0bot.swu.ac.th/upload/topic_document/1245412248.pdf · 2009-06-19 · abstract the research papers...
TRANSCRIPT
!"#!$%! &'()"*+)+,-+. /0
%! 12%3!4567#%! 89: )#,;%)+<-:=7%<-:> ?=)+8$> ?@A1<A"
Study on Effect of Radiation on Flowering and Ornamental Plants
in Thailand
B/?4.:&'()"
1. ,C-!5; @D5E7#,%C5 D)&D$:!FB #%!
$!"G$- ,CAH'1)%='IF,J/ 4.: K&-&'()"
$!#,!&&,'$; <6&:G)$HC0 4.: K&-&'()"
#!$&'()"$;L<=:AC$,$)+,$C$%! &'()"(!%FB #%! ()=M)L#1.$"0&'()"N5?%! ()=%! B&!- .:A!#GO%31!,M 0 -D!&'A"!5)"1 ;$B '$A &'F P 7#B )%30
G.1. 2550
!"#$%&'() #$!
!"#$%&'()*+,+,- .! / 01230()45)617$8*9&'()*:;"230()4230 &3<0$%=31>?2@73A509 (-2.4) <B3&'=*3;)*70C8 0'8=0&'/0D ,1 0)2@9 =CEFB%=.8A8)-A8.8230=G3&'()* +,+,- .!,3(30*9 -. ;320 :;"H(%3B8%3=CE F8B8I&*1385I31 J ,3='HKI8 AG38)2138L0<3!$H>ME,A)85' N 3 &' K 3 0) 1 AC L 0 " *. 2 59 : ; " O , / P / = L ! " &' = * 3 7 3 A 5 09 : ; " AG 3 8) 2 B , A <. 4
<B3&'=*3;)*H2@5073A509 :;"B%,1A<.4138&'()* AG38)2138 !"200<230&'()*:BI1K35' (&K.)
=CEFB% &3<H,M6,HQRS,:;" &3<,8.H 03"B9+%,<$;AG3B0)-=G3230&'()*F8 0)618C6 :;"+,+,- .!H(%3+,1#;138&'()* B8)1AM, 5G303 :;"H,2A30=CEH2CE*&+%,1 =.2=I38=CEO4%A0%31,1 9 &3<0$%5I31 J =CE8G3<3FK%,%31,'1F803*138230&'()*H;I<8C6
!"#$%&'
!"#$$%&$%' !(#)%*+,-.#(/0#) !12($,(34.45'46728'0/2"8'09$:/,&;)9$:<.=8.-
$:>%7#(9? @.=. 2510-2550 .,A((#)%*+,-.45'4"#$!(@*'@B;)$C912(%#$3#$.#(%*D#"#$ $#-(#)"#$%*+,-9$:+E#9? 3*5(@*'@B.#(%*D#"#$67#( F $%'.,A( !(#)%*+,-.45!(@*'@B;)$C9%*.-#)*@)GBH!:9IJ>#@*<=K <@L52$%&$%'2(MBM%#'$C0<"45-%",&"#$;D0$,(34 H!: !12($,(34.45'46728'0/2"8'09$:/,& @&%7#;)9$:<.=8.-'4"#$=N"K#%*+,-<"45-%",&"#$;D0$,(34",&8'0/2"8'09$:/,&H!0% +E#)%) 27 %(=B 50 3"O!
'#""%7# 64 D)*/@LD P/-37%);>J7<9Q)8'0/2" +E#)%) 18 %(=B 37 3"O! '#""%7# 44 D)*/ <9Q)8'09$:/,& +E#)%) 6 %(=B 8 3"O! '#""%7# 10 D)*/ H!:<9Q)@$$R8'0)AE#9$:/,& +E#)%) 3 %(=B 5
3"O! '#""%7# 10 D)*/ >)7%-(#).45.E#"#$=N"K#'#".453O/ ML2 '>#%*.-#!,-<"K6$=#36$B M*/<9Q)$02-!: 79 $,(34.45'4"#$)E#'#;D0=N"K#'#".453O/ML2 $,(34H"''# P/-;D0%*G4"#$S#-H&&<S4-&@!,)
$7%'",&D*A)37%).45<!4A-(;)3T#@9!2/<DLA2 P/- !12($,(34.45'46728'0/2"8'09$:/,&),A)3#'#$U+,/<9Q)"!O7'8/0/,()4A ML2 !672"#$<9!45-)H9!(12(PM$P'PV' !672!,"KR:.#(3,RW#)H!:3$4$:12(@LD
!672"#$<9!45-)H9!(34/2"H!:;& !672"#$<+$*J<6*&P6H!:@,X)#"#$12(@LD !672"#$(2"H!:2,6$#2-C7$2/12(<'!Y/ !672$:&&"#$3L&@,)GOB !672%,Z+,"$D4%*6 H!: !672!,"KR:.#(3$4$%*.-#>!,("#$<"Y&<"45-% 9I++O&,)@&%7# 9$:<.=8.-9$:3&M%#'3E#<$Y+;)"#$9$,&9$O(@,)GOB8'0/2"8'09$:/,&P/-"#$;D0$,(34H!0% 3 @LD ML2 @O.G$,"K# (Canna generalis) H@$<V45-(8[0 (Portulaca
grandiflora) H!:<&J+'#= (Chrysanthemum morifolium) +E#)%) 22, 10 H!: 6 @,)GOB 6#'!E#/,&
H!:@$$R8'0)AE#9$:/,&24">!#-D)*/ 8/0H"7 2)C<&4-3 ‘)#)#’ (Anubias nana) 2)C<&4-3 ‘M2)<+)V*3’ (Anubias congensis) 2)C<&4-3 ‘&#$B<.2$45’ (Anubias barteri) 2)C<&4-3 ‘'*)*'#’ (Anubias
minima) P!&*<!4- (Lobelia cardinalis) H!:2P"!)4'# (Aglonema simplex) <9Q)60)
Abstract
The research papers that concerned the effect of radiation on flowering and
ornamental plants in Thailand during 1967-2007 in format of journal, annual reports, printed
academic matters, thesis and special problems were compiled. The purpose of this study
was to collect both usage and effect acknowledge of radiation. It was found that studied on
effects of radiation on flowering and ornamental plants in Thailand have been recorded for
27 families 50 genera and more than 64 species. Mostly studied on flowering plants were
18 families 37 genera more than 44 species, ornamental plants 6 families 8 genera more
than 10 species and aquatic plants 3 families 5 genera more than 10 species. The
Kasetsart University has reported seventy-nine percents of the research papers. Most
reports have shown that the major ray is gamma ray by acute radiation, which through in
vitro culture. Effects of radiation on flowering and ornamental plants could be provided into
8 groups; chromosome changes, morphological and physiological changes, flower and leaf
colors changes, plant growth and development, seed germination and survival rate,
reproductive systems, life cycle and postharvest physiology. In Thailand, ornamental plants
were Canna generalis, Portulaca grandiflora and Chrysanthemum morifolium, which have
been successful improved by radiation 22, 10 and 6 cultivars, respectively. In addition,
many aquatic plants have been improved by the same method such as Anubias nana,
Anubias congensis, Anubias barteri, Anubias minima, Lobelia cardinalis and Aglonema
simplex etc.
(1)
!"#$%
&'(!
!"#$% (1)
#$%&'(!"!) (2)
#$%&'*!+,"-./# (3)
#00'1 1 #023! 1
*45678$) 1
9$(:;,"- )<=>/).!"[email protected]!96B$C 2
>/#D>(>/).!"[email protected]!96B$C 2
,"-EC&2=0'1<!F9G!B-HFI"$# 2
#00'1 2 D/. !"J8-)!296B$C0'1D.'1C9>I/) 3
#00'1 3 96K'.!"F3!D262.!"96B$C 18
"4,J##.!"96B$C 18
:!20'1J8-"-C-D98!L2.!"F3!D262.!"96B$C 18
96K'.!"96B$CJ8-D.M#>I/548 18
96K'.!"F3!D262.!" 19
#00'1 4 N8.!"96D<"!-7=>I/548 20
H5IF/.H5I,"-F$# "950$O)+""PH5I2O3!,"-F$# 0'15'"!C)!2.!"L&I"$) 'D+Q1/.!","$#,";)+$2K;=L2,"-D0?H0C
20
.!"[email protected]!96B$CD.'1C9.$#N8>/)"$) '0'15'(G/H5IF/.H5I,"-F$#L2,"-D0?H0C L2&G9),R +.?. 2510-2550
25
H5IF/.H5I,"-F$#J8-+""PH5I2O3!,"-F$#+$2K;=L75G0'1HFIB!..!"&$.23!L7I.8!C+$2K;= EFCL&I"$) '
51
>I/F'->I/D 'C>/).!"L&I"$) '.$#H5IF/.H5I,"-F$# J8-+""PH5I2O3!,"-F$# 59
#00'1 5 ";, 96B!"P= J8->I/D 2/J2- 60
#00'1 6 #""P!2;."5 69
(2)
#$%)*+!"!,
+!"!, &'(! 1 "!C&Q1/H5IF/. (Flowering plants) 0'15'.!"[email protected]!:@)N8>/).!"S!C"$) 'L2
,"-D0?H0C
20
2 "!C&Q1/H5I,"-F$# (Ornamental plants) 0'15'.!"[email protected]!:@)N8>/).!"S!C"$) 'L2,"-D0?H0C
23
3 "!C&Q1/+""PH5I2O3!,"-F$# (Aquatic plants) 0'15'.!"[email protected]!:@)N8>/).!"S!C"$) 'L2,"-D0?H0C
24
4 N8>/)"$) '0'15'(G/H5IF/.H5I,"-F$#&26F(G!) T L2,"-D0?H0C B!."!C)!2.!"[email protected]!96B$C ($O)J(G,R +.?. 2510-2550
29
5 >I/F'->I/D 'C>/).!"L&I"$) '.$#H5IF/.H5I,"-F$# J8-+""PH5I2O3!,"-F$# 59
(3)
#$%)*-!./"012#
-!.3*4 &'(! 1 72G9C)!20'103!.!"[email protected]!96B$CD.'1C9.$#N8>/)"$) '0'15'(G/H5IF/.H5I,"-F$#L2
,"-D0?H0C L2&G9),R +.?. 2510-2550 25
2 &6O2 G92>/)H5IF/.H5I,"-F$#0'123!5!S!C"$) ' 26
3 "$) '0'1L&IL2.!"[email protected]!:@)N8>/)"$) '0'15'(G/H5IF/.H5I,"-F$#L2,"-D0?H0C 26
4 96K'.!"S!C"$) '0'1L&IL2.!"[email protected]!:@)N8>/)"$) '0'15'(G/H5IF/.H5I,"-F$#L2,"-D0?H0C 27
5 +;0K"$.A! !C+$2K;=(G!) T 0'1HFIB!..!"S!C"$) '&$.23!L7ID.6F.!".8!C+$2K;= .!"L&I,"-EC&2= J8-?42C=#"6.!"S!C"$) 'J.55! J8-96B$C269D<8'C"=D0<E2E8C' 57!960C!8$CD.A("?! ("= 52
6 J+"DU'1C)HVI+$2K;=(G!) T 0'1HFIB!..!"S!C"$) '&$.23!L7ID.6F.!".8!C+$2K;= N8)!2.!"96B$CJ8-+$W2!>/)57!960C!8$CD.A("?! ("= 55
7 !"#$%&'%()*+,-./0%1 2 345678$%#%99%:;:*!;:-1)*+,-.<7(9%:=>8:*1'4>*9+?%=@8 9A79%:9B%()*+,-. 56
8 )::C6$8+D?%#AEF345678#%99%:>*9+?%=@8 9A79%:9B%()*+,-.<7(9%:G%(:*1'4H9$$% 345+?%$%H'71=+1%+F*+;:I$1+8J$ 9B8% K:*D1345 18 F*+345 24-27 $AL-+%(+ 2549 C
&M+(.H'71'A+K8%HBI9%:;:I>-$JA$H)K $NJ13J1,%+4 58
!!"# 1
!$%&
'()*+,-. !"#$ %&'()*!""+( ,&-(./"0$/#+1+ 234+(5+6578-(./"9&:!";$6578-(./"98< =8:>?@0$/34+(A!B27 5@1$
6+ 24+:%0)*!""+(C-<6!DA!B270$/;:,4B+10+"34+(0E+C-<A!B27%6F86+ %&*$/:(G&*"98< C(H$AFB& @5E+A!(>;"% +B<;" %6$/ :A><;"6!D ! "#$B4 +"I 1+61+:;:4 +"-*$6 %*$/ :"914 98< %H4( G#"#A4 +" J*K/ (AF0:7 !"#$;!*B <+9A=;%*B J*K/(6 @G#9LLM+ !"#$%;6NO !"#$G611+ !"#$(FAB ;( %&'(B<( N./" !"#$%-*4+($P1$0!P"J7?G*@=0Q =8:%R)+@ !"#$0$/1$JA+1#+1+ 2C(6+ 0@*70@*A"98<#," %H4( !"#$%;6NO !"#$G611+ S*S
5@B<;"1$6+ &M;"6!(;!(B +:5+6 !"#$=8:CH<A!B270$/1$JA+1-(+G(4(#," %H4( J;(6 $B - K;G34(B@6!/A
-(+ I 8!"(!P(% +JA B<;" ,<5!6 !"#$H(F8B4+"I A10!P"AFT$&M;"6!(;!(B +:5+6 !"#$ N./"3,<0$/0E+"+(8<+( !"#$ 5@B<;"&UFD!BFB+1><;6E+-(8B4+" I %6$/:A6!D6+ CH< !"#$;:4+"%J 4"J !8 &V557D!(1$6+ (E+ !"#$1+CH<& @=:H(OC(8<+(B4+"I ;:4+"G) 4-*+: 0!P"C(0+"6+ G)0:O ;7B#+-6 1 G*@0+"6+ %6QB %&'(B<(
6+ CH<& @=:H(O5+6 !"#$C(8<+(6+ %6QB (!P( #4A(C-W4(E+1+CH<C(6+ & !D& 7")!(T7O)KH
%(K/;"1+5+6 !"#$#+1+ 2%-($/:A(E+C-<)KH%6F86+ 6*+:)!(T7O98< =8:)!(T7O)KH0$/& @#DJA+1#E+% X5C(6+ & !D& 7")!(T7O=8:6+ CH< !"#$ G*@%&'()KH%Y QZ6F50$/#E+J!W>;"& @%0Y %H4( ><+A)!(T7O 6> 6, 6> 10,
G*@ 6> 15 %&'(><+A0$/98< !D6+ %-($/:A(E+C-<%6F86+ 6*+:)!(T7O5+6><+A5<+A>+A8;61@*F 105 #4A(C(
91<8;691<& @8!D )DA4+1$6+ CH< !"#$C(6+ & !D& 7")!(T7O91<8;691<& @8!D-*+:H(F8 G*@& @#D3*#E+% X598<)!(T7OC-14;;61+-*+:)!(T7O %H4( )70T !6Q+ )!(T7O#<1#F (7H ([+A ? %-*K;"; 7?$ )F1)O !"#$ A!(AF#+ %&'(B<( (;65+6($P:!"1$91<8;691<& @8!DH(F8;K/(I ;$61+61+: %H4( G) %N$/:"9\< >F"G8" &0711+ %DW51+Y %&'(B<(
&V557D!(6+ CH< !"#$6!D91<8;691<& @8!D98< !DJA+1#(C51+6>.P( 0!P"C( @8!D(!6AF5!:9&5(2."%6QB 6 =8:1$%&M+-1+:%)K/;Y.6Q+2."3*>;" !"#$0$/1$B4;91<8;691<& @8!D G*@6+ & !D& 7")!(T7O%)K/;C-<98<)!(T7OC-14 I N./"(!6AF5!:G*@%6QB 6 #+1+ 2(E+)KH>;"B( 1+%><+ !D6+ R+: !"#$98<0$/-(4A:D F6+ R+: !"#$ 98<G64 #E+(!6"+(& 1+?,%)K/;#!(BF 6 @0 A"AF0:+Y+#B OG*@%0J=(=*:$ G*@Y,(:OD F6+ R+: !"#$G611+G*@AF5!:(FA%J*$: O%0J=(=*:$ #2+D!(AF5!:G*@)!](+ 1-+AF0:+*!:%6QB Y+#B O =8:B<;"%#$:J4+CH<54+:C(6+ !DD F6+
C(& @%0Y90: 6+ Y.6Q+2."3*>;"6+ R+: !"#$6!D91<8;691<& @8!D(!P( 1$ +:"+(;:,41+6);#1JA C(91<8;691<& @8!D-*+:H(F8 GB4;:4+"9 6XB+1><;1,*%-*4+($P:!";:,4C(A"0$/5E+6!8 N./"#4A(C-W45@2,6%3:G) 4=8:(!6AFH+6+ G*@(!6AF5!:0$/0E+"+(%6$/:A><;"6!D !"#$%04+(!P( 0E+C-<;"JOJA+1 ,<0+"8<+(($P:!"914G) 4-*+:%04+0$/JA 8!"(!P(%)K/;C-<JA+1 ,<0+"8<+(AFH+6+ %6$/:A6!D3*>;"6+ R+: !"#$6!D91<8;691<
2
& @8!D>;"& @%0Y90:G) 4-*+:1+6:F/">.P( 5."1$JA+15E+%&'(B<;"0E+6+ AD A1G*@#!"%J +@-O;;61+%&'(;"JOJA+1 ,<=8:% XA %)K/;%&'(><;1,*)KP(Z+(%6$/:A6!D6+ CH< !"#$6!D8;691<& @8!D A12."6+ 5!80E+%&'(Z+(><;1,*%3:G) 4C-<G64H71H(B4;9&
/-0123456.789:.;&4<=;>&/*?-@
%)K/; AD A1;"JOJA+1 ,<%6$/:A6!D6+ CH< !"#$ G*@3*>;" !"#$0$/1$B4;91<8;691<& @8!DC(& @%0Y90: #E+- !DCH<%&'(G(A0+"C(6+ Y.6Q+ AF5!: G*@)!](+6+ CH<& @=:H(O5+6 !"#$6!D91<8;691<& @8!DB4;9&C(;(+JB G*@(E+;"JOJA+1 ,<0$/98<%3:G) 4G64H71H(B4;9&
9: A909:.;&4<=;>&/*?-@
0E+6+ B A5%;6#+ 0$/%6$/:A6!D6+ CH< !"#$6!D91<8;691<& @8!D %R)+@0$/1$6+ Y.6Q+AF5!:G*@1$ +:"+(0+"AFH+6+ [+:C(& @%0Y90: =8:1$6+ *")F1)OC( ,&>;"A+ #+ 0+"AFH+6+ +:"+(6+ AF5!:& @5E+& AF0:+(F)(TO &VW-+)F%YQ #F/")F1)OB4+" I %04+(!P(
345B@C$8!"#7&D/E&?5FDG4-
1. 98<;"JOJA+1 ,<%6$/:A6!D6+ CH< !"#$ G*@3*>;" !"#$0$/1$B4;91<8;691<& @8!DC(& @%0Y90:
%)K/;5!80E+%&'(Z+(><;1,*6+ CH< !"#$6!D91<8;691<& @8!D>;"& @%0Y90: =8: AD A19A< ? -<;"#178
Y,(:OAF5!:G*@6+ 5!86+ JA+1 ,<0+")_6QY+#B O 1-+AF0:+*!:Y $(J F(0 AF= ] ;"J !6QO 2 . D7J*+6 (F#FB (!6Y.6Q+ G*@(!6% $:( >;"1-+AF0:+*!:Y $(J F(0 AF = ]G*@
#2+D!(6+ Y.6Q+;K/( I A12."%6QB 6 =8:%R)+@%6QB 6 3,<3*FB91<8;691<& @8!DC(5!"-A!8(J (+:6 1$G-*4"J<(JA<+><;1,*%6$/:A6!D6+ CH< !"#$6!D91<8;691<& @8!D
3. CH<%&'(><;1,*)KP(Z+(C(6+ CH< !"#$6!D91<8;691<& @8!DH(F8;K/(I B4;9&
4. #+1+ 2(E+3*0$/98<1+CH<%&'(G(A0+"C(6+ CH< !"#$6!D91<8;691<& @8!DH(F8;K/(I
3
!! "# 2
$%&'()*+,-(./0123!"#$&"#3/45%-
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
!"#$ '*336*.+19!"1)0*E4*2%FL#2O!48,*"1*;M*(B957>J+ 2 1)F02 %&'
- !"#$,$-B2010'B'''+ (non-ionizing radiation) B95.10 !"#$%)&-+.207@)A1BCCD* 7N0+
%E*2 5'+ .#" 7#$;" %)&-+E8,;F '!4 *BE:'7)4 .)IB2:% 7EC
- !"#$,$-10'<@57189B'''+ (ionizing radiation) B95.10 !"#$7'1GH !"#$.122* !"#$.')C* !"#$7O45*.)I !"#$+8E4 '+ !"#$<+1)F02+$S2$/=57 $;1'$1N&-'@+P-"E0* “ !"#$> 2*L= (atomic radiation)”
(' FL$, 2546; Casarett, 1968)
)2-'".0/$6+"3)7 !"#$+8E7%)$; H %&' !"#$,$-2$.@)0"16*7+893*1+8E7%)$;#?'"'I4'2 7189?PS+7+&-'"3*12$+8E7%)$;#,$-B207#Q$; (Unstable) 3P"45'">)0';()!""*+#0E+718+''12*<+ =>?'" !"#$+8E7%)$; H :9; !"#$+8E7%)$; H+$S +!1> !O> F"(!+TFH+8;2+6*2*<N5UP1K*1* 1)*;(!+TFH @ &'1* 7>)$-;+.>)"?'";$+ (gene)
(' FL$, 25301) :9; !"#$+8E7%)$; H 2$ 4 N+89 %&'
!"#$.')C* @2*;QP" 1 I.#?'"'+FM*%.')C*,$-./0''12*3*1+8E7%)$;#<9 R 2$'6*+*310'<@571891* .414!EB959$ .402$%E*2#*2* Q<+1* ,I)F,I)E"/0*+E!4QF+5';2*1 '+FM*%.')C* 1 '+FM*% %&' +8E7%)$;#?'"T*4FV$7)$;2 GP-"> I1'O95E;:> 4'+ 2 '+FM*% .)I+8E4 '+ 2 '+FM*%
.)I2$> I3FBCCD* +2 @+0E;
!"#$7O45* @2*;QP" 1 I.#?'"'+FM*%'87)A14 '+,$-./0''12*3*1+8E7%)$;#<9R 2$'6*+*310'<@571891* .414!E+5';1E0* !"#$.')C* .40#*2* Q,I)F,I)E"B959$1E0* 4*2>148<++8E7%)$;#B202$'87)A14 '+ 3I71897>J+'+FM*%:> 4 '+ .)I'87)A14 '+ GP-"'87)A14 '++$S7'",$-7 $;1E0* '+FM*%O$4*
4
!"#$.122* 7>J+%)&-+.207@)A1,$-2$%E*2;*E%)&-+#!S+2*1 2$3F916*7+893*1+8E7%)$;# 2$'6*+*3,6*<@571891* .414!E+5';2*1 .402$%E*2#*2* Q,I)F,I)E"#="
!"#$+8E4 '+ 7189?PS+<+7% &-'"7 0"'+FM*%+8E4 '+ @ &'<+7% &-'">W81 LH> 2*L= :9;<+7% &-'"9!"1)0*E3I/)84'+FM*%+8E4 '+B95';0*"2*12*; .)I+8E4 '+,$-71897@)0*+!S+2$> 82*L2*1
.)I7%)&-'+,$-95E;%E*27 AE#="2*195E;
+'13*1 !"#$+8E7%)$; H ,!S" 4 N+89.)5E ;!"2$ !"#$'$1 2 N+89 ,$-28<N0 !"#$+8E7%)$; H .401A7>J+,$-+8;2+6*2*<N57>J+#8-"10'1* 1)*;(!+TFH %&' !"#$7'1GH .)I !"#$'!)4 *BE:'7)A4 (' FL$, 2530?)
