2013 02 sugar producer 17-21 history of sugar beet-2
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8/17/2019 2013 02 Sugar Producer 17-21 History of Sugar Beet-2
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See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/236845913
History of sugar beet
ARTICLE · APRIL 2013
READS
60
2 AUTHORS:
Piergiorgio Stevanato
University of Padova
63 PUBLICATIONS 275 CITATIONS
SEE PROFILE
Leonard (Lee) William Panella
United States Department of Agriculture
157 PUBLICATIONS 638 CITATIONS
SEE PROFILE
Available from: Piergiorgio Stevanato
Retrieved on: 20 January 2016
https://www.researchgate.net/profile/Piergiorgio_Stevanato?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_7https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/profile/Piergiorgio_Stevanato?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_5https://www.researchgate.net/profile/Leonard_lee_Panella?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_7https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/profile/Leonard_lee_Panella?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_5https://www.researchgate.net/?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_1https://www.researchgate.net/profile/Leonard_lee_Panella?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_7https://www.researchgate.net/institution/United_States_Department_of_Agriculture?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/profile/Leonard_lee_Panella?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_5https://www.researchgate.net/profile/Leonard_lee_Panella?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_4https://www.researchgate.net/profile/Piergiorgio_Stevanato?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_7https://www.researchgate.net/institution/University_of_Padova?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_6https://www.researchgate.net/profile/Piergiorgio_Stevanato?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_5https://www.researchgate.net/profile/Piergiorgio_Stevanato?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_4https://www.researchgate.net/?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_1https://www.researchgate.net/publication/236845913_History_of_sugar_beet?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_3https://www.researchgate.net/publication/236845913_History_of_sugar_beet?enrichId=rgreq-49ebcd37-f147-4dea-bd0d-5239ade204fe&enrichSource=Y292ZXJQYWdlOzIzNjg0NTkxMztBUzoxMDQ2NDE5OTE4MDY5ODJAMTQwMTk2MDAwMzk2MA%3D%3D&el=1_x_2
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istory of
TH STORY OF CROPS is oft
en
under-
appreciated. This is not a good thing, espe-
cially for the farmers who gr
ow
them. ln fact,
knm.nng what happened after the domestica-
tion
an
d subsequent evolution of the modern
varieties could help us better understand
the
needs of
th
e species current ly being cultivated
on our
farms.
Sugarbeets
are
one of
the
newer crops.
They were created in Germany at t he end
of
the 1700s and rapidly became the most impor-
tant destination of he species Be ta vulgaris.
The sugarbeet's wild
pa
rent is the sea beet
Beta ma
ri
tima), sti
ll
living on the European
seashor
es
and whose leaves we
re
harvested
and ea ten by preh istoric man. After d
omestica
-
tion (about 8,500 B.C.), species first was
cultivated for the leaves, but unconscious
selection performed by the ancient farmers
led to garden,
or
red, beets (around 100 B.C.)
and the fodder beets. The latter appeared in
Europe around 1500 A.D., and the sugar-
beet was likely selected from it. Progress in
sugar yield, reduction of costs and manpower
requirements, processing quality, etc. has been
impressive in these p
ast 200
years since
the
creation of
the
sugarbeet. What is
sov.rn
to
da
y
is considerably different from
the
first sugar-
beet varieties.
The history of a friendlyand part ly infor-
mal collaboration between American and Ita -
Piergiorgio Stevanato
and
Leonard W. Panella
Beginning with Italian breeder Ottavio
Munerati on century ago scientists
continue work today to improve the species.
ian official research stat ions v .ri
II
be summa-
rized here. Notwithstanding, it is sti
ll
a rather
unknown activity that resulted in findings that
improved wor ldwide perf
orma
nce of sugar-
beers. One could say thar some rrairs ofthe
modern varieties, including important disease
resistances, originated and
were
distributed
thanks to the mentioned cooperation.
ercospora leaf spot
The histor y begins exactlyone century ago,
when the Italian breeder Ottavio Munerat
i,
worki ng fur
the
Royal Sugarbeet
Station
at
Rovigo, Italy
a
small town near
the
Po
River Delta), initiated crosses between
sugarbeets and Beta maritima.
