi (Tjeder, 1955) nov.comb.

=Boriomyia barnardi Tjeder, 1955 Wesmaelius bihamitus (Yang, 1980) nov.comb.

=Kimminsia bihamita Yang, 1980a Wesmaelius brunneus (Banks, 1920)

= Boriomy ia brunnea Banks, 1920 Wesmaelius coloradensis (Banks, 1897)

= Henierobius coloradensis Banks, 1897 "Wesmaelius concinnus (Stephens, 1836)

= Hemerobius concinnus Stephens, 1836 = Hemerobius cylindripes Wesmael , 1 84 1 = Hemerobius utomarius Goszy , 1852

Wesmaelius constrictus (Parfin, 1956) =Kimminsia constricts Parfin, 1956 = Kimminsia olympica Nakahara, 196%

Wesmaelius cunctatus (Ohm, 1967) =Boriomyia cunctata Ohm, 1967

Wesmaelius davidicus (Navas, 1910) nov.comh. = Boriomy ia davidica Navas, 19 10c

Wesmaelius exoticus Makarkin, 1986 Wesmaelius fassnidgei (Killington, 1933)

=Boriomyia fassnidgei Killington, 1933 Wesmaelius fulvus (Navgs, 19 19) nov.comb.

= Hemerobius fulvus Navas, 19 19a Wesmaelius fumatus (Carpenter, 1940)

=Kimminsia funiata Carpenter, 1940 Wesmaelius fumosus (Tjeder, 1955) nov.comb.

=Boriomy ia futnosa Tjeder, 1955 Wesmaelius furcatus (Banks, 1935)

=Boriomyia furcata Banks, 1935 Wesmaelius geyri (Esben-Petersen, 1920) nov.comb.

= Boriomy ia geyri Esben-Petersen, 1920b Wesmaelius griseus (Zeleny, 1971)

= Boriomyia grisea Zeleny , 197 1 Wesmaelius hani (Yang, 1985) nov.comb.

=Kimminsia hani Yang, 1985 Wesmaelius helveticus (Aspock & Aspock, 1964)

=Boriomyia helvetica Aspock & Aspiick, 1964a Wesmaelius ignoratus Steinmann, 1967 Wesnlaelius involutus (Carpenter, 1940)

=Kimminsia involuta Carpenter, 1940 Wesmaelius kaszabi (Steinmann, 1965)

NEPAL

AFGHANISTAN CHINA CHINA

TURAN

SWEDEN

SOUTH AFRICA

CHINA

USA

USA

ENGLAND BELGIUM AUSTRIA

ALASKA USA

ITALY

PHILIPPINES USSR

FRANCE

ALGERIA

USA

SOUTH AFRICA

USA

ALGEKIA

BULGARIA

CHINA

SWITZERLAND HUNGARY

USA

Page 26: A systematic checklist of the Hemerobiidae of the world

V. J . Monserrat

= Boriomy ia kaszabi Steinmann, 1965a Wesmaelius koreanus (Kruger, 1922)

= Boriorny ia koreana Kruger, 1922 nom.nud. Wesmaelius kozlovi Makarkin, 1984 Wesmaelius lateralis (Navas, 1912) nov.comb.

=Boriomy ia lateralis Navas, 1912d Wesmaelius lindbergi (Esben-Petersen, 1931)

= Boriomy ia lindbergi Esben-Petersen, 193 1 Wesmaelius longifrons (Walker, 1853)

=Hemerobius longifrons Walker, 1853 = Hemerobius alternatus Fitch, 1855 = Hemerobius transversus Banks, 1904a = Allotomyiu borealis Banks, 1935

Wesmaelius longipennis (Banks, 1920) =Boriomyia longipennis Banks, 1920

Wesmaelius magnus (Kimmins, 1928) nov.comb. =Boriomyia magna Kimmins, 1928

Wesmaelius majusculus (Kimmins, 1959) nov.comb. =Kimminsia majuscula Kimmins, 1959

Wesmaelius malladai (Navis, 1925) =Hemerobius malladai Navas, 192% = Boriomyiu mortoni (McLachlan, 1899): Killington,

1937 = Kimminsia killingtoni Fraser, 1942

Wesmaelius mongolicus (Steinmann , 1965) =Boriomyia mongolica Steinmann, 1965a =Boriomyiu arenata Aspock & Aspock, 1966

Wesmaelius mortoni (McLachlan, 1 899) = Hemerobius mortoni McLachlan, 1899 = Boriomyia enontekiensis Klingstedt, 1929

Wesmaelius navasi (Andreu, 191 1) =Boriomy ia navasi Andreu, 191 1 = Boriomyia persica Morton, 192 1

Wesmaelius neimenica (Yang, 1980) nov.comb. =Kimminsia neimenica Yang, 1980a

Wesmaelius nervosus (Fabricius, 1793) =Hemerobius betulinus Str@m, 1788 nom.dub. = Hemerobius nervosus Fabricius, 1793 = Hemerobius nebulosus Stephens, 1836 part. = Hemerobius conspersus Burmeister, 1839

? = Mucropalpus distinctus Rambur, 1842 =Hemerobius disjunctus Banks, 1897 = Hemerobius ffostinus Navas, 1933b = Kimminsia cinerea Nakahara, 1960a = Kimminsia melaleuca Nakahara , 3 965b = Kimminsia alexanderi Nakahara, 1965b

Wesmaellus nervosus var. neurasthenica (Navas, 1914) = Boriomy ia nervosa var. neurasthenica Navis, 1914g

Wesmaelius nervosus f. melancholica (Killington, 1937) =Boriomyia betulina f. melancholica Killington, 1937