!"#$7'1GH 7>J+%)&-+.207@)A1BCCD*,$-2$%E*2;*E#!S+2*17N0+1!+ 2$%FL#2O!487N0+79$;E1!O !"#$.122* .40B20B952*3*1+8E7%)$;# .402$3F916*7+893*1N!S+?'"'87)A14 '+?'"'I4'2<9 R 7N0+ 71893*11* ;8"'87)A14 '+,$-2$%E*27 AE#="B>Q=17>D*,$-,6*95E;:)@I9!"7N0+,$-7189<+7% &-'"7'1GH7 ;H 7>J+45+
!"#$'!)4 *BE:'7)4 71893*19E"'*,84;H @ &',6* 7,$;2B953*1@)'9BC
low-pressure 8-W Hg aro lamp :9; !"#$+$S2$'6*+*31* ,I)F,I)E"4-6* 3P"2!1X*;<@5.10#>' H@ &')I''"7 L= 7>J+#0E+<@Y0 <+"*+E83!;,*"N$EE8,;* (OE0*.#"'!)4 *BE:'7)4 ,$-%E*2;*E%)&-+ ';=0 I@E0*" 2500-2900 +*:+724 3I2$> I#8,T8M*(#="#F9 (' FL$, 2530?)
8.9/3/2:)2-'" @+0E;E!9 !"#$,$-+8;2<N5<+>Z33FO!+ :9;7X(*I !"#$,$-+6*2*<N5<+1* > !O> F"(!+TFH 3I2$@+0E;7>J+ . 9 .)I 71 ;H GP-",!S". 9 .)I 71 ;H 7>J+@+0E;#6*@ !OE!91* 9=91)&+()!""*+3*1 !"#$:9;E!4QF4!E1)*" 72&-' !"#$3*145+16*7+89 !"#$/0*+4!E1)*" !"#$O*"#0E+3IQ=1E!4QF4!E1)*"9=91)&+BE5 GP-"3I2$> 82*L2*1@ &'+5'; ?PS+1!ON+89?'"E!4QF.)IN+89?'" !"#$ #0E+,$-7@)&'3I,I)F,I)E"/0*+4!E1)*"''1B> (' FL$, 2546)
1 . 9 (radiation absorbed dose; rad) @2*;QP" > 82*L !"#$,$-E!4QF4!E1)*",$-B95 !O !"#$7'1GH@ &' !"#$.122* 9=91)&+()!""*+3*1 !"#$BE5 100 7'8 H1 40'2E) 1 1 !2 ?'"E!4QF+!S+
>Z33FO!++8;2<N5@+0E; 71 ;H (gray; Gy) O'1> 82*L !"#$,$-E!4QF4!E1)*",$-B95 !O
:9; 100 . 9 = 1 71 ;H Q5*> 82*L#="?PS+1A<N5@+0E;,$-<@Y0?PS+ %&'7>J+18:). 9 @ &'18:)71 ;H
:9; 1000 . 9 = 1 18:). 9
1000 71 ;H = 1 18:)71 ;H
5
;),<3=.74%-)2-'" (#6*+!1"*+> 2*L=7(&-'#!+48, 2549%)
1. > I:;N+H95*+1* E83!; B95.10 7>J+U=+;H1)*"[\1'O 2.)IQ0*;,'97,%:+:);$ E87% *I@H E83!; .)I(!]+*,*"95*+E8,;*U*#4 H.)I7,%:+:);$ 7>J+45+
2. > I:;N+H95*+1* .(,;H.)I#*T* L#F? B95.10 <N54 E3 E8+83X!; .)I !1K*: %40*" R
7N0+ %'('1 7+&S'"'1 2I7 A" 7>J+45+
3. > I:;N+H95*+'F4#*@1 2 7(&-'<N5<+1* 4 E3#'O@* 0'" ';O1( 0'" 7N0+ ';.41 5*E E!9%E*2@+*.+0+ @ &'%E*2O*"?'"E!4QF <N5<+1* ^0*7N&S': %<+'F4#*@1 27% &-'"2&',*"1* .(,;H 7>J+45+
4. > I:;N+H95*+1* 71K4 <N5<+1* Q+'2'*@* 1* ;&9'*;F1* 71AO71$-;E/!1 /)B25 .)I7+&S'#!4EH40*" R <N5<+1* > !O> F"(!+TFH(&N <N5<+1* >D'"1!+16*3!9U!4 =(&N 7>J+45+
5. > I:;N+H95*+'&-+ R 7N0+ 95*+:O *L%9$ 2$1* <N5 !"#$7(&-'7> $;O7,$;OE87% *I@HN+89.)I> 82*L?'"T*4F'"%H> I1'O?'"E!#9F,$-+6*2*# 5*"> *#*,:O *L ,6*<@5#*2* Q IOF.@)0",$-2* 7(&-'> I:;N+H<+1* #&O%5+,*":O *L%9$40'B>
&()&+(3>2.?@74%->A=
1* 1)*;(!+TFH %&' 1* 7>)$-;+.>)"#* (!+TF1 2?'"7G))H 7189?PS+1!O:27)1F)?'" DNA
.)I#*2* QQ0*;,'9B>;!" F0+40'B>B95 :9;/0*+1* .O0"7G))H (7U K_E!N , 2536; ($ +FN, 2546) GP-"B20<N5> *1W1* LH4*2>148?'"1 IOE+1* .;13*11!+ (segregation) @ &'1* E24!E1!+<@20 (recombination) ?'";$+ (Broertjes, C. and A.M. van Harten, 1978) 1* 1)*;,6*<@5#8-"2$N$E842$)!1KLIO*"';0*".4140*"B>3*1O (OF FKGP-")!1KLI,$-7>)$-;+B>2$,!S",$-9$@ &'B209$ (#6*+!1"*+> 2*L=7(&-'#!+48, 2549?)
:9;>148(&N3I2$1* 7>)$-;+.>)"@ &'1)*;(!+TFH';=0O5*"4*2T 2N*48 .40'!4 *1* 71891* 1)*;(!+TFH%0'+?5*"4-6* 7 $;1E0* “1* 1)*;(!+TFH4*2T 2N*48” .40';0*"B 1A4*27 *#*2* Q7(8-2'!4 *1* 1)*;(!+TFH<@5#="?PS+ :9;1* 7@+$-;E+6*95E;#8-",$-10'<@571891* 1)*;(!+TFH 7N0+ !"#$@ &'#* 7%2$ 7 $;1E0* “1* 7@+$-;E+6*<@571891* 1)*;(!+TFH” )!1KLI1* 7>)$-;+.>)",$-71891* 7@+$-;E+6*<@571891* 1)*;(!+TFH3IB20.4140*"3*11* 1)*;(!+TFH4*2T 2N*48 .401* 7@+$-;E+6*<@571891* 1)*;(!+TFH3I7>J+1* 7(8-2:'1*#@ &'%E*2Q$-<@571891* 1)*;(!+TFH GP-"7>J+1* # 5*"%E*2@)*1@)*;?'"1* 1)*;(!+TFH %&' 2$:'1*#7189)!1KLI1)*;@)*; =>.OO?PS+ (' FL$, 2546)
!"#$,6*<@571891* 1)*; (mutation) GP-"7>J+'8,T8()?'" !"#$,!S",*"4 ".)I,*"'5'2,6*<@571891* 1)*;B95 2 I9!O %&' 1* 1)*; I9!O;$+ (gene mutation) .)I1* 1)*; I9!O:% :2:G2
(chromosome mutation)
- 1* 1)*; I9!O;$+ (gene mutation) 7>J+1* 1)*;,$-7189?PS+1!O;$+ GP-"'*37($;"@+P-" +8E%)$:'B,9H .)5E,6*<@5)!1KLIC`:+B,>a7>)$-;+.>)"B> 7N0+ 1* 7(8-2@ &'1* ?*9@*;B>?'"7O#
6
- 1* 1)*; I9!O:% :2:G2 (chromosome mutation) 7>J+1* 1)*;,$-7189?PS+1!O => 0*" 36*+E+ .)I?+*9?'":% :2:G2
1* 1)*;,!S"#'" I9!O 7189B95<+7G))H#&O(!+TFH (sex cell) @ &' 7G))H 0*"1*; (somatic cell)
GP-"@*17>J+1* 7>)$-;+.>)",$-7189?PS+<+7G))H#&O(!+TFH 1A#*2* QQ0*;,'9B>;!")=1@)*+B95 ,6*<@5B95)=1,$-2$)!1KLI.4140*"B>3*1(0'.20 .40Q5*7>J+1* 7>)$-;+.>)",$-7189?PS+<+7G))H 0*"1*; 3IB20#*2* QQ0*;,'9B>;!")=1@)*+B95 45'"<N5E8T$1* ?;*;(!+TFH.OOB20'*U!;7(U (vegetative
propagation) 7,0*+!S+ ,$-3IQ0*;,'9)!1KLI1* 1)*;B>;!")=1@)*+B95 (' FL$, 25301)
)2-'"&2 &();)2 ;)@->2.?@7>A=
1* > !O> F"(!+TFH(&N<@5B95(!+TFH<@20,$-.4140*"B>3*1(!+TFH7982 GP-")!1KLI,$-7189?PS+<@20+$S'*37>J+)!1KLI(!+TFH,$-9$.)I.>)1<@20B>3*17982 7N0+ #$9'1 )!1KLI =>, "?'"9'1 #$<O
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ottschalk and Wolff (1983) B95 *;"*+BE5E0* (!+TFH(&N,$-> I#O%E*2#6*7 A33*11* <N54!EN!1+6*<@571891* 1)*;(!+TFH ,!S"@29 485
(!+TFH +!S+ 94 7>' H7GA+4H 7>J+/)2*3*11* <N5 !"#$ :9;7>J+1* <N5 !"#$.122* 40.6 7>' H7GA+4H 1* <N5 !"#$N0E;<+1* > !O> F"(!+TFH(&NB9596*7+8+1* 2*7>J+7E)*1E0* 70 >` .)5E B95(!+TFH<@20
> I2*L 2,200 1E0*(!+TFH ,$-+6*2*>)=17>J+1* %5* :9;7>J+T!Y(&N > I2*L 1,000 (!+TFH B259'1
B25> I9!O > I2*L 550 (!+TFH #6*@ !O> I7,UB,;7 8-2+6*7,%+8%+$S2*<N5 72&-' >` (.U. 2508
7>J+45+2* :9;1 2E8N*1* 71K4 B95+6*72)A9?5*E2*X*; !"#$,$-#6*+!1"*+()!""*+> 2*L=7(&-'#!+48 (N&-'7982) 3+1 I,!-"%!97)&'1B95?5*E(!+TFH<@20 3 (!+TFH %&' ?5*E(!+TFH 1? 6, 1? 10 .)I 1? 15
(#6*+!1"*+> 2*L=7(&-'#!+48, 25491)
)2-'"!"#.03BC=5C.&();)2 ;)@->2.?@7>A=
!"#$,$-+8;2<N5<+1* > !O> F"@ &'# 5*"(!+TFH(&N<@20 B95.10 !"#$.122*.)I !"#$7'1GH 7+&-'"3*1@*<N5B95%0'+?5*""0*;.)I2$%FL#2O!48<+1* ,I)F,I)E"/0*+E!4QFB95#=" ,6*<@5#*2* Q/0*+7?5*B><+(&NB959$ (#8 +FN, 2540; M!, 2*U, 2548) .)I72&-'X*; !"#$<@51!ON8S+#0E+?'"(&N 3IB20(O !"#$,$-7>J+'!+4 *;41%5*"';=0<+(&N ,6*<@5>)'9M!;40'/=5,$-+6*(&NB>>)=1.)I9=.) !1K* ';0*"B 1A4*2
7
!"#$.122*7>J+ !"#$,$-B95 !O%E*2+8;2<+1* +6*2*<N5> !O> F"(!+TFH(&N2*11E0* !"#$7'1GH 7+&-'"3*1 !"#$.122*2$%E*2;*E%)&-+4-6*1E0*3P"2$'6*+*3<+1* ,I)F,I)E"/0*+E!4QFB95#="1E0* (Casarett, 1968)
'9/.4%->A=!"#'(B()D.E(B(C=5C.&()F(3)2-'" #0E+?'"(&N,F1#0E+,$-?;*;(!+TFHB95#*2* Q+6*2*X*; !"#$7(&-'> !O> F"(!+TFHB95 ,!S"+$S?PS+1!O(&N.40)IN+89 GP-"#*2* Q.O0"B959!"+$S
1. (&N,$-?;*;(!+TFH:9;'*U!;7(U
7>J+1* ?;*;(!+TFH:9;1* <N572)A9 1* ,6*<@57189(!+TFH<@20R #*2* Q,6*B95:9;/#271# 4!E/=5.)I71# 4!E72$;?'"(&N40*"N+89@ &'40*"#*;(!+TFH1!+ .)5E%!97)&'1)=1/#2,$-2$)!1KLI4*2,$-45'"1* E8T$1* +$S3I,6*<@5B95(!+TFH,$-2$)!1KLI<@20R .4045'"<N57E)* .40Q5*@*1+6*72)A92*X*; !"#$1A3I7>J+'$1E8T$@+P-",$-N0E;7(8-2:'1*#<+1* 1)*;(!+TFHB95#="?PS+
2. (&N,$-?;*;(!+TFH:9;B20'*U!;7(U
7>J+1* ?;*;(!+TFH:9;1* <N5#0E+'&-+ R ,$-+'17@+&'B>3*172)A9 7N0+ ?;*;(!+TFH:9;1* <N518-" @+0' 7@"5* B@) @!E *1 <O E2QP"45+,$-B953*11* 7(*I7)$S;"7+&S'7;&-' GP-"(&N,$-2$1* ?;*;(!+TFH.OO+$S3IB20#*2* Q> !O> F"(!+TFH(&N95E;E8T$1* <9R +'13*17@+$-;E+6*<@571891* 1)*;(!+TFH:9;1* <N5 !"#$@ &'#* 7%2$
3. (&N,$-?;*;(!+TFHB95,!S" 2 E8T$ 7>J+1* ?;*;(!+TFH:9;1* <N572)A9.)I#0E+'&-+R .404*2T 2N*48(&N,$-#*2* Q
?;*;(!+TFH95E;E8T$1* B20<N57(U #0E+<@Y03I2$7>' H7GA+4H1* 48972)A94-6* ,6*<@52$36*+E+72)A9B202*1(',$-3I?;*;(!+TFH @ &'2$7>' H7GA+4H1* "'1?'"72)A94-6* 9!"+!S+(&N1)F02+$S3P"#*2* Q<N572)A9@ &'#0E+'&-+B95
/0?"&()F(3)2-'" 1* X*; !"#$<@51!O(&N@ &'N8S+#0E+?'"(&N #*2* Q,6*B95 2 E8T$ (' FL$, 2539) %&'
1. 1* X*; !"#$.OO7X$;O()!+ (acute radiation)
7>J+E8T$1* X*; !"#$1!O4!E';0*"(&N<+> 82*L !"#$,$-#="<@57# A3#8S+<+ I;I7E)*'!+#!S+
7(&-'28<@5(&N2$:'1*#G0'2.G2%E*27#$;@*;<+N0E",$-B95 !O !"#$ 1* X*; !"#$.OO+$S3IX*;7X(*I#0E+@+P-"#0E+<9?'"(&N1AB95 ,6*<@5'!4 *1* 1)*;(!+TFH:9;,!-EB>%0'+?5*"#=" ?+*9?'"#0E+,$-1)*;<@Y0 ,6*<@5"0*;40'1* %!97'*)!1KLI+!S+ R ''12*
2. 1* X*; !"#$.OO7 &S' !" (chronic radiation)
7>J+E8T$1* X*; !"#$1!O4!E';0*"(&N<+> 82*L !"#$+5';R 40'@+0E;7E)* .40<@548940'1!+7>J+ I;I7E)*;*E+*+ 7(&-'<@5,F1 I;I?'"1* 73 8Y748O:4 @ &'.O0"7G))HB95 !O !"#$ GP-"1* X*; !"#$.OO+$S3I7189%E*27#$;@*;.10(&N+5';1E0*1* X*; !"#$.OO7X$;O()!+<+1 L$,$-B95 !O
8
!"#$> 82*L7,0*1!+ .401A#0"/)<@5#0E+,$-1)*;(!+TFH2$?+*97)A1 3P"7>J+1* ;*1.101* %5+@*@ &'
%!9.;1#0E+,$-1)*;''12*