According to other authors, Munerati
hypothesized that several traits of the ,.nld
parent disappea red gradually in the crop va-
riety due to the u nconscious selection applied
after domestication. It is well known that
the
cultivated species arc much more delicate and
require more protection and ca re than their
wild parents. Tn other words, the enhanceme nt
of rhe qualities requested by rhe farmers was
purchased with th e loss of
wil
d and potential
ly
useful rraits.
Munerati speculated that the recovery
of
such
tr
aiLs, including likely resistances against
some severe diseases, could be a feasible and
powerful means for increasing yield.
G
eo
rge Coons (left) and Lee
Li
ng , of the Food
and
Agr iculture
Or
ganization in
Ro
me, visit
on
the bank
of
t
he Po
River near the Adriatic
Sea (Porto Levante, Ita ly, 1951).
www.Su
garProducer.com
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• • • HISTORY OF
SUGARBEETS
Around 20 years later, by means of
intensive crosses between sugar beets and
seabeets collected in the Po Delta, Munerati
obtained
sugarbeets endowed with resistance
to cercospora leaf spo t (CLS). But
the
derived
hy
brids
still displayed a
num
b
er
of negative
characteristics from the wild
parent
s. The leaf
canopy
was very vigorous but multicrowned,
and the roots were observed to be fangy and
irregu lar in shape. The b est lines tended to
flower l
ater and
displayed high sugar
content
but poor roOLy ield. For these reasons,
the
hybrids agajn underwent severa l cycles of
selection in order to eliminate the undesired
characteristics.
George H. Coons was a plant
patho
logist
for the
Di
vision of
Sugar
Plant
In
vestigations,
USDA Bureau of Plant Industry, from 1925
to 1955. lie was involved in the diseases of
were released only
in a stricdy official
way and exclusively
to Italian,
German
and Japanese seed
companies. In the new
American environ-
ment, the variety was
repeatedly and
inten
-
sively tested
under
severe CLS attack.
According to Coons,
the Italian accessions
showed resis
tan
ce to
CLS
and
even under
diseased conditions
a llowed substanti al
improvement of s
ugar
production. lt is well
sugarbeet
and
also in d
evelopment
of breed- kno
wn that even
after
ing lines, parental lines and varieties with about 70 years, thls
resis
tance to
cercospora
leaf
spot, virus yellows is
the
most
important
and cur
ly top virus. I Ie
met with
Munerati
in
Italy in 1925 and 1935. After the second visit
at
Rovigo, he wrote, Munerati gave me seed of
his best variety I
esistant
to CLSI RO
58
1.
genetic r
es
istance
available against CLS.
hizomania
Enrico Biancardi displays a very developed eta maritima
Porto levante, Italy, 2011).
direL tly
know the disease, the
ca
use of which
was first discovered
in
Italy in 1
966
, 17 years
after his death.
At the time,
it
must be recalled; lines
and varieties selected by the Rovigo Station
Similar ly successful result s
occur
red in the
field of rhizomania resistance. Munerati
didn
t
However, in
CLS
-resistant materials, and
therefore de rived from his genotypes, the very
first type of resistance to rhizomania , called
8 Sugar Producer
FEBRLARY 20
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"l)'Pe Alba," was found. The name was given
because the seed company Alba bought t he
main
part ofMunerati's germplasm
in
1949.
The superior performance of multigerm
variety "Alba P"was observed initially in tr ials
grown in 1957, well before the discovery of
the agents causing the disease. More recently,
based upon observation
of
segregating popula-
tions, this kind of resistance was classified as
quantitative by Enrico Biancardi
at
Rovigo.
The more resistant variety, "Rizor," was
released in 1985 by
SES- Laly
after being di
s-
covered and developed by Marco De Bia
gg
i.