Wesmaelius nubilus (Kimmins, 1929) nov. comb. =Boriomyia nubila Kimmins, 1929a

MONGOLIA

MONGOLIA

SIBERIA

MOROCCO

USA USA USA USA

USA

GUATEMALA

UGANDA

BULGARIA

ENGLAND

MONGOLIA TURKEY

EUROPE LAPLAND

SPAIN PERSIA

CHINA

NORWAY FRANCE

ENGLAND GERMANY

SPAIN USA USA

JAPAN ALASKA ALASKA

USSR

ENGLAND

NATAL

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Wesmaelius obscuratus (Navis, 1936) nov. comb. = Hemerobius (Boriomy ia) obscuratus Navas, 1936e KENYA

Wesmaelius ogatai (Nakahara, 1956) nov. comb. =Kimminsia ogatai Nakahara, 1956 JAPAN = Kimminsia subnebulosa (Stephens, 1836):

Kuwayama, 1956b JAPAN Wesmaelius persimilis (Ohm, 1967)

=Boriomyia persimilis Ohm, 1967 GREECE Wesmaelius pinincolus (Ohm, 1967)

= Boriomyia pinincola Ohm, 1967 CANARY ISLANDS Wesmaelius posticatus (Banks, 1905)

=Boriomyia posticata Banks, 1905a USA Wesmaelius praenubilus (Fraser, 1951) nov.comb.

= Kimminsia praenubila Fraser, 195 1 MADAGASCAR Wesmaelius pretiosus (Banks, 1908)

=Boriomy ia pretiosa Banks, 1908 USA Wesmaelius pseudofurcatus Makarkin, 1986 USSR Wesmaelius quadrifasciatus (Reuter, 1894)

=Hemerobius concinnus var. quadrifasciatus Reuter, 1894 FINLAND

Wesmaelius ravus (Withycombe, 1923) =Boriomyia rava Withycombe, 1923 ENGLAND

Wesmaelius reisseri Aspock & Aspock, 1982 SPAIN Wesmaelius schwarzi (Banks, 1903)

= Hemerobius schwarzi Banks, 1903 USA = Boriomyia ultirna Banks, 1930c USA

Wesmaelius sinicus (Tjeder, 1937) nov.comb. =Boriomy ia sinica Tjeder, 1937 CHINA

Wesmaelius subnebulosus (Stephens, 1836) =Hemerobius subnebulosus Stephens, 1836 ENGLAND = Hemerobius fuscus Stephens, 1836 ENGLAND = Hemerobius nebulosus Stephens , 1836 part. ENGLAND = Boriomyia maorica Tilly ard, 1923 NEW ZEALAND

Wesmaelius subnebulosus var . obscura (Navas, 19 17) = Hemerobius subnebulosus var. obscura Navfis,

1917a SPAIN Wesmaelius subnebulosa var. lucida (Navis, 1919)

=Hemerobius subnebulosus var. lucida Navis, 1919b SPAIN

Wesmaelius subnebulosus f. melancholica (Killington, 1937)

=Boriomy~a subnebulosa f. melancholica Killington, 1937 ENGLAND

Wesmaelius sufuensis Tjeder, 1968 SINKIANG Wesmaelius tjederi (Kimmins, 1963)

= Boriomyia tjederi Kimmins, 1963 YUGOSLAVIA Wesmaelius transsylvanicus (Kis, 1968)

=Boriomyia transsylvanica Kis, 1968 ROMANIA Wesmaelius trivenulatus (Yang, 1980) nov.comb.

=Kimminsia trivenulata Yang, 1980a CHINA Wesmaelius tuofenganus (Yang, 1985) nov.comb.

=Kimminsia tuofengana Yang, 1985 CHINA

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Wesmaelius ulingensis (Yang, 1980) nov.comb. =Kimminsia ulingensis Yang, 1980a

Wesmaelius vartianae (Aspock & Aspock, 1966) nov.comb. = Boriomy ia vartianae Aspock & Aspock, 1966

Wesmaelius yemenicus (Yang, 1980) nov.comb. =Kimminsia yemenica Yang, 1980a

Wesmaelius yukonensis Klimaszewski & Kevan, 1987 Wesmaelius zhiltzovae Makarkin, 1986

Zachobiella Banks, 1920 Zachobiella angusta Kimmins, 1940 Zachobiella hainanensis Banks, 1939 Zachobiella jacobsoni Esben-Petersen, 1926 Zachobiella marmorata Navas, 1926b Zachobiella pallida Banks, 1939 "Zachobiella punctata Banks, 1920 Zachobiella striata Nakahara, 1966 Zachobiella submarginata Esben-Petersen, 1929b

CHINA

AFGHANISTAN

YEMEN CANADA

USSR

AUSTRALIA HAINAN

SUMATRA SUMATRA

AUSTRALIA PHILIPPINES

FORMOSA AUSTRALIA

I wish to express my grateful thanks to Dr M.W. Mansell for correcting the English manuscript.