G+!"#$&0:4HI.&2 $J++7K(38+2-!"#>A=L:5)2 )2-'" %E*2/89>148,$-7189?PS+1!O7G))H @)!"3*1,$-B95 !O !"#$ %&' !"#$3I#0"/)1 I,O40':% "# 5*".)I@+5*,$-?'"7G))H E2QP"#0"/)1 I,O40'1* .O0"7G))H95E; 1* 7>)$-;+.>)"3I7189?PS+2*1@ &'+5';?PS+';=01!O> 82*L !"#$,$-B95 !O '!4 * !"#$ N+89?'"7G))H 7>J+45+ (' FL$, 2540) /)?'" !"#$1!O7G))H.O0"B957>J+ 2 N+89 %&' /)40'1* .O0"4!E?'"7G))H /)40'1 IOE+1* 72.,O')8GP2?'"7G))H (#8 +FN, 2527)
- /)40'1* .O0"7G))H 1* ,$-7G))HB95 !O !"#$> 82*L+5'; R 3I,6*<@57189%E*2)0*N5*<+1* .O0"7G))H :9; !"#$2$
/),6*<@52$1* )936*+E+7G))H,$-16*)!".O0"4!E)" 7 $;1E0* B2:4481 9$7);H (mitotic delay) GP-"3I7>J+';=0N0E"@+P-"M*;@)!"7G))HB95 !O !"#$ @)!"3*1+!S+7G))H3I7 8-22$1* .O0"4!E<@20 7(&-'7(8-236*+E+7G))H :9;1* .O0"4!E.OOB2:4G$#?PS+ 7 $;1E0* B2:448% :'7E' HN=4 (mitotic overshoot) %E*2)0*N5*<+1* .O0"4!E?'"7G))H 2$#*7@4F@)*;> I1* 7N0+ #* 7%2$,$- 71$-;E?5'"<+1* .O0"7G))HQ=11 I,O1 I7,&'+:9; !"#$ 1 IOE+1* # 5*":> 4$+@;F9NI"!1 B202$1* # 5*":> 4$+?PS+2*<N5<+1* .O0"7G))H 1 IOE+1* 7?5*2* E21)F021!+?'":% :2:G2 (chromosome assembly) )0*N5*)" @ &'7(8-2%E*27@+$;E (stickiness) <@51!O:% :2:G2 #0"/)<@5:% :2:G22$%E*2;*1)6*O*1<+1* .;1''13*11!+7(&-'B>#=07G))H,$-7189<@20 (daughter cells) ';0*"B 1A4*2%E*2)0*N5*?'"7G))H<+1* 7?5*#=0B2:4G8# ?PS+';=01!O> 82*L !"#$,$-7G))HB95 !O .)I I;I?'"7G))H<+?LIB95 !O !"#$O*" I;I (stage of cell cycle) 3IBE40' !"#$2*11E0* I;I'&-+ R ';0*"<9';0*"@+P-" - /)40'1 IOE+1* 72.,O')8GP2?'"7G))H
1* 7>)$-;+.>)"?'"72.,O')8GP2,$-7189?PS+M*;<+7G))H(&N,$-B95 !O !"#$ 7>J+/)2*3*1 !"#$ B>,6*)*;7;&-'@F5240*" R M*;<+7G))H2*11E0*,6*<@57'+BG2H40*" R ,$-71$-;E?5'"1!O1 IOE+1* 72.,O')8GP22$1* 7>)$-;+.>)" 4!E';0*"7N0+ !"#$,6*)*;7'+BG2H,$-71$-;E?5'"<+1 IOE+1* @*;<3
'!+7>J+/)7+&-'"2*3*1 !"#$,6*'!+4 *;40'7;&-'@F52B2:4%'+79 $;GP-"2$7'+BG2H71*I489';=0 ,6*<@57'+BG2HB95 !O%E*21 I,O1 I7,&'+95E; 7G))H *1(&N,$-B95 !O !"#$3I#0"/)<@51* %EO%F21* 7?5*''1?'"#* /0*+7G))H:9;E8T$.'1,$C , *+#>' H4 7>J+B>';0*";*1)6*O*1 #0"/)<@51* 9=9GP2B'''+B95+5';)"
!"#$,6*<@571891* 7>)$-;+.>)":% "# 5*"?'":% :2:G2B95"0*; )!1KLI,$-7>)$-;+.>)"71893*11* ,$-:% :2:G2Q=1,6*<@5.41@!17+&-'"3*1 !"#$ .)5E2$1* 7N&-'240'1!+ @ &''*3,6*<@5N8S+#0E+?*9@*;B> (deletion) 1AB95 1* ?*9@*;B>?'":% :2:G2,6*<@5;$+36*+E+@+P-"?*9@*;B>95E; #6*@ !O1* 7N&-'240'1!+ Q5*7N&-'240'1!+7@2&'+79821AB202$1* 7>)$-;+.>)"7189?PS+1!O#8-"2$N$E84+!S+ R .401*
9
7N&-'2';0*"/89>148 7N0+ 2$N8S+#0E+:% :2:G27(8-2?PS+2* (duplication) 1* 7N&-'2.OO1)!O,8U,*" (inversion) @ &'71891* 7N&-'2#)!O%=0 I@E0*":% :2:G240*"%=01!+ (translocation) 3I7>J+/)<@5#8-"2$N$E84+!S+ R 71891* 7>)$-;+.>)"B95 (7U K_E!N , 2536; #8 +FN, 2536) 1* 7>)$-;+.>)"+$S3I2$/)2*1+5';40*"1!+B>?PS+';=01!ON+89?'";$+ .)I36*+E+;$+,$-71$-;E?5'"
1* X*; !"#$<@51!O(&NB20E0*3I7>J+.OO7X$;O()!+@ &'.OO7 &S' !" 1!O#0E+@+P-"#0E+<9?'"(&N@ &'(&N,$-16*)!"73 8Y748O:4 3I,6*<@57189%E*2/89>148';0*"<9';0*"@+P-"@ &'@)*;';0*"( 5'21!+B95 %E*2/89>1489!"1)0*E'*37>J+B>,!S"/)9$@ &'/)7#$; GP-"'*37>J+> I:;N+H@ &'B207>J+> I:;N+H1AB95 GP-"%E*2/89>1489!"1)0*E#*2* Q.O0"''1B959!"+$S (' FL$, 2530?)
1. /)?'" !"#$,$-2$40'#!L_*+E8,;*?'"(&N
%E*2/89>14840*" R ,$-7189?PS+1!O => 0*")!1KLI?'"(&N,$-B95 !O !"#$ '*37189?PS+1!O *1 )6*45+ <O 9'1 /) #0"/)<@5(&N71891* .% I.1 + %&' 45+74$S;)" <O7)A1)" ?5'>)5'"#!S+ ,!S"+$S7+&-'"3*1 !"#$B>;!O;!S"1* .O0"7G))HO 87EL;'9 .)I;!O;!S"1* ;&94!E?'"7G))HO 87EL?5'>)5'"
2. /)?'" !"#$,$-2$40'1* 7>)$-;+.>)"#$ (&N,$-B95 !O !"#$7'1GH @ &' !"#$.122* 2!171891* 7>)$-;+.>)"71$-;E1!O#$ :9;7X(*I
';0*";8-"#$<O 7 $;1E0* “mosaic” #0E+<@Y07189?PS+7+&-'"3*11* 1 I3*;4!E?'"%)': Cb))HB20#2-6*7#2'
+'13*1+$S'*3(OE0*<O7>)$-;+#$7?$;EB>7>J+#$'&-+ R 7N0+ #$7@)&'"@ &'#$?*E 7>J+45+ +'13*1+$S#0E+'&-+?'"(&N1A#*2* Q71891* 7>)$-;+#$B957N0+79$;E1!+ 7N0+ #$?'"45+ #$9'1
3. 1* ;!O;!S"1* 73 8Y748O:4
!"#$B>;!O;!S"1* 73 8Y748O:4?'"(&N7>J+/)2*3*17+&S'7;&-'73 8YQ=1,6*)*;:9;4 " 4. 1* 7 0"1* 73 8Y748O:4
2$ *;"*+E0* !"#$<+> 82*L4-6* R #*2* QB>7 0"1* 73 8Y748O:4?'"(&NB95 > 82*L !"#$,$-B>7 0"1* 73 8Y748O:4B95%&' > 82*L !"#$4-6*1E0* 5000 7 8+4H71+ 1* .#9"''1QP"1* 7 0"1* 73 8Y748O:4'*3.#9"''12*<+)!1KLI40*" R 7N0+ 45+(&N#="?PS+ 2$ *1;*E "'17 AE ''19'17 AE
/)/)84#=" @ &'2$'*;F1* 71AO71$-;E#!S+)" 7>J+45+
-(./0123!"#$&"#3/45%-&2 &()F(3)2-'"&2 LB5:%&LB5;),:2
"*+E83!;,$-,6*1* X*; !"#$<@51!O(&NN+8940*"R <+> I7,UB,; +0*3I7 8-22$/=5UP1K*2*4!S".40 (.U. 2503 @)!"3*12@*E8,;*)!;71K4 U*#4 H B95 !O'+F2!48<@53!94!S"@+0E;()!""*+> 2*L=?PS+ 7(&-'E83!;71$-;E1!O1* <N5()!""*++8E7%)$; H7(&-'#!+48<+,*"71K4 .)IN$EE8,;* #6*@ !O1* UP1K*E83!;1!OB259'1B25> I9!O 7 8-22$"*+E83!;71$-;E1!O1* <N5 !"#$1!OB259'1B25> I9!O <+>` 2505 @)!"3*17>b9<N57 &'+ F1? !"#$.)5E (' FL$ , 2550) :9;#0E+<@Y07>J+UP1K*QP"/)?'" !"#$.122*,$-2$40'1* 7>)$-;+.>)")!1KLI40*" R ?'"B259'1B25> I9!O :9;,6*1* X*; !"#$.122*<@51!OB259'1B25> I9!O .)5E#!"714)!1KLI,$-2$%E*27>)$-;+.>)"B> .)I> 82*L !"#$7,0*<9,$-2$%E*27@2*I#240'
10
1* X*; !"#$<@51!O B259'1B25> I9!ON+89+!S+ R .)5E2$)!1KLI,$-7>)$-;+.>)"B>:9;,$-45+(&NB204*;
.)I<+1* UP1K*N0E". 13I7>J+(&S+_*+?'""*+E83!;,$-71$-;E1!O1* X*; !"#$7(&-'> !O> F"(!+TFH(&N<+'+*%440'B>
OFY2$ (2518) UP1K*/)?'" !"#$.122*3*1 Cs137
,$-2$40'1* 7>)$-;+.>)")!1KLI1* 73 8Y748O:4?'"45+.)I9'1?'"@+5*E!E3!1 ( 98c :9;+6*45+@+5*E!E2*X*; !"#$<+'!4 * 800, 1,200,
1,600, 2,000, 2,400 .)I 3,200 roentgens /)1* ,9)'"(OE0* 1* 73 8Y748O:4?'"45+@+5*E!E2$.+E:+52)9)"72&-'> 82*L !"#$.122*,$-X*;B>7(8-2?PS+ > 82*L !"#$.122*,$-2$/)40')!1KLI3*+ '"9'1.)I>)$?'"9'1@+5*E!E2*1,$-#F9 %&' 1,200-1,600 roentgens > 82*L !"#$.122*,$-2$/)40')!1KLI?'"<O2*1,$-#F9 %&' 1,600-2,000 roentgens > 82*L !"#$.122* 1,200 roentgens 2$/)40'1* ,6*<@5 9'1@+5*E!E7>J+@2!+ .)I> 82*L !"#$.122* 3,200 roentgens ,$-#*2* Q,6*<@545+@+5*E!E4*;B95QP" 60 7>' H7GA+4H ?PS+B> 3*11* UP1K*(OE0*1* 7>)$-;+.>)"?'"9'1@+5*E!E3I2$)!1KLI,$-B20%",$- .)I)!1KLI?'"3*+ '"9'1 >)$ .)I<O ?'"@+5*E!E3IB20%",$- ' E L (2524) UP1K*/)?'" !"#$.122*,$-2$40'.1)98:')!# :9;1* +6*@!E.1)98:')!#
2*X*; !"#$.122*,$-> 82*L 1,000, 2,000, 3,000, 4,000 .)I 5,000 . 9 .)5E+6*B>>)=1 /)1* ,9)'"(OE0* > 82*L !"#$.122* 2,000 . 9 2$/),6*<@57(8-2%E*2;*E?'"N0'9'1 ?+*99'1
36*+E+9'140'N0' .)I F0+'*;F1* ''19'1?'".1)98:')!# > 82*L !"#$.122* 1,000 . 9 2$/),6*<@57(8-21* 73 8Y748O:4?'"45+ .)I,6*<@536*+E+@!E.)I@!E;0';2*11E0*> 82*L !"#$'&-+ R #6*@ !O> 82*L !"#$.122* 5,000 . 9 2$/),6*<@51* 73 8Y748O:4?'"45+)9)" <O2$%E*2/89>148 .)IB20''19'1
4!S".40>` (.U. 2526 7 8-22$1* +6*(&NN+8940*"R :9;7X(*IB259'1B25B25> I9!O<+#M*(>)'97N&S'2*,9)'"X*; !"#$.122*7(&-'UP1K*1* 7>)$-;+.>)"?'")!1KLI40*"R ?'"(&NN+89+!S+
#F E8N (2526) UP1K*/)?'" !"#$.122*,$-2$40'1F@)*O@8+(!+TFH)* H:1<+#M*(>)'97N&S' :9;X*; !"#$.122*<@51!O1F@)*O(!+TFH)* H:1> 82*L 0, 0.5, 1.0, 1.5, 2.0, 2.5, 3.0, 3.5, 4.0, 4.5 .)I 5.0 71 ;H /)1* UP1K*(OE0*72&-'> 82*L !"#$7(8-2?PS+ 3I,6*<@51F@)*O@8+(!+TFH)* H:12$.+E:+52)9)" > 82*L !"#$.122* 0.5 71 ;H ,6*<@5'!4 *1* 7189;'9#="?PS+ )!1KLI<O 1* 7 $;"4!E?'"<O %E*2;*E>)5'" .)I1* .4118-"?5*"/89>148 .)I> 82*L !"#$.122* 3.5 71 ;H ,$-,6*<@57189)!1KLI/89>1482*1,$-#F9 .40B20(O1* 7>)$-;+.>)"?'"36*+E+:% :2:G2
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
11
%E*2#="45+ 7#5+/0*U=+;H1)*"9'1 .)I?+*9<O?'"45+ B20.4140*"1!+1!O control .)I3*11* UP1K*>)*;;'9 %E*2;*E?'">*1<O .)I36*+E+:% :2:G2,$->)*; *1B95/)B2040*"1!O
control 7N0+1!+ .40#$9'1<+1* ,9)'",$-X*; !"#$.OO7X$;O()!+ 1 .)I 3 . 9 2$1* 7>)$-;+.>)"%&' 3*19'1#$?*E1)*;7>J+9'1#$?*E?$9N2(= " '" (2528) UP1K*/)?'" !"#$,$-2$40'7;'O$ *#*;(!+TFH;F: > :9;,6*1* 7(*I72)A97;'O$ * <+'*@* #=4 MS ,$-7482 kinetin 10 28))81 !2 40')84 .)I IAA 0.5 28))81 !240')84 72&-'7189@+0'3P"+6*B>X*; !"#$.122*> 82*L 0, 1, 2, 3, 4, 5, 6, 7, 8, 9, .)I 10 . 9 (OE0*72&-'> 82*L !"#$.122*7(8-22*1?PS+ '!4 *1* '94*;3I)9)" .40> 82*L !"#$.122*4!S".40 5-10 . 9 ,6*<@5'!4 *1* 4*;#="?PS+ 100 7>' H7GA+4H .4072&-',6*1* X*; !"#$.122*<+'!4 *79$;E1!+.4036*+E+% !S"40*"1!+
:9;,$-+6*@+0'7;'O$ *B>'*O !"#$'!4 * 1 . 9 36*+E+ 1, 2 .)I 3 % !S" .40)I% !S"@0*"1!+ 15 E!+
(OE0*7189)!1KLI/89>14840*"R 7N0+ 1* .411'7>J+1 I3F1 <O.;17>J+ 2 .X1 <O@+*? F? I @ &'<O/'2;*E O*"9'12$#$.9"71897>J+7#5+@ &'.QOO+1)$O9'11)$O<91)$O@+P-" O*"9'12$?+*9<@Y01E0*>148 72&-',6*1* UP1K*36*+E+:% :2:G2(OE0*36*+E+:% :2:G2O 87EL>)*; *12$%0*B20.4140*"3*145+>148 #F2+* (2528) UP1K*1* 1)*;(!+TFH?'"O$:17+$;:9;1* 7(*I7)$S;"7+&S'7;&-' 0E21!O1* X*; !"#$.122* :9;+6*72)A9O$:17+$;(!+TFH Bella Vista 2*7(*I<+#=4 '*@* MS ,$-7482 NAA .)I BA