In
field trials grown in 1980, e Biaggi included
some
CLS
-resistant genotypes that woukl
have originated from germplasm selected by
Munerati. ln a trial located near Ravenna, an
unexpected and strong rhizomania infection
occurred. The yield
of
five
genotypes was
much better
than
other entries.
From these lines, the healthiest beets
were submitted
to
the
normal selection and
breeding procedures.
In
1984, a collaborative
study was set up with the Bologna University,
where the agents
that
caused rhizomania
were discovered. It was established
that th
e
fungus Polymyxa betae infects the beet rootlets
carrying
and
inoculating
the
virus, BNYVV.
The virus moves rapidly through the root
bundles of susceptible genotypes, whereas the
movement in
the
roots of " Uzor" appeared to
be hindered. Early in the selection program,
the " Uzor type" resistance was recognized as
monogenic and dominant.
In the summer of 1983, Alvin Erichsen, a
breeder working for Holly Hybrids, observed
very low sugar yield in a trial near Tracy, Calif.,
with the exception of
th
ree experi mental
hybrids which produced
five
times more than
1he suscepl ible check
USH
l l An ELISA Lest on
L
he
roots confirmed the presence ofBNYVV.
The hybrids had different pollinators, but the
same female parent, obviously carrying s
ome
unknown
re
sistance. During 1985 and 1986,
the mentioned hybrids were evaluated by
Robert Lewellen at Salinas, Calif., under severe
rhizomania conditions. Compared with resis-
tant
varieties of European and Japanese origin,
the
trials confirmed the high level
of
resistance
derived from
the
Holly seed
bearer
line.
Just as in the first trial, the three hybrids
segregated in a pattern
ty
pical for a single
dominant gene, subsequently called Rz. The
resistance gene reduces
the
BI\iYVV
replication rate in the root. It was observed
that the resistant (RzRz) genotypes had
Robert Lewellen discovered the Holly
resistance gene in
1985-1986.
significantly lower virus tite r than susceptible
(rzrz) genotypes. In other words, "Rizor" and
"Holly'' resistances both seemed to limit the
spread and replication of the virus inside the
root. Recently, due to
th
e similitude
of
the
resistance traits, it was speculated
that
Holly
resistance may have originated from old Italian
material as well.
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www. Suga rProd ucer.com
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8/17/2019 2013 02 Sugar Producer 17-21 History of Sugar Beet-2
5/6
• • • HISTORY
OF SUGARBEETS
Monogerm seed
Viacheslav Savitsky was a leading sugar-
beet breeder and geneticist in the former So-
viet Union. He and his wife, Helen, immigrated
to
the
United States
after
World
War
II.
He
len
was
an
excellent microscopist
an
d cytologist.
Before WWH, Savitsky had frequent and close
contacts with Munerati, who understood Ru s-
sian very
we
ll
and therefore was
updated
on
the research occurring in the USSR on select-
ing lines for monogerm seed. Surely around
1935,
Savitsky was provided with Munerati's
CLS-resistant lines.
Around
the
same time, Savitsky discovered
a quantitarive form of monogermity defined
as Russian, but whose transmission was dif-
ficult to control in commercial seed multiplica-
tion. /\.monogenic type
of
monogermity was
isolated as well. But
the
breeding work was
stopped
by the
war. In
1946,
Munerati wrote,
I hastily asked Savitsky for more details and
for a small quantity of the new type of seed
[carrying th e monogenic trait] so I would be
able to test
it
here. Obviously the request
could not be answered.
Thanks to Coons, in 1947, Savitsky was
employed in the United States by the Beet
Sugar Development Foundation and su bse·
Vi
ac hes lav and Helen Sav itsky
{Sa lt La
ke
City, 1959
quently by the
USDA
. His first task at the Salt
Lake City Station was t o find possible sources
of mono
germ seed for the domestic sugar-
beet growers and industry. At the time,
the
development of monogerm seed was becoming
essential due
t
the costs of manually singling
the beet crop. n this research, Savitsky collabo-
rated with several American
breeder
s, includ-
ing Eubanks Carsner, John McFarlane, Forrest
Owen, George Coons, Ray PendletOn and
others. The group was charged with finding
monogerm beets in different parts of the
U.S.