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World catalogue of Hemerubiidue

INDEX TO TAXA (Synonyms, nomina dubia and nornina nuda in italics)

abditus - 219 abyssinicus - 219 acuminata, Megalomina - 225 acuminatus, Megalomus - 225 acuminatus, Wesmaelius - 238 acunai - 225 acuiTai - 225 acutipennis - 226 adelgivorus - 219 afinis, Hemerobius (Stephens) - ufpnis, Henierobius (Wesmael) - affinis, Psectra - 234 africana, Notiobiella - 232 africanus, Micromus - 227 Afromicromus - 2 17, 226 albadelta - 231 albipennis - 2 19 albostigma - 231 alexanderi - 240 algida - 2 1 8 algonquinus - 221 Allemerobius - 217 AIIotomyla - 2 17, 2 17 alpestris - 219 alpinus - 229 alternans - 227 alterrzatus - 240 altissimus - 239 amabilis - 227 amazonicus - 236 Ameromicromus - 2 17, 226 amiculus - 236 amicus - 237 amseli - 239 amurensis - 219 Anapsectra - 217 andina - 233 andreinii - 225 angularis - 227 angulatus, Mcgalomus - 225 angulatus, Micromus - 227 angusta, Zachobiella - 242 angustatus - 225 angustior - 229 angustipennis - 227 angustus,Carobius - 2 17 angustus, Micromus - 230 angustus, Sympherobius - 236 Annancluliu - 217, 234 annulicornis - 232

Anomicromus - 217, 226 Anotiobiella - 217, 219 urltennatu - 235 antigonus - 2 19 anzobica - 231 aper - 219 uphidivorus - 227 apicalis - 221 arakuwae - 229

222 Archaeomicromus - 2 17, 226 221 arenuta - 240

ariasi - 236 arizonicus - 236 asiaticus - 239 assamensis - 23 l atlanticus - 227 atomurius, Hemerobius - 239 atomarius, Megalomus - 225 atrifrons - 219 audax - 227 australis, Hemerobius - 219 czustrali.s, Micromus (Froggatt) - 230 australis, Micromus (Hagen) - 227 Austromegalomus - 21 7 Austr-omicromus - 21 7, 226 axillaris - 236 axillatus - 225 azoricus - 219

baguiensis - 219 baikalensis - 239 balachowskyi - 225 balticus - 239 barberi - 236 barnardi, Notiobiella - 232 barnardi, Wesmaelius - 239 beameri - 236 beckwithii - 237 bella, Notiobiella - 232 bcllulus - 227 hellus, Sympherobius - 238 benardi - 227 berothoides - 2 18 Bestretu - 217, 218 betulinus - 240 bifasciatus, Micromus - 227 bifasciatus, Sympherobius - 236 bihamitus - 239 bilineata - 218 binigripunctatus - 219

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World ccztalogue of Hetnerohiidae

deceptor - 220 Eutnicrr,rnus - 2 19, 226 decisum, Neuronema - 231 Eurobius - 219, 236 clecisus, Hemerobius - 23 1, 23 1 exceptatus - 220 decisu.~, Neuronema - 23 1, 231 excisa - 218 decora - 232 exiguus - 228 decorata, Psectra - 234 exoterus - 220 ckcoratus, Megalomus - 225 exoticus - 239 decussutus - 225 exterior - 225 delamarei - 227 externa - 234 de1icatulu.s - 234 deltoides, Megalomus - 231, 232, 232 falcata, cayomyia - 219 dentata - 232 falcata, Nusalala - 233 dilutus - 236 falcatus, Micromus (Banks) - 228 diminutus - 230 falcatus, Micromus (Zimmerman) - 228 diptera - 234 falciger - 219 discolor - 2 19 falculoides - 218 discr~tus - 223 fallax - 237 disjunctus - 240 fasciata, Psectra - 234 dispar - 227 fusciutu.~, Hemerobius (Goszy) - 219 disparilis - 220 fascintus, Hemerobius (Stephens) - 223 dissimilis - 227 fassnidgei - 239 distincta, Hemerobius hageni var. - 221 fenestratus - 220 distinctus, Micromus - 227 ferox - 220 distinctus, Mucropalp~~s - 240 fidelis - 217 distinctus, Sympherobius - 236 $mhriutus - 225 divisa, Drepanacra binocula var. - 218 jlrl\.,u - 231 divisus, Hemerobius - 220 flaveolus - 217 domesticus - 237 jle.xuo.sus - 222 domingensis - 220 flinti - 225 dorsatus - 220 forcipatus - 228 Drepanacra - 217 formosanus, Megalonius - 225 Drepanacrella - 21 8 ,fi~rtrzosanus, Micromus - 228 Drepanepteryx - 218 fortini - 238 drepanoides - 228 fractus - 228 Drepanomima - 2 18 franzeni - 234 Drepunopteryx - 2 19, 2 18 friedeli - 224 duporti - 228 froggatti, Drepanacra - 2 17 dyari - 224 froggatti, Micromus - 230 Dyshemerobius - 219 frontalis (Hagen) - 220

frontalis (NavBs) - 220 eatoni - 220 ,fro.rtinus - 240 Egnyonyx - 219, 219 fu.jimotoi - 220 elatus - 220 fulva, Notiobiella - 232 elegans (Goszy) - 229 fulvescens - 228 elegans (Stephens) - 237 fulvus, Wesmaelius - 239 ellenbergeri - 223 fumatus - 239 elongatus - 220 firnosus, Hemerobius - 224 enontekiensis - 240 fumosus, Wesmaelius -- 239 erecta - 233 furcatus - 239 errans - 223 fusca, Hemerobius nitidulus var. - 222 escomeli - 233 fusca, Psychobiella - 235 Eucurobius - 219, 234 fusca, Subboriomyia - 236