';0*")I 1 ppm .)I7)$S;"<+'*@* #=4 MS ,$-7482 NAA 1 ppm .)I kinetin 5 ppm .)5E+6*B>X*; !"#$<+> 82*L 0, 2, 4, 6, 8 .)I 10 . 9 4!9.O0";'93+B95 F0+,$- 3 .)5E+6*''1>)=1 /)1* UP1K*(OE0* 45+#0E+<@Y0>148 2$O*"45+,$-2$)!1KLI?5'>)5'";&9;*E<+N0E". 1.)5E40'2*3I73 8Y>148 #6*@ !O45+,$-B95 !O> 82*L !"#$,$-.4140*"1!+ 3I(O)!1KLI,$-1)*;(!+TFH %&' (O45+74$S; 45+.% I 45+,$-2$<O90*" 45+,$-2$<O@+*/89>148 .)I)!1KLI?'"7G))H>*1<O?+*9<@Y0<@Y01E0*45+ control
71&'O#'"7,0* .)I#$?'"9'1O$:17+$;2$%E*2.4140*"1!+2*1 .)I?+*9?'"9'12$,!S"?+*9,$-<@Y01E0*7982.)I7)A1)"1E0*7982
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
NF48+, (2531) UP1K*/)?'" !"#$.122*,$-2$40'7OY32*U 9 (!+TFH :9;1* +6*7OY32*U2*X*; !"#$.122*.OO7X$;O()!+ > 82*L 0.5 .)I 1 . 9 36*+E+ 1 .)I 2 % !S" 3*1+!S++6*2*4!9.O0"7>J+?5' .)5E7)$S;"<+ MS ,$-7482+S6*2I( 5*E 20 7>' H7GA+4H .)I+S6*4*) 40 1 !240')84 .)I#!"7141* 73 8Y748O:4 (OE0* 1* 73 8Y748O:43I)9+5';)"4*2> 82*L !"#$.)I36*+E+% !S"?'" !"#$,$-7(8-2?PS+ .)I10'<@57189)!1KLI/89>148 7N0+ ?5'Q$-#!S+ <O7@)&'" .)I45+,$-B95 !O !"#$> 82*L 1 . 9
12
(OE0*2$1* 73 8Y748O:4,$-+5';2*1 B20#*2* Q+6*''12*>)=1B95 ;17E5+ Yellow Cappa,
Hakutengo .)I Murera (White) +6*45+ F0+,$- 1 >)=172&-'B95 F0+,$- 2 (OE0*7>' H7GA+4H1* ';=0 '93I)9)"72&-'> 82*L !"#$.)I36*+E+% !S"?'" !"#$7(8-2?PS+ )!1KLI?'"9'17OY32*U3*145+,$-X*; !"#$ (OE0*7#5+/0*+U=+;H1)*"?'"15*+9'1 _*+ '"9'1 9'1 9'1N!S+<+ 36*+E+9'1N!S++'1 36*+E+9'1N!S+<+ '*;F1* >Z1.31!+ > 82*L+S6*,$-9=9B><N5 2$.+E:+527(8-2?PS+ +'13*1+$S !"#$#*2* Q,6*<@571891* 7>)$-;+.>)"#$9'1<+7OY32*U 5 (!+TFH 3*1 9 (!+TFH B95/)9!"+$S 7OY32*U(!+TFH Yellow
Cappa 72&-'X*; !"#$ 0.5 . 9 36*+E+ 1 .)I 2 % !S" 2$1* 7>)$-;+.>)"3*1#$7@)&'"7>J+#$#+82 ,$-> 82*L !"#$7,0*1!+ 7OY32*U(!+TFH Ping Pong 3*17982#$?*E7>J+#$N2(= 7OY32*U(!+TFH Hakutengo 7>)$-;+.>)"3*1#$?*E7>J+#$N2(= 72&-'B95 !O> 82*L !"#$ 0.5 . 9 36*+E+ 2 % !S" 7OY32*U(!+TFH Ta Shin Hon 72&-'B95 !O !"#$> 82*L 0.5 . 9 36*+E+ 1 % !S" 2$1* 7>)$-;+.>)"#$9'13*1#$20E"7>J+20E"+S6*4*) 36*+E+ 1 .)I 2 % !S" .)I7OY32*U Murera 72&-'B95 !O> 82*L !"#$ 0.5 .)I 1 . 9 36*+E+ 1 .)I 2% !S" 2$1* 7>)$-;+.>)"#$9'13*1#$?*E7>J+#$N2(= 7@)&'" N2(=#)!O 8SE+S6*4*) .)I#$+S6*4*)'0'+ .)I<+>`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control .)I72&-'B9545+ F0+,$- 3 (OE0*2$)!1KLI =>, "?'"9'1/89>148 #$?'"1)$O9'1N!S++'12$1* 7>)$-;+.>)"3*1#$?*E7>J+#$7@)&'".)I#$?*E.QO7@)&'" .)I72&-'+6*45+ F0+,$- 4
.)I 5 2*>)=1 (OE0*1* 1)*;(!+TFH?'"#$9'1N!S++'1;!"%"';=0 .)I1* 1 I3*;#$9'12$.+E:+52)9)" #6*@ !O?+*9?'"9'1 36*+E+9'1N!S++'1.)I9'1N!S+<+3IB20.4140*"3*1 control .)I3*11* UP1K*:% :2:G2O 87EL>)*; *1 (OE0*:% :2:G2?'"7OY32*U,$-/0*+1* X*; !"#$45+@+P-"7,0*1!O 57 #0E+45+,$-7@)&''$1#*245+2$36*+E+:% :2:G2 7,0*1!O 54
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
13
9'1G5'+#$N2(=7?52?'O1)$O?*E 9'1G5'+#$O*+7;A+1)$O9'17)A17>J+['; 9'1)*#$O*+7;A+ .)I9'1)*#$N2(=?*E
7% &'E!);H (2541) UP1K*/)?'" !"#$.122*40'1* 7(*I7)$S;"7+&S'7;&-'?'"45+>,F22* :9;+6*45+>,F22*,$-B953*11* 7(*I7)$S;"7+&S'7;&-'2*X*; !"#$.122*N+897X$;O()!+ <+> 82*L 0, 1, 3, 5,
7, 9 .)I 11 18:). 9.)5E +6*2*.;17>J+45+7)$S;"<+'*@* #=4 MS ,$-2$ MS E84*28+ .)I7482 BA 5
28))81 !240')84 +S6*4*) 30 1 !240')84 (OE0*'!4 *1* 4*; 50 7>' H7GA+4H 7,0*1!O> 82*L !"#$ 2.8
18:). 9 .)I> 82*L !"#$ 1 18:). 9 3I2$#0E+#=" 36*+E+ .)I)!1KLI40*"R <1)57%$;"1!O45+,$-B20B95 !O !"#$ 45+,$-B95 !O !"#$> 82*L 3 18:). 9 3I2$1* 73 8Y748O:4)9)" 45+,$-B95 !O> 82*L !"#$ 3, 5,
7, 9 .)I 11 18:). 9 3I7189)!1KLI/89>148 7N0+ <O7@)&'" <OB202$#$ <ONS6* <O@+* .?A" 1 'O B207 $;O 3*1+!S++6*2*7)$S;"3+B95 F0+,$- 2 (OE0*> 82*L !"#$ 5, 7, 9 .)I 11 18:). 9 4*;7+&-'"3*11* >+7>de'+3*1.O%,$7 $; @)!"3*1+!S+72&-'45+7982.41@+0'<@20 1A+6*B>7)$S;"<+'*@* #=4 79827>J+ F0+,$- 3 .)5E7,$;O1!O F0+,$- 1 (OE0* 7>' H7GA+4H1* 7(8-236*+E+#="?PS+ 7,0*1!O 242.19 .)I 209.47
7>' H7GA+4H 72&-'+6*2*E!9#0E+#=".)I36*+E+@+0',$-7189?PS+ (OE0*,$-> 82*L !"#$ 0 .)I 1 18:). 9 B20.4140*"1!+ #0E+,$-> 82*L !"#$ 3 18:). 9 3I2$#0E+#=".)I36*+E+@+0',$-7189?PS+4-6*)" +")!1KLH (2541) UP1K*/)?'" !"#$.122*,$-2$40'O$:17+$;7 A1GH :9;+6*;'9O$:17+$;7 A1GH ,$-73 8Y<+#M*(>)'97N&S',$-2$#=4 '*@* MS B>X*; !"#$> 82*L 0-100 71 ;H .)5E4!9#0E+./0+<O
.)I15*+<OB>7)$S;"O+'*@* #=4 MS ,$-7482 BA 1 28))81 !240')84 .)I NAA 0.1 28))81 !240')84 /)1* ,9)'"(OE0*7>' H7GA+4H1* ';=0 '9.)I1* 73 8Y748O:4)9)"72&-'> 82*L !"#$7(8-2?PS+
> 82*L !"#$4!S".40 40 -100 71 ;H ?PS+B>,6*<@571891* 4*; 100 7>' H7GA+4H .)I72&-'7> $;O7,$;O1* 73 8Y748O:41)F02,$-X*; !"#$1!O1)F02 control (OE0*1)F02,$-X*; !"#$2$1* 73 8Y748O:4+5';1E0*1)F02
control .)I(O)!1KLI,$-/89>148<+1)F02O$:17+$;7 A1GH,$-X*; !"#$> 82*L 20 71 ;H <+ F0+,$- 4 %&'
<O1)2 <O;*E7 $;E <O,$-2$#$.9"7?52.)I2$> I#$?*E2*11E0*>148 @ &')!1KLI,$-7189;'97>J+1 I3F1
.4072&-'UP1K*%E*2;*E7G))H>*1<O.)I36*+E+:% :2:G2?'"45+,$-B95 !O1* X*; !"#$1!O45+>148(OE0*B20.4140*"1!+
U8 8U!198c (2542) UP1K*/)?'" !"#$,$-2$40'1* 1)*;(!+TFH?'"O!E@)E" (!+TFH#!44OFK;H ,$-7)$S;"<+#M*(>)'97N&S' :9;,6*1* UP1K*@*#=4 '*@* ,$- 7@2*I40'1* 7)$S;"7+&S'7;&-'O!E@)E" (!+TFH#!44OFK;H <+#M*(>)'97N&S' :9;#=4 ,$-7@2*I#2 %&' ½ MS ,$-7482 IAA 3 fM 0E21!O 2iP 15
fM .)I?;*;(!+TFH:9;E8T$1* >Z1N6*:9;<N5#=4 '*@* 7982 .)5E+6*B>X*; !"#$7(&-'N!1+6*<@571891* 1)*;(!+TFH :9;1* X*; !"#$7'1GH.)I.122* (OE0*1)F02,$-X*; !"#$.122*3I7>J+.OO7X$;O()!+3I<N5> 82*L !"#$ 0, 2, 3, 4, 5, 6 . 9 #0E+1)F02,$-X*; !"#$7'1GH3I<N5 !"#$> 82*L 0, 1, 2, 3, 4 .)I 5
. 9 /)1* UP1K*(OE0* !"#$.122*.)I !"#$7'1GH ,$-> 82*L !"#$ 2 . 9 ,6*<@57189)!1KLI<@20 21
)!1KLI 45+,$-2$)!1KLI/89>14871893*145+,$-B95 !O1* X*; !"#$.122*.)I !"#$7'1GH > 82*L 1-2
. 9 (O45+,$-2$)!1KLI1* 73 8Y748O:49$1E0*45+>148 2$ *1.?+";*E B@)#2O= LH.)I<@545+<@20,$-2*11E0* #0E+45+,$-B95 !O1* X*; !"#$.122*.)I !"#$7'1GH 3-5 . 9 45+#0E+<@Y03I.#9")!1KLI
14
/89>148 GP-"7>J+)!1KLI,$-B20#0"7# 821* 73 8Y748O:4 7N0+ 7>J+.15E (vitrification) G$97@)&'",!S"45+
15*+<O.O+25E+(!+1!+7>J+15'+ B20.414*?5*" B202$ *1.)I.411' 45+,$-B95 !O !"#$> 82*L.122* 6 . 9 3I4*;@29M*;<+ 4 #!>9*@H 72&-'UP1K*:% :2:G2O 87EL>)*; *1(O 2n=16 !"#$,$-,6*<@5718945+1)*;(!+TFH,$-2$:% :2:G2 anueploid 2$ 2 1)F02 %&' 1)F02,$-2$36*+E+:% :2:G2 2n=18 .)I 2n=20
U8 8U!198c .)I E8N!; (2543) UP1K*/)?'" !"#$40'1* 1)*;(!+TFH?'"O!E@)E"(!+TFHOFLg 81
:9;,6*1* X*; !"#$.122*> 82*L 0, 2, 3, 4, 5 .)I 6 krad .)I7)$S;"7>J+7E)* 12 #!>9*@H /)1* UP1K*(OE0* !"#$.122*,$-7@2*I#2#*2* QN!1+6*<@545+O!E@)E"7189)!1KLI/89>148 %&' 0-2
krad )!1KLI,$-/89>148 %&' 2$?+*97)A11E0*45+,$-B20B95 !O1* X*; !"#$ 2$3F9> I?+*97)A1.)I2$.QO#$?+*9<@Y0O+<O 7N0+ #$7?$;E #$7@)&'" #0E+> 82*L !"#$,$- 4, 5 .)I 6 krad 4*;,!S"@29M*;<+ 2
#!>9*@H /)?'"36*+E+:% :2:G2 2n=16 ?+*9>*1<O 9.78 B2% '+
!N+$;H (2544) UP1K*1* N!1+6*7U K_$15*+,'"<@51)*;(!+TFH:9;E8T$1* X*; !"#$.122* :9;1* +6*72)A92*X*; !"#$.122*.OO7X$;O()!+ <N5> 82*L !"#$ 0, 10, 20, 40, 80, 160, 240 .)I 320 71 ;H /)1* UP1K*(OE0*72&-'> 82*L !"#$7(8-2?PS+'!4 *1* "'1?'"72)A9)9)" 72)A9"'1N5*)".)IB20#*2* Q73 8Y748O:440'B>B95 72&-',6*1* X*; !"#$ % !S",$- 2 <N5> 82*L !"#$ 0, 80, 90, 100
.)I 110 71 ;H /)1* UP1K*(OE0* > 82*L !"#$,$- I9!O 90 71 ;H ,6*<@545+1)5*2$N$E84 '9B95 50
7>' H7GA+4H .)I(O1* 7>)$-;+.>)"?'"#$./0+<O?'"1)F0245+1)5*,$-X*; !"#$ 80 90 .)I 110 71 ;H E: M*# (2544) UP1K*/)?'" !"#$.122*40'1* 73 8Y748O:4.)I1* /)84@!E;0';?'")8))$-(!+TFH Red Night .)I(!+TFH Apeldoorn <+#M*(>)'97N&S' :9;+6*@!E;0';?'")8))$-(!+TFH Red Night
.)I(!+TFH Apeldoorn 2*X*; !"#$,$-> 82*L 0, 2, 3, 4, 6 .)I 8 71 ;H .)5E+6*2*7)$S;"<+#=4 '*@* MS ,$-7482 NAA 0.1 28))81 !240')84 +S6*4*) 60 1 !240')84 /)1* ,9)'"(OE0* (!+TFH Red Night
3*11* .;1B>7(*I7)$S;":9;1* /0*% P-"@!E (OE0*,$-> 82*L !"#$.122* 0-4 71 ;H 2$1* 73 8Y748O:4B95 #0E+@!E,$-B95 !O> 82*L !"#$ 6-8 71 ;H B202$1* 7>)$-;+.>)"M*;<+ 30 E!+ Q5*.;1''17>J+1)$O@!E,$-B95 !O !"#$,$-> 82*L 0-8 71 ;H B202$1* 71897>J+@!E;0'; 7+&-'"3*1@!E;0';,$-+6*2*7(*I7)$S;"+!S+2$?+*97)A1718+B> 45'"<N57E)*+*+<+1* 73 8Y7>J+@!E;0';
1hKL* (2545) UP1K*1* N!1+6*<@571891* 1)*;(!+TFH<+ Ornithogalum dubium Houtt.