Savitsky knew that the variety Michigan
Hybrid
18''
was derived from the Polish CLS-
resiswnt varie ty Buszczynsky CLR,'' which
had been obtained from Italian genotypes. He
likely recalled
in
conversations wiLh Munerali
thaL
in such materials
tesLed in Lhe USSR
, a
few monogerm plants were found. Cyclesof
inbreeding had been used in Italy to help in
the identification
of CLS
resistance
and
this
method
possibly allowed the rare recessive
monogerm trait
to
be expressed.
In fact, in a four-acre seed production field
north of Salem, Ore., sown
with
Michigan
Hybrid
18,
Savitsky found five plants bearing
monogerm seeds. In
1953,
the selected off-
spring
of the
beet
SLC
101
was
made
available
to the American and European breeders, and
afte r a few years, the monogerm varieties be-
came a ll
that
was sown in developed countries.
Proof
of
Munerati's hidden and post
mortem involvement in this discovery might
be
given by the fact that Coons brought an
ST.C
101 seed sample to Rovigo
in
JuJy 195], two
years before its officiaJ release.
The
occasion
Maximize yield and sugar
with
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rh
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Sysstem-Ready v s an advanced formulation of
zinc
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manganese
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rap
id
absorption and complete plant movemen1 at nutnent
peak demand
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Zinc ts
the
cornerstone
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leaf
and
root development while
manganese Is key to
increased photosynthesis,
nitrogen assimilation, stress tolerance, andother essential
bio
-chemical reactions within the plant
Harves
t tonnage
depends
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weU
early-season nutritional needs are met. Peak nutrient demand
timing
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foliar
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il,
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I
ncreased beet tonnage and
sugar
are the
result.
20 Sugar Producer FEB RUARY 2013
on
t make sugar
beets
miss out on crucial Zn and
Mn.
Make
your beets
Syss
t
em-Ready
.
Cali
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8/17/2019 2013 02 Sugar Producer 17-21 History of Sugar Beet-2
6/6
was
th
e grand opening
of
the new research
fa-
cilities
at the Rovi
go Station, completely built
\vith
U.S.
funds. This could be mo re evidence
for the collective
but
unpubl
is
hed knowledge
and cooperation
be
tv.re
en scienti sts in
Tta
ly and
the
U.S.
at
this time.
Starting in the 1980
s the
old, fruitful
and tradilional collaboration was continued
between Lewellen
and
Bi
anc
ard i
working
at
the stalions of Salinas and Rovigo respectively
and
increa
se
d time
af
ter
Lime
. The
partn
er-
ship has
be
en
expanded to the s
ta
tions of
USDA-ARS at
on Collins, Colo.; East Lansing,
Mich.; and Fargo, N.D . [with Leonard Panella,
Mitch McGrath and Lar
ry
Campbell respective-
ly
I
wi
th
important results
and
publications, as
always, wi thout s
pe
cific finandal support.
n
example is the book on Beta maritima recently
published by Biancardi, Panella
and
Lewellen
reviewed by
Sugar Produ r
April2012}.
Piergiorgio Stevanato, currently
wo
rking
at
the Padua
University, is the
new
scientist in
this continuing
coll
aboration that has resulted
in new research pape rs being pub
li
shed on
germplasm resources, enhancement , genetics
and molecular characterization. And so, the
sto
ry
goes on. •
Acknowledgemen
ts
Th
is article was
wr
i
tte
n in collaboration with
Rober
t
Lewellen and
Enrico Biancard i who
ex
-
per
ienced more
or less
directly the
seco
nd half of lhe abcve stor
y.
Because
many
of the details
described are unpublished the ir he lp should be remembered with gra titu
de.
Pi
erg
iorgi
o Steva
nato
Leonard Panella on the shore
of
the
Adriatic Sea Aibarella , Italy 2 11 .
www. SugarProducer.
com
21