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V. J. Monserrat

fuscata, Drepanepteryx - 2 18 fuscatus, Micromus - 228 fuscescens - 237 fuscinervis, Mucropalpus - 222 fuscinervis, Sympherobius - 237 fuscotriatus - 228 fuscus, Hemerobius - 241

galloisi - 234 Gunchetus - 219, 232 gayi - 237 Gayomyia - 219 geyri - 239 gibbosa - 2 19 gilvus - 220 glacialis - 220 gloriosa - 232 gossypii - 221 gracilis - 237 gradata, Nusalala (Navas, 1925) - gradata, Nusalala (Navis, 1930) - gradatus, Micromus - 228 graeffei - 234 grahami - 221 gratiosus - 237 gratus - 228 greeni - 221 griseus, Hemerobius - 222 griseus, Wesmaelius - 239 gyirongana - 235

Haarupiellu - 219, 233 haemuticus - 222 hageni, Hemerobius - 221 hageni, Psectra - 234 Hagenobius - 219, 2 19 hainanensis - 242 haitiensis - 226 haleakalae, Nesomicromus - 228 haleakalae, Nesothauma - 228 handschini - 221 hani - 239 hardyi - 217 hargreavesi - 232 harmandinus - 221 hedini - 221 helveticus - 239 Hemerobiella - 219 Hemerobius - 219 Hemerodomiu - 224, 2 19 hengduanus - 221 hespericus - 221

hesperus - 221 Heteromicromus - 224, 226 hirtus - 225 hopii - 221 hospes - 231 huangi - 231 humilior - 217 humilis, Drepaneptery X - 2 17 humilis, Sympherobius - 237 humuli (Hagen) - 222 humuli (Linnaeus) - 22 1 humuli (Zetterstedt) - 222 humulinus - 221 hyalinatus - 223 hyalinus, Hcmcrobius - 221 hyalinus, Sympherobius - 237

iuponica, Bestreta - 218 Idiomicromus - 224

233 ignoratus - 239 233 igorotus - 228

immuculutus - 22 1 impar - 238 imperfectus - 227 impudica - 233 inconspicuus - 237 incursus - 221 indica, Neuronema - 231 indicus, Hemerobius - 221. Indomicromus - 224, 226 infulcutus - 218 infirma - 233 infumatus - 227 infuscutus - 225 iniqua - 235 innoceus - 237 insignis - 225 insipidus - 230 instabilis - 217 insulanus, Austromegalomus - 2 17 insulanus, Micromus - 228 insulanus, Sympherobius - 237 insularis, Drepanacra binocula v. - 218 insularis, Micromus - 228 intervenalis - 237 intricatus, Hemerobius - 227 intricatus, Megalomus - 225 invcrsus - 221 involutus - 239 irrebita - 231 irregularis, Hemerobius - 222 irregularis, Psectra - 235

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World catalogue of Hemerobiidae

irrorata, Neuronema - 231 irroratus, Hemerobius - 224 irroratus, Mucropalpus - 222 israeli - 232 italicus - 238

jacobsoni, Micromus - 228 jacobsoni, Zachobiella - 242 jamaicensis - 221 japonicus, Hemerobius - 22 1 japonicus, Micromus - 228 juvunus, Hemerobius - 221 javanus, Micromus - 228 jeanneli - 235 jilinensis - 231 jonas - 227 jucundus - 221

kanoi - 224 kapuri - 228 kaszabi - 239 kervillea - 234 khatzdulensis - 235 khasiana - 218 killingtoni, Kimminsia - 240 killingtoni, Sympherobius - 237 kimminsi - 225 Kimminsia - 224, 238 Kimminsiella - 225 klapaleki - 237 kobayashii - 221 kokaneeanus - 221 kollari - 218 kootenuyensis - 223 koreanus - 240 kozlovi - 240 krugeri - 233 Kulirlgu - 225, 231 kulinga - 231 kuscheli - 217 kuwayamai - 231 kwanshiensis - 231

Luchlanius - 225, 236 lacunaris - 221 laetus - 238 lumhereti - 238 laminata - 231 lampus - 231 lunc.rolatus, Megalomus - 2 17 lanceolatus, Micromus - 228 lanosus - 228

lapponicus - 222 lateralis - 240 latipennis - 228 latus - 225 lautus - 222 laxus - 228 lentus - 222 liana - 23 1 lii - 222 limbatellus (McLachlan) - 220 limbatellus (Stitz) - 22 1 limbatellus (Wallengren) - 220, 224 limbatellus (Zetterstedt) - 224 limbatus - 224 limbus - 237 limpida - 237 lindbergi - 240 linearis - 228 lineatus - 227 lineosus - 229 linguatus - 225 lioni - 225 lobipennis - 228 longifrons - 240 longipennis - 240 longispinosus - 228 longitudinalis - 2 18 longulus - 229 lorianus - 228 lucida - 241 ludicrus - 227 luigiorzii - 225 luisae - 232 luojiaensis - 237 lutescens Hernerobius (Burmeister) - 222 lutescens, Hemerobius (Fabricius) - 222 lutescens, Hemerobius (Zetterstedt) - 222 lutescens, Mucropalpus - 22 1 luzonensis - 229

maculata, Conchopterella - 217 maculata, Notiobiella - 232 maculatipes - 228 muculatus, Hemerobius - 221 maculipennis - 237 maculipes - 229 maculosa, Psectra - 235 madeirae - 222 magmangana - 235 magnus -- 240 maindronina - 235 majusculus - 240