:9;+6*<O'0'+@ &'_*+<O,$-4891!O3F916*7+89 *12*7(*I7)$S;"O+#=4 '*@* MS #0E+,$-N!1+6*<@57189;'9<@20B959$ %&' <O'0'+ 72&-'7(*I7)$S;"<O'0'+O+#=4 '*@* MS ,$-7482 kinetin 2 28))81 !240')84 0E21!O NAA 0.1 28))81 !2 .)I+S6*4*)G=:% # 30 1 !240')84 #*2* QN!1+6*<@57189;'9<@20B959$ .)I#=4 '*@* ,$-N!1+6*<@57189@!E.)I *1B959$ %&' #=4 '*@* MS ,$-7482 kinetin 2 28))81 !240')84 0E21!O NAA 0.2 28))81 !240')84 .)I+S6*4*)G=:% # 60 1 !2 40')84 .)I+6*;'9'0'+B>X*; !"#$.122*.OO7X$;O()!+> 82*L 0, 1, 2, 4, 6, 8, 10, 12, 14 .)I 16 18:). 9 /)1* ,9)'"(OE0*> 82*L !"#$,$-,6*<@5'!4 *1* 73 8Y748O:4)9)"% P-"@+P-" %&' 2.21 18:). 9 > 82*L !"#$,$-7(8-2?PS+2$/)
15
,6*<@51* 73 8Y748O:4?'";'9)9)" > 82*L !"#$4!S".40 4 18:). 9?PS+B> 3I;!O;!S"1* 73 8Y748O:4.)I;'93I4*;<+,$-#F9
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
23.54 71 ;H > 82*L !"#$,$-,6*<@5'!4 *1* 73 8Y748O:4?'">,F22*(!+TFH(&S+72&'")9)" 50 7>' H7GA+4H %&' 37.56 71 ;H 3*11* #!"714)!1KLI/89>148(OE0*>,F22*,$-B95 !O !"#$ 30 .)I 50 71 ;H ?'",!S" 2 #*;(!+TFH3I2$1* 73 8Y748O:4)9)"2*172&-'7,$;O1!O>,F22*,$-B95 !O> 82*L !"#$ 0 71 ;H .)I7189)!1KLI/89>148 %&' <O7@)&'"G$9 #0E+45+>,F22*,$-B95 !O> 82*L !"#$4!S".40 60 71 ;H ?PS+B>B20(O1* 73 8Y748O:4 ,!S" 2 #*;(!+TFH
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
'!YNF)$ (2546) UP1K*#=4 '*@* ,$-7@2*I#2.)I/)?'" !"#$.122*,$-2$40')!1KLI,*"#!L_*+E8,;*?'"1)5E;O!E#$#52 :9;+6*>)*;;'91)5E;O!E#$#52B>7(*I7)$S;"<+'*@* #=4 MS
,$-7482 6-Benzylaminopurine (BA) %E*27?52?5+ 2.5, 5.0, 7.5 .)I 10 ppm (OE0* '*@* #=4 MS ,$-7482 BA %E*27?52?5+ 5.0 .)I 7.5 2$%E*27@2*I#240'1* (!]+*?'"@+0' 3*1+!S++6*>)*;;'91)5E;O!E#$#52<+#M*(>)'97N&S'2*X*; !"#$.122* ,$-> 82*L !"#$ 0, 5, 10, 15, 20, 25 .)I 30
71 ;H (OE0*,$-> 82*L !"#$7(8-2?PS+2$1* '9N$E84 .)I1* .41@+0')9)" 7>' H7GA+4H1* 4*;,$- 30
.)I 50 7>' H7GA+4H %&' 25.5 .)I 28.5 71 ;H +6*>)*;;'91)5E;O!E#$#52B>>)=1 I;I7E)* 8 79&'+
45+,$-B20 !O1* X*; !"#$2$%E*2#="?'")6*45+7,$;2#="#F9 %&' 68.19 7G+48724 1)5E;O!E,$-B95 !O> 82*L !"#$ 30 71 ;H 2$36*+E+<O.)I36*+E+@+0'#="#F9 29.71 <O .)I 6.5 @+0' <+?LI,$- I;I7E)*>)=13+QP"<OT"%)$-.)I I;I7E)*>)=13+QP"''19'1?'"45+1)5E;O!E#$#52,$- !O> 82*L !"#$.122* 30 71 ;H<N57E)*#!S+,$-#F9 197.14 .)I 198.24 E!+ #0E+1)5E;O!E,$-B95 !O> 82*L !"#$ 5 71 ;H<N5 I;I7E)*9'1O*+O+45++*+,$-#F9 77.67 E!+ 72&-'UP1K*)!1KLI,*"#!L_*+E8,;*(O1)5E;O!E,$-
16
B95 !O> 82*L !"#$ 20 71 ;H 36*+E+ 2 45+ ,$-2$%E*2/89>148.)I#*2* QQ0*;,'9B>;!" F0+40'B>B95 :9;,$-45+. 12$)6*45+7,$;290*")*;B>3+QP" 0'"<O@29 45+,$-#'"2$)6*45+> I#$+S6*4*).)I1*O<O.;13*1)6*45+7,$;2
(8N84 (2546) UP1K*/)?'" !"#$.122*40'1* 73 8Y748O:4.)I)!1KLI,*"#!L_*+?'"O*+N&-+@+= :9;1* +6*72)A9O*+N&-+@+= (!+TFH9'1#$7@)&'" 2*X*; !"#$.122*.OO7X$;O()!+
> 82*L 0, 100, 200, 400 .)I 600 71 ;H .)5E+6*B>>)=1 E!9%E*2#=".)IO!+,P1)!1KLI,*"#!L_*+?'"45+O*+N&-+@+= B95.10 7#5+/0*+U=+;H1)*" 36*+E+9'140'45+ 36*+E+1)$O40'9'1 36*+E+9'1 .)I<O40'45+ /)1* UP1K*(OE0*72&-'> 82*L !"#$7(8-2?PS+%E*2"'1?'"72)A93I)9)" > 82*L !"#$ 500 71 ;H ,6*<@5%E*2"'1)9)" 50 7>' H7GA+4H 1* 73 8Y748O:4?'"45+O*+N&-+3IE!995E;%E*2#="?'"45+ (OE0*%E*2#="?'"45+O*+N&-+@+=3I)9)"72&-'> 82*L !"#$7(8-2?PS+
(!+THE9$ (2546) UP1K*/)?'" !"#$.122*,$-2$40'1* 73 8Y748O:4?'":(T8cO!))!"1H<+#M*(>)'97N&S' :9;1* +6*#0E+?'")6*45+:(T8c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
60 E!+ 2*11E0*72)A9,$-B20.1I7>)&'1.)I.N0+S6* 8 N!-E:2" .)I72)A9,$-.1I7>)&'1 1* +6*72)A9,$-.1I7>)&'12*X*; !"#$,6*<@52$'*;F1* ''19'1. 1?'"45+,*+4IE!+ .)I36*+E+<O72&-'71894*9'1. 1+5';,$-#F9 .402$36*+E+9'1@)!"7(*I72)A9 60 E!+2*1,$-#F9
T+*T8> (2547) UP1K*/)?'" !"#$.122*40'1* 7@+$-;E+6*<@571891* 1)*;(!+TFH40'45+9*E1 I3*; :9;1* 7(*I7)$S;"7+&S'7;&-'9*E1 I3*;95E;'*@* #=4 MS ,$-2$ BA 2 28))81 !2/)84 NAA 0.25 28))81 !2/)84 7(&-'N!1+6*<@57189;'9'0'+> 82*L2*110'++6*B>X*; !"#$> 82*L 0, 20,
40, 60, 80 .)I 100 71 ;H 7(&-'@*> 82*L !"#$,$-7@2*I#2<+1* 7@+$-;E+6*<+1* 71891* 1)*;(!+TFH (OE0*(&N F0+. 1 2$1* 73 8Y748O:4)9)"72&-'> 82*L !"#$#="?PS+ > 82*L !"#$,$-,6*<@545+9*E1 I3*;
17
2$1* 73 8Y748O:44-6*#F9 %&' 100 71 ;H > 82*L !"#$,$-,6*<@545+9*E1 I3*;2$'!4 *1* 73 8Y748O:4)9)" 50 7>' H7GA+4H %&' 92.19 71 ;H 72&-'+6*(&N F0+. 12*7)$S;"<+'*@* #=4 MS (OE0*(&N F0+,$- 2
2$1* '9N$E84.)I1* 73 8Y748O:4)9)"72&-'> 82*L !"#$#="?PS+ :9;,$-> 82*L !"#$ 100 71 ;H 3I,6*<@545+9*E1 I3*;2$'!4 *1* '9N$E844-6*#F9 72&-'7,$;O1!O45+,$-B20B95X*; !"#$ #0E+> 82*L !"#$,$-,6*<@545+9*E1 I3*;<+ F0+,$- 2 )9)"% P-"@+P-" %&' 55.90 71 ;H )!1KLI,*"#!L_*+E8,;*,$-/89>148 ,$-(O<+ F0+. 1,$-B95 !O1* X*; !"#$> 82*L 60 71 ;H %&' <O7@)&'" .)I<+ F0+#'",$-B95 !O1* X*; !"#$> 82*L 20, 40 .)I 60 71 ;H %&' @"81"'/89 => @ &'7 $;E;*E
38 I+!+,H (2547) UP1K*/)?'" !"#$.122*,$-2$40'1* 73 8Y748O:4.)I)!1KLI,*"#!L_*+E8,;* ?'" Kalanchoe sp. :9;+6*45+'0'+B>X*; !"#$.122*N+897X$;O()!+ <+> 82*L 0, 20, 40,
60, 80, 100, 150, 200, 250, 300, 350 .)I 400 71 ;H /)1* UP1K*(OE0*> 82*L !"#$,$-,6*<@5'!4 *1* 73 8Y748O:4)9)" 50 7>' H7GA+4H 72&-''*;F 45, 90 .)I 105 E!+ %&' 402.25, 316.06,
263.68 71 ;H .)I1* 73 8Y748O:4:9;E!9%E*2;*E?'"<O GR50(45), GR50(125) %&' 267.25 .)I 258.13 71 ;H .)I(O)!1KLI#!L_*+E8,;*?'"45+1)5*,$-/89>148 %&' <OO897O$S;E 2$)!1KLI7>J+ =>@!E<3 <O.[9 4891!+ <O7 $;E .)I(OE0*> 82*L !"#$ 4!S".40 200 71 ;H?PS+B> 3I2$/)40'1* )91* 73 8Y748O:4.)I%E*2';=0 '9?'"45+ Kalanchoe sp.
18
!!"# 3
$%&"'()*+(,-%-'()$%./0
)123 '()$%./0
!"#$%&'()*+,-( !"#$%&'+.$/0""'!' 123/+,-( !"45(4#5!4#!6%"$/7!689!: !",;%%<0&(=)3,"! >?'@A6!0""'!'BC5+CD(#A!6)E5?+=D%%"$/?FG"=)3,"! >?'@A 6)4#!68&6:&(HI &(?'A!/G" JK'G6A8(B%#A!=L!G6M2/+,-(+.A((&*( C"N?G6A6) !"%&K=L!+:N3?4#04<67&#O,"BK P
!"#$%&'+.$/0""'!' 4N? "@,O00 !"#$%&'=)3G6A6) Q<A6+,")'0+=)'0 +,-( !"R2 S!89!:, 7$E?/7&#O,"7A!/ P +.A( !"R2 S!4#!64$K+CD( 4#!6+.N3?E?/0<44Q C"N? !"R2 S!Q& STF !" "F%!'E?/J"47!6UK@E?/J"4B(.A#/+#Q!C(23/ (,;VV!, 2546)
45(-!"#367)707,$6(8-'()*+(,-%-'()$%./0
1. 6C!#$='!Q&'R")(4"$(="#$J"W ?/4"& SI %. (4"(!'
2. 8L!(& C?86<K6C!#$='!Q&'+ S7"R!87"I #$='!+E70!/+E( %. "</+=:6C!(4" 3. 8L!(& C?86<K6C!#$='!Q&'+ S7"R!87"I #$='!+E7 L!O:/O8( %. (4",X6
4. R@('I0"$ !"Y!'"&/8)O 66!OQF#$%&'($#+4Q)'"I+=4J(JQ') 8M!0&(#$%&'OQF:&W(! 6C!#$='!Q&'+ S7"R!87"I
5. 8L!(& /!(,"6!T@+:N3?8&(7$ "F="#/#$='!R!87"IOQF+=4J(JQ')
"F'F+#Q!B( !"B( !"KL!+($( !"#$%&' 7&*/O7A+KN?( <69!:&(HI 2550 M2/+KN?( 8$/C!46
:.R. 2550
$%&"'()$%./0367,'9 :;<=16
1. OCQA/E5?6@Q ,"F ?0K5#'
1.1 OCQA/45(4#5!/!(#$%&'=)3+ )3'#E5?/ &0 !"B.5"&/8) &0G65K? G65,"FK&09!'B(,"F+=RG=' GK5O A
1.2.1 C(A#'/!(7A!/ P =&*/9!4"&XOQF+? .( =)3KL!+($( !"#$%&'=!/K5!( !"B.5"&/8) 1.2.2 E5?6@Q=)38N045(%! X!(E5?6@Q=!/+4"N?EA!'4?6:$#+7?"I 1.2.3 !"8?0M!6%! Z@5=)3R2 S!#$%&'=!/K5!( !"B.5"&/8) &0G65K? G65,"FK&0
JK'7"/
19
2. ,"F+9=E?/E5?6@Q ,"F ?0K5#'
2.1 E5?6@Q,X69@6$ GK5O AE5?6@Q=)3GK5%! !"8?0M!6%! Z@5=)3R2 S!#$%&'=!/K5!( !"B.5"&/8) &0G65K? G65,"FK&0JK'7"/
2.2 E5?6@Q=<7$'9@6$ GK5O A E5?6@Q%! +? 8!"/!(#$%&'=)36) !"7):$6:IB(#!"8!" OQF/!(#$%&'=)37):$6:IB("@,E?/"!'/!( !"#$%&',"F%L!,[ #$='!($:(HI ,;VC!:$+RS 8$3/:$6:I7A!/ P =&*/=)3+,-(9!S!G='OQF?&/ US
3. +4"N3?/6N?=)3B.5B( !"+ D0"#0"#6E5?6@Q
3.1 +? 8!" 8L!+(!+? 8!" =)3+ )3'#E5?/ &0 !"B.5"&/8) &0G65K? G65,"FK&0
3.2 :!C(F+K$(=!/G,45(C!OQF"#0"#6+? 8!"=)3+ )3'#E5?/B(8M!(=)37A!/ P
$%&"'()*+(,-%-'()
1. "#0"#60=4#!6OQF/!(#$%&'7A!/P =)3+ )3'#E5?/ &0 !"B.5"&/8) &0G65K? G65,"FK&0B(,"F+=RG='
2. =L! !"#$+4"!FCIOQF8"<,E5?6@Q7A!/P =)3+ )3'#E5?/ &0 !"B.5"&/8) &0G65K? G65,"FK&0 +.A(
!"B.5,"FJ'.(I ZQ "F=0=)3+ $KE2*( E5?6@QG65K? G65,"FK&0=)3(L!6!Y!'"&/8) G65K? G65,"FK&0:&(H<IBC6A=)3GK5%! !"Y!'"&/8) +,-(75(
3. =L! !"+E)'(0=($:(HI,"$=&R(I (reviewed paper) + )3'# &0ZQE?/ !"B.5"&/8) &0G65K?
G65,"FK&0 +:N3?(L!+8(?E2*(+#D0G1=IE?/R@('I#$%&'OQF !"%&K !"4#!6"@5=!/:U SR!87"I +:N3?+Z'O:"AO A.<6.(7A?G,
4. !"%&K=L!X!(E5?6@Q+ )3'# &0.($KE?/G65K? G65,"FK&0=)36) !"Y!'"&/8)B(,"F+=RG='
+:N3?+Z'O:"AO A.<6.(7A?G,
20
!!"# 4
$%&'()*+,('-./01234%
!""!##$%#$&'!($) *+,-!("!#./-#*'01"*%2&-,3"2&-4#5,*% #$&6*7'8##92&-(7:!4#5,*%.(4#5;6<26+ 61=2,-&1"!#>18)&8?;@+A8#B#5C$B!'4D 8.<. 2510 – 2550 6*7''!($) *+E3'C(B$+'!(#!/"!#AF5;3"/( .($!#0!#6!'$)/!"!#AF5'!($) *+61=>18)&8?.(#G4E3'#!+'!("!#$) *+4#5 :!4D $)6+!()8(H? 4IJC!8);<K 0)='8)&8?>B!' L 8%$B!
53162&5317(-68 ()3!89:;((<531=9>'7(-68 !"#3"('?:'=&'(@A1(8:B"+;C#2&'(7(8 7(D:;8=ED/@=7(-+!F5!?
2&-,3"2&-4#5,*% #$&6*7'8##92&-(7:!4#5,*% 61=&1#!+'!("!#M!+#*'01;8N=3"!#4#*%4#O'8*(HO?.(4#5;6<26+ 8%$B! &1 :!($( 27 $'<? (family) 50 0"OF (genus) &!""$B! 64 /(), (species)
P,+0B$(.CJB;4Q(2&-,3" :!($( 18 $'<? 37 0"OF &!""$B! 44 /(), (>!#!'61= 1) ;4Q(2&-4#5,*%
:!($( 6 $'<? 8 0"OF &!""$B! 10 /(), (>!#!'61= 2) AF5;4Q(8##92&-(7:!4#5,*% :!($( 3 $'<? 5 0"OF 10 /(), (>!#!'61= 3)
G'(':!"# 1 #!+/N=32&-,3"61=&1"!#<R"K!SR'@FE3'"!#M!+#*'01.(4#5;6<26+
/N=3$)6+!<!0>#? /N=326+ /N=30!&*J /N=3@G-<R"K! 1. Adianum spp. * /$(/& Desert Rose 3#O91 (2546)
3#$##9 (2548)
2. Alpinia purpurata E)'A,' Red ginger ;%J &!< (2541)
;"< (2544)
3. Anthurium andraeanum C(-!$*$ Flamingo flower %OJ&1 (2518)
$)//O>! (2537)
4. Begonia spp. %1P";(1+ Begonia 0O&(! (2528)
('F*"K9? (2541)
5. Brassolaeliocattleya hybrids "F-$+2&-AT6F1+!FG"@0& - T##/)> (2538)
6. Bulbophyllum longissimum * 0)'P>;T#!+!$ - 3#O91 (2546)
7. Calanthe rubens 3*7$8$'&91 - H*($? (2546)
21
G'(':!"# 1 (GH2)
/N=3$)6+!<!0>#? /N=326+ /N=30!&*J /N=3@G-<R"K! 8. Canna indica 8O6H#*"K! Canna ' )(>? (2520)
(*(6(! (2525)
0)#(O/ AF5T95 (2543)
<G(+?%#)"!#M!+#*'01A"&&!AF5$) *+()$;TF1+#?;6TP(PF+1 (2546)
9. Chrysanthemum morifolium ;%J &!< Chrysanthemum /O>)(6# (2531, 2532)
&*FF)"! (2542)
81#(O/ AF5T95 (2544)
<G(+?%#)"!#M!+#*'01A"&&!AF5$) *+()$;TF1+#?;6TP(PF+1 (2546)
Lamseejan et al. (2000)
10. Curcuma alismatifolia * 46O&&! Siam Tulip ;T#N3$*F+? (2541)
$*($)0! (2545)
$)/J! (2546)
3#O91 (2546)
11. Curcuma sparganifolia 46O&&! Siam Tulip 3,)<# (2535)
12. Dendrobium albosanguinium ;3N73'@!;$1+' - H*($? (2546)
13. Den. chrysotoxum ;3N73'T:! - H*($? (2546)
14. Den. cruentum ;3N73'4!"("A"-$ - H*($? (2546)
15. Den. hybrids * "F-$+2&-C$!+FG"@0&8*(HO?>B!' L
-
H(5 (2548)
3#O91 (2546)
16. Dianthus caryophyllus T!#?;(/*=( Carnation /*+/O&8F (2526)
3#,1AF5/*+/O&8F (2527)
17. Etlingera elatior ,!CF! Torch Ginger $)6+!"OF (2543)
3U)/!>) AF5T95 (2543)
)#!U#9? (2545)
$#8()> (2545)
4"#9? (2548)
18. Gladiolus spp. & hybrid A"F,)P3F*0 Gladiolus 3#$##9 (2524)
4#!(3& AF5T95 (2525)
19. Gloriosa superba ,3',R' Climbing lily 8#6)8+? (2542)
22
G'(':!"# 1 (GH2)
/N=3$)6+!<!0>#? /N=326+ /N=30!&*J /N=3@G-<R"K! 20. Grammatophyllum speciosum $B!(;8/#CR' Tiger orchid H*($? (2546)
21. Habenaria militaris * "F-$+2&-"FOB&F)7(&*'"# - 3#O91 (2546)
22. Helianthus anuus 6!(>5$*( Sunflower <)#)#*>(? (2546)
23. Hippeastrum johnsonii $B!(01=6)< Amaryllis 3>)(*(6? (2545)
24. Hygrophylla difformis ,!$"#5 !+ Water Wisteria H(!H)4 (2547)
25. Kalanchoe blossfeldiana "OCF!%C)( Flaming Katy 0O#$)/ (2526)
$*(;MF)& (2546)
)#5(*(6? (2547)
26. Kalanchoe laciniata V-3'0!&+B!( - 3#O91 (2550)
27. Lilium spp. F)FF1= Lily 3!U!U#9? (2539)
$P#U!0 (2544)
28. Musa laterita "F-$+%*$010-& - 3*J/OF1 (2546)
29. Musa sp. "F-$+#*>"*6F1 Wild Callimusa $(),! (2547)
30. Nelumbo lutea %*$CF$'3;&#)"*(
01;CFN3' American yellow
lotus
$)/*+AF5"("8# (2549)
31. Nelumbo nucifera %*$CF$' Sacred lotus ;3&3# (2530)
<)#)<*",)W (2542)
<)#)<*",)WAF5$)/*+ (2543)
Lama et al. (2005)
32. Nymphaea ‘Jongkolnee’ %*$ '"F(1 Water lily Puripunyavanich and
Boonsirichai (2005)
33. Oncidium ‘Popoki’ * 33(X1;,1+&AT#5 - 3#O91 (2546)
34. Ornithogalum dubium 33()P6"!F*& - "YK9! (2545)
35. Petunia hybrida 8)6G;(1+ Petunia 8##91 (2543)
36. Phalaenopsis violacea * Z!AF((34X)0
2$P3#!;X1+
-
H*($? (2546)
3#O91 (2546)
37. Polianthes tuberosa XB3("F)=(26+ Tuberose 3,)<# (2533)
38. Portulaca grandifora A8#;X1=+'2[- Potulaca 3#O91AF5($FM$1 (2534)
<G(+?%#)"!#M!+#*'01A"&&!AF5$) *+()$;TF1+#?;6TP(PF+1 (2546)
Wongpiyasatid and
Homchan (2000)
39. Plumeria spp.* F1F!$,1 Temple tree 3#O91 (2546)
23
G'(':!"# 1 (GH2)
/N=3$)6+!<!0>#? /N=326+ /N=30!&*J /N=3@G-<R"K! 40. Rhynchostylis gigantea * /-!'"#5 - H*($? (2546)
3#O91 (2546)
41. Rosa hybrida "OCF!% Rose 8*/#)(6#? (2538)
42. Schoenorchis fragrans* ;3N73' )\$ - 3#O91 (2546)
43. Vanda hybrids * "F-$+2&-A$(,-!FG"@0& - 3#O91 (2546)
44. Zinnia augustifolia %!(/N=(C(G Narrowleaf Zinnia 8)/)> (2546)
G'(':!"# 2 #!+/N=32&-4#5,*%61=&1"!#<R"K!SR'@FE3'"!#M!+#*'01.(4#5;6<26+
/N=3$)6+!<!0>#? /N=326+ /N=30!&*J /N=3@G-<R"K! 1. Agave sp. 3!"!;$B Century plant 3#O91 (2550)
2. Aglaonema sp.* A"-$"!J (! Aglaonema 8*(H?$,1 (2546)
3#O91 (2546)
3. Chlorophytum amaniense ;<#K]1"-!(63' - #*/(1+? (2544)
4. Dracaena fragrans $!0(! Philodendron
"Red Emerald"
3#O91 (2550)
5. Dracaena sanderiana * 2@B"$(3)& - 3#O91 (2546)
6. Dracaena surculosa 2@BZF)44^(0? Florida beauty 3#O91 (2550)
7. Ficus benjamina var.
variegata
26#+-3+.%ACF&,B!'