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malgassicus - 229 malladai - 240 manapouriensis - 230 manchuricus - 237 mangkamanus - 222 maori - 217 maorica - 241 marginata, Nusalala - 233 marginata, Sympheromima - 238 marginatus, Hemerobius - 222 marginatus, Megalomus - 225 mariliae - 232 marmorata, Zachobiella - 242 marnioratipennis - 237 marmorutus, Sympherobius - 232 marquesanus - 229 rnarshami - 237 matsucocciphagus - 237 medleri - 217 Megaloniina - 225 Megalornus - 225 melaleucu - 240 melancholica, Wesmaelius

nervosus f . - 240 melancholica, Wesmaelius

subnebulosus, f . - 241 melunoguster - 238 melanostictos - 222 menendezi - 238 Mmoptery.~ - 226, 217 ~Menutus - 226 Meomicromus - 226, 226 meridionulis - 229 Mesohemerobius - 226, 235 niexicana, Notiobiella - 232 mexicanus, Hemerobius - 222 micans - 222 Micromus - 226 minima - 235 minimus - 229 minor, Megalomus - 225 minor, Nesomicromus - 227 miranda - 237 Mixomicromus - 231 modestus, Archaeomicromus - 230 modestus, Sympherobius - 232 moestus, Hemerobius (Hagen) - 222 moestus, Hemerobius (Banks) - 224 moestus, Megalomus - 225 molinarii - 237 molokaiensis - 227 mombassina - 235

mongolicus - 240 mononeurus - 222 Monorobius -- 23 1 , 2 19 montanus, Hemerobius - 222 montanus, Micromus - 229 monticellii - 225 monticola - 222 montserraticus - 226 morosus - 229 mortoni (Killington) - 240 mortoni (McLachlan) - 240 mozambica - 235 Mucropalpus - 231, 2 19 multifurcata - 232 multinervosus - 229 multipunctatus - 229

nairobicus - 222 nakaharinus - 221 nanus - 225 natalcnsis - 222 navasi, Megalomus - 225 navasi, Neuronema - 232 navasi, Nusalala - 233 navasi, Wesmaelius - 240 nuvigutorum - 230 neadelphus - 222 nebulosus, Hemerobius - 240, 241 nebulosus, Megalomus - 225 Nefasitus - 23 1 , 236 tzeglectus - 222 neirnenica - 240 nenzoralis - 229 nemorensis - 222 Nenus - 23 1, 226 neocaledonicus - 229 Neomicromus - 23 1 , 226 Neosympherobius - 23 1 neotropica - 233 nepalensis - 232 nervosus - 240 nesaeus - 236 Nesbe - 23 1 Nesobiella - 231 Nesomicromus - 23 1 , 226 Nesothanuma - 23 1, 226 nesoticus - 230 neurasthenia - 240 Neuronema -- 231 nevadensis - 223 nigrans - 222 nigratus - 226

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World catulogue of Hemerobiidae

nigricornis, Hemerobius - 222 nigricornis, Sympherobius - 238 nigrifrons - 229 nigrinus - 224 nigroscriptus - 230 nigrostigma - 222 nilghiricus - 229 Ninga - 232, 231 Ningutu - 232, 231 Niremberge - 232, 236 nitidula, Notiobiella - 233 nitidulus, Hemerobius - 222 Noius - 232 Nomerobius - 232 norfolkiensis - 21 7 notatus - 237 Notiobiella - 232 noualhieri - 226 noumeanus - 232 novitius - 229 nubilus - 240 numerosus - 229 Nusalala - 233 nyingchiana - 235 Nyrma - 234

obliqua, Psectra - 235 obliquu.~, Micromus - 229 obliteratus - 221 oblonga, Psectra - 235 oblongus, Micromus - 229 obscura, Neuronema - 232 obscura, Wesmaelius

subnebulosus var. - 241 obscuratus - 241 obscurellu~ - 237 obscurus, Hemerobius - 221 obscurus, Megalomus - 226 obscurus, Mucropalpus - 222 obtusus - 221 occidentalis - 237 occiduus - 221 oceanicus - 232 ochracea, Notiobiella - 233 ochraceus, Hemerobius - 222 Oedobius - 234, 2 18 ogatai - 241 okinntvunus - 228 okinawensis - 238 olympica - 239 ornbrias - 229 omeishana - 232

oropaea - 225 orotypus - 223 outereloi - 237 ovaiis - 222 Oxybiella - 234

pacificus - 222 paddiae - 233 paganus (Linnaeus) - 229 paganus (Stephens) - 222 Paleaeomicromus - 234 palle.scens - 223 pallida, Drepanacra binocula var. - 218 pallida, Nusalala - 233 pallida, Zachobiella - 242 pallidulus - 223 pallidu.~, Hemerobius - 222 pullidus, Hemerobius

parvulus var. - 223 pallidus, Megalomus - 221 pallipes - 223 Palmobius - 234, 236 paradoxus - 229 parallelus, Micromus - 229 parallelus, Nenus - 229 Paramicromus (Nakahara) - 234, 226 Paramicrornus (Kruger) - 234, 226 purvulus, Hemerobius - 223 parvulus, Megalomus - 226 purvus, Megalomus - 226 parvus, Sympherohius - 237 paucinervis - 237 pehlkeanus - 223 pekinensis - 238 pellucidus (Walker) - 237 pellucidus (WalIengren) - 237 perelegans, Hemerobius - 223 perelegans, Micromus - 229 perifericu.5 - 224 perk~nsi - 230 perparvus - 238 persicu - 240 persimilis - 241 peterseni - 233 phaeostictus - 227 phalaenoides - 218 phaleratus (Hagen) - 223, 224 phaleratus (Hoffmanseg) - 223 philippinu,, - 230 Phlebiornu~ - 234, 226 Phlehonetna - 234, 218 P ~ C C M J - 223

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pic.tipe.s - 229 pictus, Megalomus - 226 pictus, Sympherobius - 238 pielina - 232 pigm. - 225 pinarensis - 233 pini (Leach) - 223 pini (Stephens) - 223 pinidumus - 223 pinincolus - 241 pinnatus (Navas, 1914) - 223 pinnatus (Navas, 1918) - 221 Pirionus - 234, 225 piunatulus - 22 1 placidus, Hemerobius - 223 piacidus, Micromus - 229 plaga - 218 plagatus - 229 Pleomegalomus - 234, 225 pleshanovi - 2 18 popii - 223 Porter - 234. 219 posticatus - 241 posticus - 229 preanubilus - 241 pretiosa, Psectra - 235 pretiosus, Wesmaelius - 241 princeps - 225 productus - 219 Psectra - 234 pseudofurcatus - 24 1 Pseudnmicromus - 235, 226 Pseudopsectra - 235, 226 Psychobiella - 235 psychodoides - 232 pulchellus, Carobius - 217 pulchellus, Micromus - 230 pumilio - 238 punctata, Megalomus (Okamoto) -

punctata, Notiobiella - 233 punctata, Zachobiella - 242 punctatus, Hemerobius (Goszy) - punctatus, Hemerobius (Stephens) punctatus, Megalomus (Kimmins) pupillus - 238 pusillus - 230 pygmaeus - 238 pyraloides - 226

quadrifasciatus - 24 1 quettanus - 223 quxamana - 235

radialis - 223 rafaeli - 226 raniosus - 230 ravus - 241 reconditus - 223 reisseri - 241 remiformis - 230 resinata - 21 8 reticulata - 233 Reuterobius - 235, 219 rhegmatica - 234 ricarti - 223 riudori - 238 rizali - 223 rosea - 233 rubrinervis, Drepanacra

binocula var. - 218 rubrinervis, Micromus - 230 rubrostigma - 233 rudebecki - 223 rufescens - 223

sabulosus - 230 sunctus - 237 sauteri (Esben-Petersen) - 230 sauteri (Nakahara) - 227 scalaris - 220 schedli - 223 schmidti, Palaeomicromus - 234 schmidti, Sympherobius - 237 Schneiderobius - 235, 219 schwarzi - 241 sriopterus - 220 scriptus - 238 semeriai - 233 semireticulatus - 223 Semohemerobius - 235 serrinus - 226 setosulus - 226 shensiensis - 236 shibakawae - 223 shikotanus - 223 siamica - 235 signatus Hemerobius (Hagen) - 223 signatus, Hemerobius (Kriiger) - 223 signatus, Sympherobius - 239 similis, Sineuronema - 236 similis, Sympherobius - 238 simplex - 224 simulans - 223 sinensis - 236 Sineuronema - 235

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World catalogue of Hemerobiidae

sinicus - 241 sinuata, Hemerobiella - 219 sinuatus, Megalomus - 226 sjostedti, Hemerobius - 223 sjostedti, Micromus - 230 skottsbergi - 224 smitheri - 238 sobrius - 230 solanensis - 224 sordida - 235 Spadobius - 236, 236 sparsus - 238 speciosa - 217 Spilomicromus - 236, 226 spirzigerus - 221 Spinomegalomus - 236, 225 spinosa - 233 stangei, Megalomus - 238 stangei, Sympherobius - 238 stellata - 233 Stenolomus - 236, 219 Stenomicromus - 236, 226 Stenomus - 236 stenoptevx - 228 stephensii - 224 stictica - 219 stigma - 224 stigtnaterus - 224, 223 stigmatica - 233 striata, Zachobiella - 242 striatellus - 237 striatus, Hemerobius - 224 strigosus - 224 subacutus - 235 subanticus - 230 Subboriomyia - 236 subcostalis - 238 subfalcatus - 224 subfasciatus, Carobius - 2 17 subfasciatus, Hemerobius - 221 submarginata - 242 subnebulosa (Kuwayama) - 24 1 subnebulosus (Stephens) - 241 subochraceus - 230 suboculata - 218 subolivacea - 233 su~cicus - 223 sufuensis - 241 sumatranus, Hemerobius - 224 surnatrunus, Micromus - 230 swczcyi - 230 Sympherobius - 236

Sympheromima - 238

tagalicus - 224 Tanca - 238, 226 tasmaniae - 230 tasmanica - 218 tateyamai - 224 terzdinosus - 227 tenellus - 237 tener - 226 tessellatus, Micromus - 230 tessellatus, Sympherobius - 238 texanus - 238 thaumasta - 238 theryanus - 230 tibialis - 224 tienmushana - 232 tillyardi - 225 timidus - 230 tineoides - 226 tjederi, Neuronema - 232 tjederi, Wesmaelius - 241 tolimensis - 224 topali - 224 tortricoides - 226 transsylvanicus - 241 transversus - 240 triangularis - 224 trifasciatus - 224 trifurcatus - 217 tristis - 236 tristriatus - 224 tr~venulatus - 241 tsinlinga - 231 tumida - 233 tunkunensis - 224 tuofenganus - 241 tuomurensis - 238 turneri - 233 tututrix - 221

ugandensis - 233 ulingensis - 242 ~ ~ l t i m u - 241 umbratus - 238 urrrbrosus - 230 uncata - 234 uniformis - 226 unipunctata - 232 unita - 233 uruguaya - 234 usingeri - 230