Gogen fig
0O/!,! (2550)
8. Musa acuminata ‘BEP’ "F-$+;%8 - 3!#+! (2548)
9. Philodendron sp. &#">A,' - 3#O91 (2550)
10. - 4!F?& - 3#O91 (2550)
24
G'(':!"# 3 #!+/N=38##92&-(7:!4#5,*%61=&1"!#<R"K!SR'@FE3'"!#M!+#*'01.(4#5;6<26+
/N=3$)6+!<!0>#? /N=326+ /N=30!&*J /N=3@G-<R"K! 1. Alternathera reineckii @*";4Q, - #*"/(" AF5T95 (2549)
2. Anubias bartiri * 3(G;%1+0 Anubias ,!$0$##T? (2545)
3#O91 (2546)
3. Cryptocoryne balansae %3((7:! Balansa's
Crytocoryne
0O )>#! (2548)
4. Cryptocoryne blassii %3(A,' Braune von
Bangkok
0!#)(1 (2548)
5. Cryptocoryne wendtii
“green”
.%8!+, $!(7:! Amazon sword
plant
4^_(3O%F (2548)
6. Echinodorus argentinensis 3;&X3( - 3O2# (2542)
7. Echinodorus armazonicus * 3;&X3(.%+!$ - 3#O91 (2546)
8. Echinodorus bleheri * 3;&X3( - 3#O91 (2546)
9. Echinodorus parviflorus * 3;&X3( - 3#O91 (2546)
10. Lobelia cardinalis PF%1;F1+ Lobelia 2>#;&H (2547)
<#!+O6H (2547)
11. Sagittaria pusillus * X! )63;#1+ Sagittaria 3#O91 (2546)
.3'?+.GD * TN3 0B$(C(R=';4Q(E-3&GF61=&1#!+'!(.(;3"0!#4#5"3%"!#`a"3%#& PT#'"!#$) *+;/)'SB!+63,;6TP(PF+1 “"!#8*b(!8*(HO?2&-,3"2&-4#5,*%,-$+#*'01A"&&!0GB;"K>#"#” 61=.C-;"K>#"#61=;E-!#B$&PT#'"!#(:!2&-,3"2&-4#5,*%E3'>(&!M!+#*'01 *,P,+U!T$)/!#*'014#5+O">?AF523PXP64 T95$)6+!<!0>#? #B$&"*%<G(+?%#)"!#M!+#*'01A"&&!AF5$) *+()$;TF1+#?;6TP(PF+1 0S!%*($) *+AF58*b(! &C!$)6+!F*+;"K>#<!0>#? AF5 "#&0B';0#)&"!#;"K># "#56#$';"K>#AF50C"#9? $*(61= 26-27 &)SO(!+( 2546 9 <G(+?%#)"!#M!+#*'01A"&&!AF5$) *+()$;TF1+#?;6TP(PF+1 &C!$)6+!F*+;"K>#<!0>#?
25
79%
3% 3%
1%
1%3%
5%
1%4%
&C!$)6+!F*+;"K>#<!0>#? &C!$)6+!F*+;/1+'.C&B
&C!$)6+!F*+E3(A"B( &C!$)6+!F*+;6TPF+1#!/&'TF
&C!$)6+!F*+0'EF!(T#)(6#? Oc!F'"#9?&C!$)6+!F*+
0S!%*(;6TPF+18#5 3&;"F-!; -!TO96C!#F!,"#5%*' &C!$)6+!F*+;6TPF+18#5 3&;"F-!H(%O#1
0:!(*"'!(4#&!9G;8N=30*(>)
&'(FI&J')*K8?+&"#?)&8 $%02:(8:B"!"#3"GH253162&5317(-68 @=7(-+!F5!? @=AH):7L ;.F. 2510-2550
!""!#<R"K! 8%$B!"!#M!+#*'01.C-A"B2&-,3"2&-4#5,*% 0B$(.CJB&1$*>SO4#50'T?;8N=3"!#4#*%4#O'8*(HO? T),;4Q(#-3+F5 95.56 AF5;8N=34d3'"*(":! *,A&F'<*>#GCF*'"!#;"e%;"1=+$ T),;4Q(#-3+F5 4.44
- .=H)?:'=!"#!>'&'(FI&J'
&C!$)6+!F*+;"K>#<!0>#? ;4Q(C(B$+'!(61=6:!"!#<R"K!$) *+SR'@FE3'#*'0161=&1>B3
2&-,3"2&-4#5,*%.(4#5;6<26+ .(/B$'4D 8.<. 2510-2550 &!"61=0O, T),;4Q(#-3+F5 79 E3'@F'!(6!'$)/!"!#61=>18)&8?;@+A8#B #3'F'&!2,-A"B &C!$)6+!F*+;/1+'.C&B #-3+F5 5 0S!%*(;6TP(PF+18#5 3&;"F- ! ; - !TO96C!#F!,"#5%* ' T), ;4Q(#- 3+F5 4 0: !(*"'!(4#&!9G ;8N= 30*(>) &C!$)6+!F*+0'EF!(T#)(6#? AF5 Oc!F'"#9?&C!$)6+!F*+ T),;4Q(#-3+F5 3 &C!$)6+!F*+;6TP(PF+1#!/&'TF &C!$)6+!F*+8#5 3&;"F-!H(%O#1 AF5&C!$)6+!F*+E3(A"B( T),;4Q(#-3+F5 1
(U!861= 1)
M';!"# 1 C(B$+'!(61=6:!"!#<R"K!$) *+;"1=+$"*%@FE3'#*'0161=&1>B32&-,3"2&-4#5,*%.(4#5;6<26+
.(/B$'4D 8.<. 2510-2550
26
;&Fe, 4%8N/6*7'>-( 3%0B$(E+!+8*(HO?
61=2&B./-;8< 30%
;8!5;F17+'.(0U!84F3,;/N73 63%
A"&&!99%
;3"X?1%
- BH)=02:53162&5317(-68 !"#=>'3'N'?(8:B" 8%$B!0B$(E3'8N/61=(:!&!M!+#*'01 0B$(.CJB;4Q(0B$(E3'8N/61=;F17+'.(0U!84F3,;/N73
(;8!5;F17+';(N73;+N=3) T),;4Q(#-3+F5 63 #3'F'&!TN3 0B$(E+!+8*(HO?61=2&B./-;8<3N=( L ;/B( ")=' F:!>-(
.% ;C'-! C*$ fFf T),;4Q(#-3+F5 30 ./-;&Fe, T),;4Q(#-3+F5 4 AF5./-8N/6*7'>-( T),;4Q(#-3+F5 3
>!&F:!,*% (U!861= 2)
M';!"# 2 /)7(0B$(E3'2&-,3"2&-4#5,*%61=(:!&!M!+#*'01
- (8:B"!"#@A1 )*E"&'(N'?(8:B" O%-7(*3'<(8:B"!"#@A1 "!#<R"K!SR'@FE3'"!#M!+#*'01"*%2&-,3"2&-4#5,*% 8%$B!0B$(.CJB./-#*'01A"&&!.(
"!#<R"K!&!"61=0O,SR'#-3+F5 99 ./-#*'01;3"X?.("!#<R"K!;81+'#-3+F5 1 AF52&B8%$B!&1"!#<R"K!SR'@FE3'#*'013N=( L 61=&1>B32&-,3"2&-4#5,*%.(4#5;6<26+;F+
M';!"# 3 #*'0161=./-.("!#<R"K!SR'@FE3'#*'0161=&1>B32&-,3"2&-4#5,*%.(4#5;6<26+
27
A%%;M1+%8F*(79%
2&B#5%O14%
A%%;#N73#*'7%
$)H1"!#M!+#*'01.C-A"B2&-,3"2&-4#5,*%;8N=3/*"(:!.C-;"),"!#"F!+8*(HO? 8%$B!0B$(.CJB./-$)H1"!#M!+#*'01A%%;M1+%8F*( T),;4Q(#-3+F5 79 #3'F'&!TN3 2&B#5%O$)H1"!#M!+#*'01 T),;4Q(#-3+F5 14 AF5"!#M!+#*'01A%%;#N73#*' T),;4Q(#-3+F5 7 >!&F:!,*% (U!861= 4)
M';!"# 4 $)H1"!#M!+#*'0161=./-.("!#<R"K!SR'@FE3'#*'0161=&1>B32&-,3"2&-4#5,*%.(4#5;6<26+
4#)&!9#*'0161=M!+.C-A"B2&-,3"2&-4#5,*% 8%$B! 4#)&!9#*'01A%%;M1+%8F*(61=./-3+GB.(/B$'>=:!0O,-0G'0O, TN3 1-600 ;"#+? P,+0B$(.CJB4#)&!961=()+&./-&!"61=0O, 53+GB.(/B$' 10-100
;"#+? 0B$("!#M!+#*'01;8N=34d3'"*(":! *,<*>#G2&-,3"2&-4#5,*%CF*'"!#;"e%;"1=+$ 8%$B!&1"!#<R"K!"*%"F-$+2&- 3+GB.(/B$' 1.5-2 ")PF;"#+? (>!#!'61= 4)
- $%02:(8:B"!"#3"GH253162&5317(-68
"!#./-#*'01"*%2&-,3"2&-4#5,*% 0B$(.CJB&1$*>SO4#50'T?;8N=3/*"(:!.C-;"),"!#"F!+8*(HO?.C-2,-F*"K95.C&B L 61=,1>!&>-3'"!# ;8N=34#5P+/(?.("!#4#*%4#O'8*(HO? XR='@F"#56%61=;"), !""!#./-#*'01"*%2&-,3"2&-4#5,*%(*7( 8%$B!2&-,3"2&-4#5,*%A>BF5/(),&1"!#>3%0(3'>B3#*'0161=TF-!+"*( A>B&1T$!&6(6!(>B34#)&!9#*'0161=M!+.C-A>">B!'"*( P,+@FE3'#*'0161=&1>B32&-,3"2&-4#5,*% &1,*'(17
1. @F>B3"!#;4F1=+(A4F'E3'PT#P&PX& ;/B( "!#A>"C*" C#N3E!,C!+24E3'PT#P&PX& "!#;8)=&C#N3F, :!($(PT#P&PX& ;4Q(>-(
2. @F>B3F*"K956!'0*9]!(E3'8N/ ;/B( F*"K95AT#5A"#( "!#;4F1=+(A4F'E3'F*"K95E3'.% #G4#B!'E3'.% E(!,E3'4!".% "!#;4F1=+(A4F'F*"K95#G4#B!'E3'F:!>-( fFf
3. @F>B3"!#;4F1=+(A4F'01,3"AF5.% 2,-A"B F*"K95,B!'E3'0B$(>B!' L
"!#;4F1=+(A4F'01,3" #G4#B!'E3',3" E(!,E3',3" ;4Q(>-(
4. @F>B3"!#; #)J;>)%P> XR='8%$B!#*'01&1@F6*7';#B'AF5+*%+*7'"!#; #)J;>)%P>E3'2&-,3"2&-4#5,*%
5. @F>B3;&Fe, 8%$B!#*'01&1@F>B3"!#'3"AF5;43#?;Xe(>?T$!&3+GB#3,E3';&Fe,
2&-,3"2&-4#5,*%
28
6. @F>B3#5%%"!#0N%8*(HO? 8%$B!#*'01&1@F6:!.C-2&-,3"2&-4#5,*%;"),"!#;4Q(C&*(
7. @F>B3$*g *"#/1$)>E3'2&-,3"2&-4#5,*% ;/B( #*'01&1@F>B3#5+5;$F!33",3"
#5+5;$F!"!#%!(E3',3" #5+5;$F!;"e%;"1=+$ fFf
8. @FE3'#*'01>B3F*"K956!'0#1#$)6+!CF*'"!#;"e%;"1=+$E3'2&-,3"2&-4#5,*%
8%$B!#*'01 56:!.C-2&-,3"2&-4#5,*% ;/B( "F-$+2&-&13!+OCF*'"!#;"e%;"1=+$0*7(F' A>B&1#!+'!(.(;%J &!< 8%$B!#*'01&1A($P(-&6:!.C-;%J &!<&13!+O"!#4I"A "*((!(ER7(
XR='"!#<R"K!@FE3'#*'0161=&1>B32&-,3"2&-4#5,*%.(4#5;6<26+(*7( 2,-#$%#$& !"@F'!("!#<R"K!AF5$) *+P,+0#O42$-.(>!#!'61= 4
29
!"!#$%& 4 !"#$%&$'()(*+
(,-#.+/0#1.+/2
%30&4
5670,-8$
9 :6
2%3;)
<.)=
>81%8=$8618%<?1@
8A7>&=
,&B$C,-2
D E.<
. 25
10-2
550
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
1!/A=4&A'('/+
#&L5M!(
(254
6)
2!8==#0)
(*;!(B=$
:6'N
8E2!#0;5FB#
C1++
8
5-30
- 2%7+8J%&$'()
H8:O/;17018%,8=
30 C!3 5
0 ;2#%K;PQ6,
K (LD
30 C!3
LD
50)
;)-81&4
25.5
C!3
28.
5 ;1%=K ,8+!H80&4
- +(18%C,1O
6-#!0!$;+F*#2%7+8J
%&$'(;E7*+
"?B6
- +(18%%#05(A7,!0!$;+F*#2%7+8J
%&$'(;E7*+
"?B6
- )H8:O/;17018% 7021
,7)8$'&J
R86A7)=
8C!3'8+8%SS-8=)
#0:O/1&4
O6-#
%M-6,-#.2.0/
- %3=3;A!818%##10#1 C!3%3=3;A
!818%486"#$0#1
C1++
8 C44;I(=4E
!&6
20-8
0
1!/A=;4E
#8%=8
(254
8)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++
8C44
;%FB#%&$
18.7
-77.
6
- 1!/A=;4E%#05(A7,
50 ;2
#%K;PQ6,
K (LD
50) )(*2
%7+8J
%&$'(
33.5
;1%=K
- ;6FB#;=F*#1!/A=;4E )(*.0/%&4%&$'(,-#;6F*#$
18.
7 ;1%=K E4A-848$,/6
+(TA8+
7021
,7:4+
(!&1@J
3470$#+/A6;"/8O81&6 C,-'-A6:OL
-+(!&1@
J3T!/8=T!?$
1&61&4
,/6)(*.+-.0/I
8=%&$'(
T%%57,
(253
8)
U2%U,T#%K+>81
18%;E
83!(B=$
;6FB#;=F*#
C1++
8 20
-130
- 2%7+8J%&$'( %3OA-8$
80 C!3 1
30 ;1
%=K ;O
+83'+:61
8%5&16H8:O/;17018%
1!8=E&6
VMK - %&$'()
H8:O/1
8%;>%7L
;,74U,"#$U2%U,T#%K+1!/A=.+/5/8!$ 4
8$'-A6
;2!(*=6;2W6
'(6BH8,8!C!3,8=
!>3%M6C%$"?B6
;+F*#:5/2
%7+8J
%&$'()
(*'G$"?B6
1!/A=.+/
#%6M5
(254
5)
0#1
C1++
8C44
;I(=4
E!&6
500-
2000
- 2%7+8J%&$'(
1.5-
2.0
kGy '8+8%S1H8>&0;E!(B=.X.0/)M1
%3=318%
;>%7L
;,74U, C,-'-$ !,-#18%!0#8=M18%2Y1C>1&6"#$1!/A=.+/ 2
-7 A&6
30
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
V&6AK
(254
6)
U2%U,T#%K+)
(*.0/
>8118%;E83;+!Q0
C1++8C44
;I(=4
E!&6C!3
C44;%FB#%&$
20-8
0 - ,/6;#FB#$TH8 +
(6BH8O6
&1 TA8+'G$!H8C!3TA8+=8A:4!0!$
,8+#&,%8%&$'(
)(*.0/%&4;E7*+
"?B6
- %30&4
%&$'(
60 ;1
%=K )H8:O/A-86;E5%O?$
+("680"#$,/6)
(*:OL-1A-8,/6)
(*.+-.0/%&4%&$'(C!3%#0,8=)&B$O+
0 - ;+!Q05/8$T-#+)(*
-8618%I8=%&$'(C44
;%FB#%&$
#&,%8 6
27.5
+(18%E& Z6
8;2W6
,/61!/8"680:OL
- T70;2W6
55.
93 ;2
#%K;PQ6,
K 'M6(=K
(254
7)
0#1
C1++
8 15
00
- +(
!)H8:O/;17018%O!M0%-A$"#$0#1#=-8$%A0;%QA
1!/A=.+/ (,
-#)
V63 (
2548
)
U2%U,T#%K+
C1++
8C44
;I(=4
E!&6C!3
C44;%FB#%&$
2-10
0
- 2%7+8J%&$'()
H8:O/1
8%;>%7L
;,74U,!0!$ 3
0 ;2#%K;PQ6,
K (GR
30) ;+F*#;)(=
41&4
18%.+-.0/%&4%&$'("#$ D
en. W
hite
San
an,
Den
. Pin
ky S
em 'R
inna
pa',
Den
. Son
ia'N
o.17
Red
', D
en. Son
ia 'E
arsa
kul' TF#
73.
93 8
1.56
64.
00
C!3 6
9.05
;1%=K ,8+!H80&4
- 18%I8=%&$'(2
%7+8J
,*H8 2
-10 ;1%=K 1&4
PBLs
'8+8%SE
&Z68;2
W6,/6.0/
N8=:6
10 ;0
F#6 %&$
'(+( !:O/TA8+'G$!H8
TA8+'G$)%$,/6
C!3TA8+=8A:4
!0!$
- 18%I8=%&$'(C44
;I(=4
E!&62
%7+8J
20-
100 ;1%=K)H8:O/ TA8+'G$!H8 T
A8+
'G$)%$,/6 TA8+=8A:4 C!3TA8+=8A%81!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- 18%I8=%&$'(C44
;%FB#%&$
,/6)
(*.0/%&4
800
;1%=K +(18%;>
%7L;,74U,)
(*5/81A-8
C!3E4!&1@
J3 702
1,7
31
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
1!/A=%&,1&)!(
A6708
(254
7)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++8
C44
;I(=4
E!&6
10-7
0
- 2%7+8J%&$'()
H8:O/;17018%,8=
50 ;2#%K;PQ6,
K (LD
50) ;)-81&4
27 ;1%=K
;+F*#;E7*+
2%7+8J%&$'(>3)H8:O/+
(18%%#05(A7,C!318%C,1O6
-#!0!$
- ;+F*#=/8=2!G1E
4A-8 ,/6
)(*I8=%&$'(+("6
80C!3TA8+'G$6/#=1A-8,/6)
(*.+-.0/
I8=%&$'(
- 1-#:O/;17018%;2!(*=6C2!$0/86'&J
R86A7)=
848$2%318%
;5-6 '( 18%;%
(=$,&A
"680 %G2%-8$"#$:42%30&4;2!(*=6
.2
- ;170#
818% 70
21, 7)
(*:4 ;5
-6 ;1700
-8$"8A :4O6
8 :4O=
&1 C!3+(!&1@J
3 70
21,7"#$'(:42
%30&4
;2!(*=6C2!