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V. J . Monserrat

ustulatus - 229 withycombei - 224 wuyianus - 238

vagans - 224 vagus - 230 vaillanti - 224 Vuju - 238, 232 valida - 233 variegatus - 230 variolosus - 230 vartianae - 242 venosus - 238 venustus, Hemerobius - 220 venustus, Sympherobius - 237 vicentei - 237 vicina - 233 villosus - 227 vinaceus - 230 viridinervis - 233 viridis - 233

weisong - 238 weryiae - 230 Wesmaelius - 238

xia - 230 xizangensis - 224

yadongana - 236 yemenicus - 242 yukonensis - 242 yunica - 236 yunnanica - 218 yunnanus, Megalomus - 226 yunnanus, Micromus - 230 yunpinus - 238 yunu - 235

Zachobiella - 242 zanganus - 224 zelenyi - 238 zernyi - 224 zhamana - 236 zhiltzovae - 242 zonarius - 230

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Mitteleuropas im Spiegel der Neuropteren-Fauna von Linz und OberOsterreich, sowie Bestimmungs- schussel fur die mitteleuropaischen Neuropteren und Beschreibung von Coniopteryx Lentiae nov. spec. Naturkundliches Jahrbuch Sradt Linz 1964: 127-282.

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World catalogue qf Hemerobiidae

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BANKS, N. 1909. Hemerobiidae from Queensland, Australia. Procreding.~ of the Entornologicul Society r?f Washington l l : 76-82.

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Arulgonrwi tlc, Cicv~cic~.r Nrttur-ules 12 : 77-9 1 . NAVAS, I,. 1013b. NCvroptCrcs. In: Mi.t.sion I'Eqrru~rur 10: 69-77. Gauthier-Villars. Paris.

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(I.ogro7io). Bolrtirr de /U Socieclrd Arugonesn dc, Ciarlc,irr.c Nuturclla,~ 13: 29-37. NAVAS, L. l 9 14b. Resultats scicntifiqucs. Inscctes NCvroptkres 1. Plenipennia et Mecoptera. In: Vquge dc

Ch. Allucrud er R. Jecrrlnel cJrl Afriqrrr Oric~nrtzle (1911-1912). 52 pp. Librairie Alhert S c h u l ~ , Paris. NAVAS, L. 1 9 1 4 ~ . Ncuroptcra asiatica. 111. RcI'N~, RNJSO ~I'Etrrornologic 14: 6-13. NAVAS. 1,. 1914d. New Neuroptera from the United States. BrllIc,tirr O ~ I I ~ P Brooklyn Ertronlologicc~l Society.

9: 13-18. NAVAS, L. 19 14e. Neurcipteros nuevos o poco conocidos. 11. Mf,rrroricl.s rle Itr Real Actr(1c~mia ( ir Ciencins y

Artes dc, B(rrce1onc1 1 1 : 105- 1 19. NAVAS, L. I9 14f. Neurcipteros sudarnericanos. I . Brotc,riu (Sen'e Zoologicrr) 12: 45-56. NAVAS. L. 1914g. RCsultats Scicntifiques de I'Expedition des Freres Kuznecov (Kouznetzov) a I'Oural Arcti-

que en 1909, sous la direction de H. Backluntl. Mc;moirc,.\ (Ie ltr Acuclc'rrrir Imprrirlle de.s Scirnt.e.v do Perrogrud 8 , vol. 28. 6: 1-2.

NAVAS, 1,. 191 5a. Contribute alla conosccnm degli insctti dcll'olivo dell Eritraen. Bt,lletit~o del Lnhorcrtorio di Zoologiu Gmercrle 6, Agrrrricr del11 R. Scuolr s~i j ) . ci'Agric~olturu Portici 9: 263; 332-333.

NAVAS, L. 191 5b. Ncurciptcros nuevos o poco conocidos. V . Me,moritrs dc, /U Rrcll Acc~~lc~mic~ clc. Cirncicrs v Artes de B(zrccj1oncz I l : 455-480.

NAVAS, L. 1 9 1 5 ~ . Neucipteros nuevos o poco conocidos. VI. Memorias rk. lu recil Ac.crderniu de Cirnc.ias Arts de Burcelorla 12: 1 19- 136.

NAVAS, L. 1915d. Neurcipteros nuevos dc E s p a h . I. Rc~vistcl </c 1'1 Recrl Accrderniu de Cic,nt.ic~s E.rurtcrs, Fi,si- r.n.s Y Nt~turcrles de Mndritl 14: 245-253.

NAVAS, L. 1915e. Excursioncs por Cataluiia. Julio 1914. Boletin de h Socird(ul Arrrgones(~ de Ciencim Nururu1e.c 14: 27-32; 35-59; 66-80.

NAVAS, I.. 1915f. Neue Neuropteren. Entorrrolo,yi.sche Mirteilurrxerr 4: 146153. NAVAS, L. 1916. Neurcipteros nuevos o poco conocidos. VII. Mrrnorirr.~ de 11 Real Ae.rrderrrir1 rlc Cierrc.ias

y Artrs de Bun-eloncr 12: 219-243. NAVAS, L. 1917a. Ncurcipteros nucvos o poco conocidos. VIII. M(,morius dc lo Rani Ac,udcrnicr de Cienc,ic~s