$.2#
=-8$;0
-65&0>H86A6
2 ,/6
U0=,/6
C%1 +(1
8%;%(=$"#
$:42
%30&4
5701&6
C6-6 +('
(;O!F#$#-#6
#+'/+
;!Q16
/#=
"#4+
('(;"(=A C!32!8=+(%#=O
=&1;2W6T
!F *6 1!(448$ ,
/6)(* 2
:42%30&4'
( ;O!F#$#+;"(=AS&0;"/8.20/86:6+
('(;O!F#$#+'/+
"#4
'(;"(=A 2!8='M0+(C,/+
'(;"(=A 1!(4O
68C!3T-#6"/8$C"Q$
1MO!84
E&5%76)%K
(253
8)
=#0=8A
10
+7!!7;+,
% )(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8 C44
;I(=4
E!&6
20-4
0 - ;2#%K;PQ6,
K18%#=G-%#0!0!$;+F*#2%7+8J
%&$'('G$"?B6
- '(0#1;2!(*=6C2!$.2
- %G2%-8$0#1;2!(*=6C2!$.2
- >H86A6A&6)
(*:5/:618%##10#1;2!(*=6
C2!$.2
32
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
'M%A75
(252
6)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++
8C44
;I(=4
E!&6
5-50
- 18%;>
%7L;,74U,!0!$ ;+F*#2%7+8J%&$'(;E7*+
"?B6
- 2%7+8J%&$'(
5 ;1%=K )H8:O/;170=#0+81"?B6
- )H8:O/!&1@J
318%;%(=
$,&A"#$:4
TA8+=8A2!/#$
C!318%C,117*$"/8$
7021
,7 - 2%7+8J%&$'(
35 ;1
%=K )H8:O/;170!&1@J
3 702
1,7+81)
(*'M0
O+8=;O,M TA8+ 7021
,7"#$!&1@J
3:4C!318%C,117*$"/8$'8+8%S;1701&4
)-#6E
&6VMK)(*;!(B=$:6'N8E2
!#0;5FB#U0=.+-,
/#$I8=%&$'(.0/
A&6;I!7+
(254
6)
17*$C!3:4
C1++
8C44
;I(=4
E!&6
10-6
0
- ;2#%K;PQ6,
KTA8+#=G-%#0:6:4C!317*$!0!$;+F*#;E7*+
2%7+8J%&$'(+81"?B6
- ;+F*#;E7*+
2%7+8J%&$'(>3)H8:O/1
8%C,117*$:O+
-C!3>H86A6:4>8117*$:O+
-!0!$
- !&1@J
3 702
1,7)(*E
4TF# :4+("680;!Q1;2W6E
M-+ 17*$+(!&1@
J3 702
1,7 E4
)(*2%7+8J
%&$'(
20, 40
C!3
60 ;1%=K U0
=TA8+
7021,7>3;E7*+"?B6,
8+2%7+8J%&$'()
(*;E7*+"?B6
1MO!84O
76
>7%36&6)
K (2
547)
,/6#-#6
C1++
8
20-4
00
- )H8:O/;170!&1@
J3 702
1,7 .0
/C1- :4
470;4(B=A
+(!&1@J
3;2W6%G2O
&A:> :4
C[0,701&6 :4;%(=
A
- %&$'(
10 ;1
%=K )H8:O/;170
pla
ntle
t 46
:4.0/;%QA
- 2%7+8J%&$'('G$1A-8
200
;1%=K +( !,-#18%!018%;>%7L
;,74U,C!3TA8+#=G-
%#0"#$,/6
33
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
#%A%%J
(252
4)
O&A
C1++
8C44
;I(=4
E!&6
10-5
0
- 18%#84%&$'()
(* 20 ;1%=K 686
7 A&6 +(
!)H8:O/;E7*+
TA8+=8A"#$5-#0#1
"6800#1 >H86A60
#1,-#5-#
)H8:O/#
#10#1;%QA"?B6
;17018% 7021
,7"#$5-#
0#1C=1##1;2W6
2 5-#
- 18%#84%&$'()
(* 10 ;1%=K 686
7 A&6 +(
!)H8:O/ ;E7*+18%;>%7L
;,74U,"#$
,/6 :O
/>H86A6O
&AC!3O&A=-#=;E7*+
+81"? B6
- 18%#84%&$'()
(* 50 ;1%=K 686
7 A&6 )H8:O/18%;>
%7L;,74U,!0!$ :4
7021
,7 C!3.+-##10#1
C1!07U#!&'
#%A%%J
(252
5)
O&A)(*;!(B=$.A/:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
20-1
00
- %&$'(
20 ;1
%=K >3;E
7*+>H86A6O&A
>H86A60#1,-#5-# "6800#1 %3=3;A!8
18%486"#$0#1
C!3%-6#8=M18%,&00#1"#$C1!07U#!&'
- 1-#:O/;17018% 7021
,7"#$1/860
#1
- %&$'(+811A-8 6
0 ;1%=K >3.+-1
-#:O/;1705-#0#1
- %&$'(
40 ;1
%=K >3'-$ !=&4=
& B$18%;>%7L
;,74U, )H8:O/,/6
CT%3C1%6
:4O6
8"?B6
34
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
C1!07U#!&'
(,-#
)
2%86#+
C!3TJ3
(252
5)
O&A)(*;!(B=$.A/:6
'N8E2!#0;5FB#
C1++
8
20-1
00
- %&$'(
20 ;1
%=K >3;E
7*+>H86A6O&A
>H86A60#1,-#5-# "6800#1 %3=3;A!8
18%486"#$0#1
C!3%-6#8=M18%,&00#1"#$C107U#!&'
- )H8:O/;17018% 7021
,7"#$1/865
-#0#1C=1;2W6
2 5-# :6
vm
1 C!3.+-E
4:6
vm
2
- )&B$
2 E&6VMK )(*#84%&$'(+811A-8
60 ;1%=K >3.+-;1705-#0#1
- %&$'(
40 ;1%=K )H8:O/1
8%;>%7L
;,74U,"#$,/6
!0!$
,/6CT%3C1%6 :4+(
TA8+O6
8;E7*+"?B6 TA8+1A/8$
TA8+=8A
C!3>H86A6:4 !0!$:6
vm
1 C,-
vm2
vm3
vm4 +(18%;>%7L
;,74U,;2W6
21,7
;4L>
+8<
(254
1)
;6FB#;=F*#)(*;!(B=$.A/:6
'N8E2!#0;5FB#
C1++
8 C44;I(=4E
!&6
10-1
10
- O6
-#"7$C0$+(T
-8 LD
50 ;)
-81&4
22 ;1
%=K
- 2%7+8J%&$'(
50 ;1
%=K "?B6.2 )H8:O/"7$C0$O=M018%;>
%7L;,74U,
- 2%7+8J%&$'(
30 ;1
%=K )H8:O/1
8%;>%7L
;,74U,!0!$ C
!3+(!&1@
J3 702
1,7
TF# +(18%##1O6
-#0/86"/8$;2
W6>H86A6+81C,-##1%816
/#=
- 2%7+8J%&$'(
10 ;1
%=K )H8:O/:4+
('(;"(=A#-#6
C!3+(%7BA'("8A;!Q1
9 2%81\
#=G-46
:4
"7$C0$
;1<
(254
4)
,/6)(*;E83;!
(B=$:6
'N8E2!#0;5FB#
C1++
8 C44;I(=4E
!&6
5-60
- T-8
LD
50(3
0) ;)
-81&4
52.2
;1%=K
- ,/6
)(*I8=%&$'(
60 ;1
%=K ,8=O+
0 :6%M-6
M1V
2
- ,/6
)(*%#05(A7,>8118%I8=%&$'(
15 ;1
%=K +(18%C,1O6
-#;2W6
1!M-+1
/#6 :4
O$71$# :4;2W6CI1
C,-!&1@
J36(BO8=.2:6%M-6
M1V
3
- 18%I8=%&$'(
5 ;1%=K E4!&1@
J3:4;2W6
%7BA'("8A:6;6FB#;=F*#
:6%M-6
M1V
3
)&B$)(*%M-61
-#6O6
/8.+-2
%81\
!&1@J
30&$1!-8A##1+8
35
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
C1++
8C44
;I(=4
E!&6
10-1
50
5&=5M+E!
(252
6)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C44;%FB#%&$
I8=,-#;6F*#$)M1
TF6 ,!#0
7 TF6
%A+
308.
32 ;1
%=K
- ;2#%K;PQ6,
K18%#=G-%#0C!3C!318%;>
%7L;,74U,!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- '("#$0#1;2!(*=6
C2!$.2
T8%K;6
5&*6
#%0( C!35&=
5M+E!
(252
7)
=#0)
(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6C!3
C44;%FB#%&$
10-1
50
- ;2#%K;PQ6,
K18%#=G-%#0C!318%;>%7L
;,74U,!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- 2%7+8J%&$'( ,&B$C,- 7
0 ;1%=K"?B6
.2 O!&$>81I8=%&$'(
12 '&20
8OK )H8:O/,
/6,8=O
+0
- '("#$0#1;2!(*=6
C2!$.2>81'("8A;2W6
0#1'("8A)(*+("(05+E
G1%3>8=)&*A
0#1
P-#61
!7*6.)=
#07<%
(253
5)
O6-#
C1++
8C44
;I(=4
E!&6
5-30
- 18%;>
%7L;,74U,SG1=&4=
&B$,8+2%7+8J%&$'()
(*;E7*+"?B6
- >H86A6O
6-#!0!$,8+2
%7+8J
%&$'()
(*;E7*+"?B6
- 1-#:O/;170!&1@
J3:40
-8$
36
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
0#$0?$
E%)7E
=K (2
542)
O&A
C1++
8 2.
5-40
0 - 2%7+8J%&$'()
(*;E7*+"?B6)
H8:O/#
&,%818%%#05(A7,C!318%;>%7L
;,74U,!0!$
U0=+(T-8
LD
50 ;)
-81&4
10.8
8-14
.22
;1%=K
- 2%7+8J%&$'(
20 ;1
%=K "?B6.2+
( !)H8:O/0#$0?$,8=
100
;2#%K;PQ6,
K - 2%7+8J%&$'(
2.5 ;1%=K +( !,-#!&1@J
3)8$'&J
R86A7)=
8C!318%
;>%7L
;,74U, TF#
>H86A6,/6
)(*$#1##1>81,84%7;AJ2!8=O&A+(;E7*+
"?B6
TA8+'G$"#$,/6!0!$ '
("#$:4 !&1@J
3"#$0#1C!3'("#$0#1;2!(*=6
C2!$
.2 TA8+'+4G%JKC!3;2
#%K;PQ6,
K18%$#1"#$;%
JG!0!$
%A+)
&B$+(O&A)(*;170
:O+-;E7*+
"?B6
- +(
!,-#UT%U+UP+ TF#UT%U+UP+
>&4TG- 702
1,7 ;1
70;2W6
rin
g m
ultiv
alen
t
C!3E4
lagg
ing
chro
mos
ome C!3 c
hrom
osom
e br
idge
U0=>H86A6
;P!!K)(*E
4TA8+
7021,7"#$UT%U+UP+;O!-86(B >3+81"?B6
,8+2
%7+8J
%&$'(
C1++
8)(*;E7*+"?B6
08A1%3>8=
V68V72
(254
7)
,/6#-#6)
(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
20-1
00
- 18%;>
%7L;,74U,!0!$;+
F*#2%7+8J
%&$'('G$"?B6 U0=+(T-8
GR
50(3
0) ;)-81&4
92.1
9 ;1%=K
- 18%%#05(A7,!0!$;+
F*#2%7+8J
%&$'('G$"?B6
- 18%;2
!(*=6C2!$"#$:4
.0/C1- :4
;O!F#$ :4+
(!&1@J
3470$# 70%G2 :4;%(=
A=8A C!3:4:OL
- 7021
,7
37
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
A7)=81M!
(254
3)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++
8C44
;I(=4
E!&6
10-8
0
- T-8
LD
50(3
0) ;)
-81&4
27 ;1
%=K C!3T-8
GR
50(3
0) ;)
-81&4
12 ;1
%=K
- ;6FB#;=F*#)(*I
8=%&$'(:62
%7+8J
20 ;1%=K"?B6
.2 ;1
7018%,8=)
&B$O+0
- ;6FB#;=F*#08O!8)(*%#05(A7,
)(*2%7+8J%&$'(
10 ;1
%=K E4!&1@
J3)(*
;2!(*=6C2!$TF# :4;2W6%#=>(4
:4;%(=
A;!Q1
:40-8$ :4CI1
C!3!&1@J
318%
C,1O
6-#;2W61
!M-+1/#6
- !&1@J
3:4;2W6
%#=>(4 C!3:4CI1
O8=.2 :6%M-6
M1V
3
#N758,7 C!3
TJ3 (
2543
)
;6FB#;=F*#)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8 .+-%34M
- )H8:O/;6FB#;=F*#,8=;+F*#;!(B=$.0/ 3
0 A&6
O6-#
C1++
8C44
;I(=4
E!&6
1-3 ;1%=K C!3
I8=%&$'(;E7*+#(1:6
%M-6 M
1V2 )%()
;+/6,
K!3 1
5 ;1%=K
- 2%7+8J%&$'('G$"?B61
8%,8='G$"?B6
- 18%C,1O6
-#:O+
- (M1V
2) !0!$
- !&1@J
3 702
1,7)(*E
4 TF#
+(18%C,1O6
-###1+8+81>65
7B6;6FB#;=F*#;2W6
1!M-+1
/#6 ,/6CT%3C1%6 :4+(!&1@
J3 702
1,7 .0
/C1- :4
0-8$ :4;%(=A=8A
:4%G2O&A:>
:4O=
71 C!3 :4CI1
>7%8N%J
K (2
545)
;O$/8
C1++
8C44
;%FB#%&$
30-1
20
- 2%7+8J%&$'( ,&B$C,- 6
0 ;1%=K )H8:O/,
/6,8=O+
0 - ,/6
)(*.0/%&4%&$'( 3
0 ;1%=K +(TA8+'G$C!3C,1O6
-#+811A-8,/6)
(*.+-.0/I8=
%&$'(
08O!8
A%E6
7, (2
545)
,/6)(*
-8618%I8=
%&$'(
10 ;1
%=K +8
;E83;!(B=$;6FB#;=F*#
C1++
8C44
;I(=4
E!&6
10-4
0 - 18%,8=;E7*+"?B6,
8+2%7+8J%&$'()
(*;E7*+"?B6
- 18%C,1O6
-#!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- !&1@J
3 702
1,7)(*E
4 TF#
!H8,/6CT%3 :4O
$71$#
:4O-#
:4+("680;!Q1!$
18%C,1O6
-#;2W6
1!M-+1
/#6
38
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
08O!8 (,-#
) 21
%JK
(254
8)
;6FB#;=F*#)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
10-4
0 - T-8
LD
50(3
0) ;)
-81&4
20.4
4 ;1%=K C!3T-8
GR
50(3
0) ;)
-81&4
16.6
5 ;1%=K
- E4
!&1@J
3 702
1,7)(*2
%7+8J
%&$'(
10 ;1
%=K TF# :4;%(=
A;!Q1
:4;2W6%#=>(4
:4O$71$# :4+(%7BA'(;O!F#$ :4;2W6
CI1 C!3:4O
-# !H8,/6
CT%3C1%6
!H8,/6+
("680;!Q1 C!3+(18%C,1O6
-#;2W6
1%3>M1;+F*#;)(=4
1&4,/6
)(*.+-.0/%&4%&$'(
)86,
3A&6
<7%7%&,6K
(254
6)
;+!Q0
C1++
8C44
;I(=4
E!&6
50-2
50
- ;2#%K;PQ6,
K18%$#1!0!$
- ;2#%K;PQ6,
K18%#=G-%#05(A7,!0!$
- TA8+'G$"#$,/6!0!$
- >H86A6:4;+F*#;170,80#1C%1!0!$
C1++
8C44
;I(=4
E!&6
20-6
0 .)%=/#=:4CO!+
0-8$
'M5808
(255
0)
17*$2Y1
5H8
C1++
8C44
;%FB#%&$
19.2
-515
.3
- 18%I8=%&$'()
&B$'#$C44+
( !)H8:O/18%;>
%7L;,74U,"#$.)%=/#=:4CO!+
0-8$!0!$ ;+F*#.0/%&42
%7+8J
%&$'()
(*'G$"?B6
- 18%I8=%&$'(C1++8C44;%FB#%&$
+( !)H8:O/;17018%1!8="#$:4O
!8=
!&1@J
3 U0=;17018%;2!(*=6C2!$%G2C44
18%0-8$"#$:4
%G2%-8$:4
'("#$
:4 C!3"680"#$:4
U0=+(!&1@J
318%1!8=)(*T
$)(* 2
!&1@
J3 O
!&$18%
"=8=E&6
VMK#=-8$,-#;6F*#$
4&A>$1!6(
Pur
ipun
yava
-nic
h an
d
Boo
nsiri
chai
(200
5)
O&A=-#=
C1++
8 50
-200
- T-8
LD
30 ;)
-81&4
70 ;1
%=K C!3
LD
50 ;)
-81&4
120 ;1%=K
- 18%;>
%7L;,74U,5/8C!3,/6
#-#6C#1A-8,/6
)(*.+-.0/I
8=%&$'(
- #&,%818%$#1;2W6,
/6:O+
-!0!$,8+2
%7+8J
%&$'()
(*;E7*+"?B6
- !&1@J
3'(1!(40
#1;2!(*=6.2
2 !&1@J
3 TF# 1!(40
#1'("8AC!31!(40#1
'(+-A$
39
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
;#+#%
(253
0)
,/6#-#6)
(*;!(B=$
N8=:6"A0
;E83;!(B=$
C1++8
- - 2%7+8J%&$'(
30 ;1
%=K +(18%;>
%7L;,74U,0(1A-8%30&4%&$'(#F*6
9
- ,/6
)(*.0/%&4%&$'('G$1A-8 5
0 ;1%=K >3;>
%7L;,74U,.+-0
(C!3,8=:6)(*'
M0 - 18%;>
%7L;,74U, %A+S
?$!&1@
J3N8=6#1)
(*'&$;1, .0/C1- %G2
%-8$ '
( C!3
"680"#$:4 0#1C!3)%$,/6 ;2W62
1,7
C1++
8C44
;I(=4
E!&6
20-6
0
4&AO!A$
<7%7<&107]
(254
2)
,/6)(*.0/>8118%2Y1
5H8.A/:6'N
8E2!#0;5FB#
;#1P
K 10
-50
- T-8
LD
50 "#$4&AO!A$
)(*.0/%&418%I8=%&$'(C1++8C!3;#
1PK TF# )(*2
%7+8J
%&$'(
20
;1%=K U0
=%&$'(C1++
-8C!3;#1P
K)H8:O/;170!&1@
J3:O+
- 21 !&1@J
3 - ,/6
)(*.0/%&4%&$'(C1++
-8C!3;#Q1)
(* 10-
20 ;1
%=K +(!&1@J
318%;>%7L
;,74U,)
(*0(1A-8,/6
21,7
+(%81C"6$=8A :OL- C!3%81[#=>H86A6+81 .O!+(TA8+'+4G%JK C!3:O/,/6
:O+-)
(*+811A-8
- ,/6
)(*.0/%&4%&$'(C1++
-8C!3;#Q1)
(* 30-
50 ;1
%=K +(!&1@J
3 702
1,7)(*.+-'-$;'
%7+18%
;>%7L
;,74U, ;5-6 ;2W6C1/A
(vi
trifi
catio
n) P(0;O!F#$)&B$,/6
1/86:4+/A6E
&61&6;2W6
1/#6
.+-C,1,8"/8$
.+-+(%81C!3.+-C,11#
- ,/6
)(*.0/%&4%&$'( 6
0 ;1%=K >3,8=O+
0N8=:6
4 '&20
8OK
- 4&AO!A$+(UT%U+UP+)
(*2!8=%81
2n=16
TA8+=8A281:4;I!(*=
2.46
.+T%#6
%&$'()
H8:O/,/6
)(*;17018%1!8=E
&6VMK)(*+
(UT%U+UP+
ane
uplo
id 2
1!M-+
TF# 1!M-+)
(* 1 +(>H86A6
UT%U+UP+
2n
= 1
8 +(TA8+=8A281:4;I!(*=
3.43
>H86A6
5 ,/6 U0=;170>8118%I8=
%&$'(C1++-8C!3;#
1PK)(*
30 ;1%=K C!3%&$'(;#1PK)
(* 40 ;1%=K C!31!M-+)
(* 2 +(>H86A6
UT%U+UP+
2n
= 2
0 +(TA8+=8A281:4;I!(*=
4.3
4 >H86A6
3 ,/6
U0=;170>8118%
I8=%&$'(C1++
-8)(* 4
0 ;1%=K
- ,/6
)(*;170>8118%I8=%&$'(C1++
-8)(* 4
0 ;1%=K +(281:4
7021,7 ;2
W6C44
cyc
locy
tic
>H86A6
1 ,/6
C!3281:44
70;4(B=
A >H86A6
1 ,/6
40
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
<7%7<&107]C!3
A75&=
(254
3)
,/6)(*;!(B=$:6
1%3S8$2!G1
C1++
8
20-6
0
- T-8
LD
50 #=G-%3OA-8$ 0
-20 ;1%=K
- '8+8%S5&16H8:O/+(!&1@
J3 702
1,7
- )H8:O/+("680"#$,/6
;!Q11A-8,/6
)(*.+-.0/%&41
8%I8=%&$'(
- 18%I8=%&$'(+811A-8 4
0 ;1%=K )H8:O/4
&A,8=)&B$O+
0N8=:6
2 '&20
8OK
4&AO!A$
(,-#
)
Lam
a et
al.