Y Ar1r.s tie Burcrloncr 13: 155- 178. NAVAS,.L. 1917b. InsecLa Nova. I, 11. M~~moric~dr~l l 'Ac~t~t~r l r~miu PontiJic.ir~ dr i N~rovi Lirrcai (2) 3: 1 1 1; 13-28. NAVAS, L. 1 9 1 7 ~ . Neurtiptcros nucvos o poco conocidos. I X . Mr~rrroric~.~ cle la Recrl A(~crclc~mia rlo Ciencicrs

y Arttjs cle Burcelonu 13: 393-406. NAVAS, L. 1917d. Ncuc Ncuroptcrcn. Entomologische Mi~tr i lur~,yen 6: 274-282. NAVAS, L. 191 7e. NCvropteres de i'lndo-Chine. 11. 1rr.set~tri. Rrrmes 8: 8- 17. NAVAS, L. 19 18a. Algunos insectos dc la Rcpublica Argentina. Revisru (16, lu Rc~rl Acrrtlc~trriu (If, Cic,nc.itr.s Er-

crc,ras, Fisicus y Nrrrurulf~s tlr M(rdri(1 16: 49 1-504. NAVAS, I*. 19 1%. Insccta nova 111. Mrtnoric' d(~I1 'AcccrtI(~rrriu Pont~/ic,itr clei Nnovi Lbrc,r,i (2) 4: 1- 1 1 . NAVAS, L. 1 9 1 8 ~ . Insectos chilenos. Boletin clc, 10 Soc.ic(ltrd Arugorrrstr tlr Cicvlcirr.~ Nuturcrlrs. 17: 212-230. NAVAS. L. 19 1%. Quelques Neurcipteres d'Algerie. In.src.tcl. Rc~rlnrs 8: 167-176. N AVAS, 1,. 19 1%. Ncuropteros de Espaiia nuevos. 11. Bol(,rirr (1. /(I Sot-ievful Errtorrroloxicrr tic E,spuEci 2: 2 1 8-223. NAVAS. L. 1920. Insectos rudamcricanos. 1, 11. 111. Anales de la Socic~dud Cierrr$crr Ar;qrtrrirrtr 90: 33-72. NAVAS, L. 192 Ia. NCvroptCres des Isles Juan Fe rnande~ et de L'lsle tle Piques. In: Skottsbcrg, C . (Ed.) The

N(rturu1 History of.luun Fi,rntrnrlr,z urrd Easter I.slarrd 3: 125-128. Uppsala. NAVAS, L. 1921b. Insectos americanos nucvos o criticos. Br-oreriu (Set-ic, ioologrru) 19: 1 1 3-124. NAVAS, L. 1922a. Algunos insectos del Museo de Paris. Rcvisrci dc /(I Acuclerrritr (If. Cic,nc,ius Excrc.trrs,

Fi.sico-Qnimicus y Ntrrnrcrles de Zrrcrgozcr 7: 15-5 1. NAVAS, I,. 1922b. Inscctos exoticos. Brotrricr (Sc.ricj Zoolo,yicc~) 20: 49-63. NAVAS, L. 1 9 2 2 ~ . Sur des NCvroptkres nouveaux ou critiques. V . Arrncrles (I(, 11 Soc.ic;tc; Stirnti/iquc7 dr

Bn~rel les 3: 250-256. NAVAS. L. 1922d. Estudis sobrc Ncurciptcrs. Ar-rius tlr I'lrrstitur de Ciolcicr.~. Barcrlorru 7 : 179-203. NAVAS, L. 1923. Insecta Nova. 1X. M(~mor i r rleII 'A(.cc~rl(,rr~iu Porlrificia clei Nuovi Lincei (2) 6 : 9-1 8 . , NAVAS, L. 1924a. Nueva especie de Neurbpteros de la fauna cspafiola. ~ol(,thr tic, lcr Soc,ir,dc~d Er~to~rrologiccr

de Espuiia 1924: 106-108. NAVAS, L. 1924b. Insecta orientalia. Ill. Metrrorie dell'A(~ccrtl~~rr~itr Pontificicl dci Nuovi Lrncri (2) 7: 2 17-228. NAVAS, L. 1924c. Algunos inscctos dc Chile. 11. Rf~vi.stu Cl~i lrnir del Hisroriu Nrrtnrcrl 28: 12-16. NAVAS, L. 1924d. Neuroptera er Mccoptera. In: %oological rc\ults o l the Swedish Expedition to Central-Africa,

1921. Arkivfur Zoologi 16 (13): 1-3. NAVAS, L. 1925a. Insectos exciticos nuevos o poco conocidos. 11. Mc,moric~s de In Rrrrl A(~c~c1c~rrrrtr rle, Cirtrcins

y Arrt,.c dc Burceloncr 19: 18 1-200. NAVAS, L. 192%. Algunos inscctos del Musco de Paris. 11. Brotrriu (Serie Zoologic~cr) 22: 75-83.

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World cutulogue of Hemerohiidue

NAVAS, L. 192%. Neuropteros del Museo d e Berlin. Revista rle la Aradernia de Ciencias E.ractus, Fisico- Quimicas y Nuturules de Zaragozu 9: 20-34.

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V.J. Mon~cjrrc~t

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Address of author: Dr Victor J. Monserrat , Departamento de Biologia Animal 1 Facultad de Biologia Universidad Complutense de Madrid 28040 Madrid Spain

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Bibliography of the Neuropterida Bibliography of the Neuropterida Reference number (r#): 4311 Reference Citation: Monserrat, V. J. 1990 [1990.??.??]. A systematic checklist of the Hemerobiidae of the world (Insecta: Neuroptera). Pp. 215-262 in Mansell, M. W.; Aspöck, H. (eds.). Advances in Neuropterology. Proceedings of the Third International Symposium on Neuropterology (3-4 February 1988, Berg en Dal, Kruger National Park, South Africa). South African Department of Agricultural Development, Pretoria. 298 pp. Copyrights: Any/all applicable copyrights reside with, and are reserved by, the publisher(s), the author(s) and/or other entities as allowed by law. No copyrights belong to the Bibliography of the Neuropterida. Notes: File: File produced for the Bibliography of the Neuropterida (BotN) component of the Lacewing Digital Library (LDL) Project, 2010.