(200
5)
;+!Q0
C1++
8 20
-100
- %&$'(
60 ;1
%=K +( !!018%;>
%7L;,74U,"#$,/6
1!/8!$ 5
0 ;2#%K;PQ6,
K - %&$'(
40 ;1
%=K ,/64
&AO!A$.+-'8+%S;170%81.0/ C!3E4
!&1@J
3 702
1,7
;5-6 4A+I
*H86BH8 =
#0C!3:4'(;O!F#$ :4C!31/86:4+
('(C0$
4&AO!A$#;+%71&6
'(;O!F#$
A75&= C!3
161E
% (2
549)
;O$/8
C1++
8C44
;I(=4
E!&6
10-3
0
- #&,%818%%#05(A7,!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6 U0=+(T-8
LD
50 2%3+8J
17 ;1
%=K
- U0=2
1,74&A'8=E
&6VMK6(B.+-'8+8%S##10#1:62
%3;)<.)=
C,-E
4;E(=
$ 1
,/6 '8+8%S##10#1:62
%3;)<.)=
.0/
- 2!8=1!(40
#1UT/$+6+
811A-8E&6VMK;07+)
(*+(2!8=1!(40#1CO!+
4865
F*6O6G
E757,
(254
6)
;+!Q0
C1++
8C44
;I(=4
E!&6
100-
600
- +(
!,-#;2#%K;P
Q6,KTA8+$#1 U0=TA8+$#1>3!0!$;+F*#2%7+8J
%&$'(
;E7*+"?B6 2%7+8J%&$'(
500 ;1%=K )H8:O/;2#%K;P
Q6,K18%$#1!0!$
50
;2#%K;PQ6,
K - 18%;>
%7L;,74U,!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
4(U1;6(=
'M+68
(252
8)
,/6)(*;E83>81;+!Q0
;!(B=$:6'N
8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
20-1
00
- ;17018%;2!(*=6C2!$"#$%G2)%$,/6 :4C!3 0
#1;2!(*=6C2!$.2>81;07+
.0/C1- ,/6;,(B=CT%3
:40-8$ :4O
68 7021
,7 ;P!!K281:4:OL
-"?B6 '(0#1
;2!(*=6C2!$.2.0/O
!81O
!8=+81
"680"#$0#1+
()&B$:OL-"
?B6C!3;!Q1!$
1A-8;0
7+
41
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
4(U1;6(=
(,-#
)
6$!&1@J
K (2
541)
'-A6=
#0)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
20-1
00
- 2%7+8J%&$'()
(* 40 ;1%=K "?B6.2 )H8:O/;17018%,8=
100 ;2#%K;PQ6,
K - C -6:4C!31/86:4>81,/6
)(* -8618%I8=%&$'(+(18%;2
!(*=6C2!$C!3E&Z
68.2;2W6=
#0.0/6/#=1A-8,/6
)(*.+-
-8618%I8=%&$'(
- !&1@J
3 702
1,7,%A>E4
>81,/6)
(*.0/%&4%&$'(2
%7+8J
20 ;1%=K ;5
-6 :41!+
:4;% (=
A=8A
:4'(C0$;"/+C!3+(2
%3'("8A+811A-821
,7 18%;1
70=#0;2W6
1%3>M1
5M,76)
% (2
531)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++
8C44
;I(=4
E!&6
5 C!3 1
0
- 18%;>
%7L;,74U!0!$,8+2
%7+8J
%&$'()
(*;E7*+"?B6
- ;170#818% 7021
,7 ;5-6 "/#S(* :4;O!F#$
- !&1@J
3"#$0#1 .0/C1- ;'/6
-8<G6=
K1!8$1/860
#1 R86%#$0#1
0#1
5&B6:6
>H86A60#15&B66
#1C!35&B6:6 ;E7*++
81"?B6
- #8=M18%2Y1
C>1&6
686"
?B6 - 18%;2
!(*=6C2!$'(0#1 ;5-6 >810#1'("8A;2W 6'(5+EG,
'(+-A$;2
W6+-A$#+
6BH8,8! "8A ;2W6 '(5+E
G ;O!F#$ O
%F# 5+E
G'!&4%7BA'(6BH8,8! ;2W6,
/6
;4L>
+8<
5M,76)
% (2
532)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++
8C44
;I(=4
E!&6
10-1
00
- 2%7+8J%&$'()
(*;O+83'+:61
8%5&16H8:O/;17018%1!8=TF#
10 ;1%=K
- 18%;>
%7L;,74U,!0!$
- 1-#:O/;170!&1@
J3 702
1,7 ;5
-6 "/#S(*'&B6 :40-8$ (
Chi
mer
a)
- ,/6
,8=.0/O
81.0/%&4%&$'(+81;176.2
- 18%;2
!(*=6C2!$'(0#1
- +(
!,-#>H86A6UT%U+UP+
E4;E(=
$ 1 :6
4 ,/6
)(*I8=%&$'(
>H86A6
UT%U+UP+
;)-81&4
57 UT
%U+UP+
21,7;)-81&4
54
42
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
+&!!718
(254
2)
1!(40
#1)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
10-9
0 - 2%7+8J%&$'()
(*;E7*+"?B6)
H8:O/1
8%;>%7L
;,74U,!0!$
- 2%7+8J%&$'(
30 ;1
%=K "?B6.2)
H8:O/:4;O!F#$T!BH8 >
6;2W6
'(6BH8,8!
C!3,8=
:6)(*'
M0 - +(T
-8 LD
50 )(*%3=3;A!8
30 A&6
;)-81&4
24 ;1
%=K
- ;+F*#6H8##12!G1 E4
A-8,/6
)(*.+-.0/I
8=%&$'(+(18%;>
%7L;,74U,0( C!318%
C,117*$+811A-8,/6)
(*I8=%&$'(
- 2%7+8J%&$'(
10 C!3
20 ;1%=K )H8:O/:4O
$71$# 70%G2
:4O6
8 "680;!Q1
1A-821,7
;4L>
+8<
(,-#)
Lam
seej
an
et a
l.
(200
0)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++
8C44
;I(=4
E!&6
10-1
10
- +(T
-8 LD
50 )(*%3=3;A!8
30 A&6
;)-81&4
14 ;1
%=K
- %30&4
%&$'( ,&B$C,- 5
0 ;1%=K "?B6.2 )H8:O/'-A6=
#0)(*;!(B=$:6'N8E2
!#0;5FB#
%M-6 M
1V4 ,8=O
+0 N8=:6
25-
30 A&6
- 1-#:O/;17018%;2!(*=6C2!$"#$0#1 .0/C1- '( "6800#1
>H86A65&B6
C!3
%G2%-8$"#$0#1
43
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
;T%F#A&!=K
(254
1)
,/6)(*.0/>8118%
;E83;!(B=$;6FB#;=F*#
C1++
85670
;I(=4
E!&6
10-1
10
- 2%7+8J%&$'(
30 ;1
%=K )H8:O/1
8%;>%7L
;,74U,C!3>H86A6O
6-#!0!$
- 2%7+8J%&$'(
30, 70
, 90
C!3
110
;1%=K )H8:O/;170!&1@J
3 702
1,7'G$
;5-6
:4;O!F#$ :4.+-+
('( :45BH8
:4O6
8 C"Q$
1%#4 .+-;%(=
4
A&6A7'8
(254
5)
,/6)(*.0/>8118%
;E83;!(B=$;6FB#;=F*#
C1++
85670
;I(=4
E!&6
30-1
00
- T-8
LD
50 "#$E&6
VMK0#=,M$C!3E&6VMKEFB6
;+F#$
+(2%7+8J;)-81&4
37.5
1 C!3
37.5
6 ;1%=K ,8+!H80&4
- +(18%;>%7L
;,74U,!0!$ U0=T-8
GR
50 "#$E&6
VMK0#=,M$C!3E&6VMKEFB6
;+F#$
+(2%7+8J
;)-81&4
23.5
4 C!3 3
7.56
;1%=K ,8+!H80&4
- ;170!&1@
J3 702
1,7 TF# :4;O!F#$ P
(0
- ,/6
)(*.0/%&4%&$'(+811A-8
60 ;1%=K .+
-E4A-8+(1
8%;>%7L
;,74U,)
&B$ 2 E&6VMK
20-1
00
- %30&4
%&$'(,&B$C,-
40 ;1
%=K "?B6.2 '-$ !:O/18%$#1"#$;O
$/8SG1=&4
=&B$
- %30&4
%&$'(
20 ;1
%=K ;1
70 le
af c
him
era
#07<%
(253
3)
;O$/8
C1++
85670
;I(=4
E!&6
5-20
- ;O$/8$#15/81A-8;O$/8)(*.+-.0/%&4%&$'(
- #&,%818%;>
%7L;,74U,5/81A-8,/6
)(*.+-.0/%&4%&$'(
- ;O$/8)(*.0/%&4%&$'(>3;1700#15/8
C!35/8+81"?B6,
8+2%7+8J%&$'()
(*+81"?B6
- %3
0&4%&$'(
20 ;1
%=K E4
1 ,/6
)(*+('(5+EG"
#$184%#$0#1;"/+"?B6
C,-,/6)
(*;>%7L
,-#+8:O/0#1)
(*+('("#$184%#$0#1;5- 6;07+
2)M++
8
A75L8
(254
6)
,/6)(*.0/>8118%
;E83;!(B=$;6FB#;=F*#
C1++
85670
;I(=4
E!&6
10-1
10
- T-8
LD
50 +(T-8
42.
01 ;1
%=K C!3T-8
GR
50 +(T-8
42.
01 ;1
%=K )(*%3=3;A!8
45 A&6 U0=18%;>%7L
;,74U,!0!$;+
F*#2%7+8J
%&$'(;E7*+
"?B6
- 2%7+8J%&$'(
30 C!3
50 ;1%=K E4!&1@
J3:4;O!F#$
:4+('("8A)8$0/864
6"#$:4 C!3,8=O+
0 O!&$>81C=1,/6)
(*;170:O+
-!$;!(B=$:6#8O8%'G,%;07+
44
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
E%%J
.+/6BH8:4E8=
'M>7,%8
(254
8)
,/6#-#6)
(*.0/>81
18%;E
83;!(B=$
;6FB#;=F*#
C1++
8 C44
;I(=4
E!&6
10-5
0
- T-8
LD
30-L
D50
TF#
18.2
-22.
2 ;1%=K
- ;2#%K;PQ6,
K18%%#05(A7,!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- %30&4
%&$'(
10 C!3
20 ;1%=K )H8:O/!&1@
J3,-8$9"
#$:4;2!(*=6.2 .0/C1- '(
:4 %G2
:4 "#4:4
2!8=:4
- %30&4
%&$'(
10 C!3
20 ;1%=K )H8:O/,
/6+(!&1@J
3CT%3C1%6C!3C,1,/6
:O+->H86A6+81
- %30&4
%&$'(
40 ;1
%=K )H8:O/,
/6,8=O+
0 N&)
%+8<
(254
8)
17*$E&6
VMK)(*I8=%&$'(
1-#66
H8.22
Y15H8
C!317*$)(*2Y1
5H81-#6
6H8.2I8=%&$'(
C1++
8
20-1
00
- T-8
LD
50 +(T-8
33-
37.2
5 ;1%=K 2%7+8J
%&$'(
20 ;1
%=K '8+8%S;O6
(*=A6H8:O/
;17018%1!8=E&6
VMKO!8=5670
- +(
!)H8:O/1
8%;>%7L
;,74U,:60
/86TA8+'G$
TA8+1A/8$)%$EM-+
TA8+=8A
17*$)(*;>%7L:6C6A"/8$ >
H86A617*$C!3>H86A60#1!0!$ C
,-.+-+
( !,-#"680
"#$0#1
- )H8:O/;17018%1!8=E&6
VMK)&B$!&1@
J3:4C!30#1 U0=;2W6
%G2C44:4
34
!&1@J
3 C!3'(0#1
3 !&1@
J3
E7)G;6(=
E%%J
( (2
543)
17*$5H8
C1++
8C44
;I(=4
E!&6
10-6
0
- T-8
LD
30-L
D50
TF#2%7+8J
%&$'(
21-3
5 ;1%=K
-;2#%K;PQ6,
KTA8+#=G-%#0 TA8+1A/8$)%$EM-+ C!3>H86A617*$!0!$
- 1-#:O/;170!&1@
J3 702
1,7 TF# ,/6
CT%3C1%6
:40-8$C!3O=
71 "6800#1
;2!(*=6.2)
&B$:OL
-"?B6C!3;!
Q1!$ 0
#1;2!(*=6'(>81+
-A$;2W6+
-A$#-#6 "8A
C!3+-A$'!&4"
8A
- 2%7+8J%&$'()
(* 60 ;1%=K >3)H8:O/E
F553$&118%;>%7L
;,74U,C!3,8=:6)
(*'M0
45
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
>$>76,K
(252
0)
O6-#
C1++
8 C44
;I(=4
E!&6
10-3
0 - !"#$%&$'()H8:O/48$O6-#.+-$#1
- EM)
V%&1@
8)(*$#1+
(18%;2
!(*=6C2!$;1
(*=A1&4
TA8+'G$
TA8+1
A/8$ C
!3TA8+=8A"#$:4
U0=;IE
83:61!M-+)
(*I8=%&$'( 3
0 ;1%=K E4!&1@
J3CS4
'("8A"680,-8$
9 "6861
&4;'/6:4
:4+&1>3;A/84%7;A
J1!8$:4
"6800#1
:648$,/6;!Q1!$
- ;2#%K;PQ6,
K"#$.+UT%'2#%K56
7021,7>81,/6)
(*.+-.0/%&4%&$'(>3'G$1A-8,/6
)(*.0/%&4%&$'(
EM)V%&1@
8
6&6)6
8 (2
525)
O6-##-#6C!3,/6
1!/8!G1 '+
C1++
8 C44
;I(=4
E!&6
5-20
- E4
18%;2
!(*=6C2!$)8$0/861
8%;>%7L
;,74U, TA8+'G$
C!3"680"#$:4
- 18%;2
!(*=6C2!$!&1@
J3N8=6#1"#$:4)&B$:6O6
-##-#6C!3,/61
!/8!G1 '+)
(*I8=%&$'()M1
2%7+8J%&$'(;170"?B6
;E(=$5&*AT%8A
- !&1@J
3)(*;2!(*=6C2!$.0/C1- +(CS4
'(;O!F#$"6
80,-8$ 9
"6861
&4;'/6:4
:4;A/8>81"#4:4;"/8'G-1!8$:4 :4O6
8"?B6C,-"680;!Q1!$
48$,/6
0#1;"/+
"?B6O%F#>8$!$;!Q16
/#= 48$,/6
0#1T!(*486.+-O+
0 - %&$'()
H8:O/;170!&1@J
3 702
1,7"#$UT%U+UP+ U0=;2#%K;P
Q6,K;P!!K)(*+
(UT%U+UP+
7021
,7+('O'&+E&6
VK1&42
%7+8J
%&$'(:6;57$;'/6,
%$)&B$:6O6
-#C!3
,/61!/8)(*.0/%&4%&$'(
46
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
CE%;P
(*=$.^/
#%MJ( C!3
6A!IA(
(253
4);
Won
gpiy
as
atid
and
Hor
mch
an
(200
0)
!H8,/6
C1++
8 C44
;I(=4
E!&6
10-4
0
- ;E7*+TA8+S
(*"#$18%1!8=C!3!&1@
J3"#$18%1!8=:O/'
G$1A-8)(*;170"?B6
;#$
,8+V%%+58,7
- 18%;2
!(*=6C2!$"#$0#1 )&B$%G2%-8$ "
680 '(
1!(40#1 !&1@J
3;1'%)&B$
;E< G/C!3;E
<;+(=
3-15
2%7+8J%&$'(
3-15
;1%=K .+
-+(18%C,1,8;1
70"?B6O
!&$>81I8=%&$'('-A6
57B6'-A6)
(*.+-.0/%&418%I8=%&$'(+(18%C,1,8C!3;>
%7L;,74U,;2W6
21,7
UEV7]4&!!&$1K
(#U1!6(+8
)
E&6VKA0(
(254
6)
57B6'-A6!H8,/6:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
1-
4
- 2%7+8J%&$'(
1-4 ;1%=K E4A-8+(18%C,1,8C!318%;>%7L
;,74U,!0!$,8+
2%7+8J%&$'()
(*;E7*+"?B6
#8N8N%JK
(253
9)
O&A=-#=)
(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
3-6
- )H8:O/+(6
BH8O6&1
,/6)(*;E7*+
6/#=1A-8,/6
)(*.+-.0/)
(*I8=%&$'(
- !0>H86A6O
&A=-#=
!7!!(*
AU%N8'
(254
4)
'-A6"
#$1!(4O
&A)(*
;!(B=$:6'N
8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
2-8
- 2%7+8J%&$'(
0-4 ;1%=K ;6
FB#;=F*#+(18%;>%7L
;,74U,.0/
- 2%7+8J%&$'(
6-8 ;1%=K =&$.+-+
(18%;2
!(*=6C2!$O!&$18%I8=%&$'(
30 A&6
47
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
A-86'(*)7<
#,76&6
)K (2
545)
,/6#-#6:6'N8E
2!#0;5FB#
C1++
8C44
;I(=4
E!&6
5-25
- T-8
LD
50 :6%3=3;A!8
3 ;0
F#6 ;)
-81&4
11.7
;1%=K
- ,/6
#-#6A-86'(*)
7<)(*.0/%&4%&$'('G$"?B6+
(C6AU6/+
;2#%K;PQ6,
K18%%#05(A7,!0!$
- 18%;>
%7LC!3E&Z6
8;2W6,
/6#-#6.0/5/81A-857B6'-A6O&A)(*.+-.0/%&41
8%I8=
%&$'(
- O!&$=/8=2!G1.+-E
4!&1@
J3 702
1,7.2>81,/6)
(*.+-.0/%&418%I8=%&$'(
;<%@R(1/86)#
$
%&56(=
K (2
544)
;+!Q0
C1++
8C44
;I(=4
E!&6
10-3
20
- #&,%818%$#1"#$;+
!Q0!0!$,8+2
%7+8J
%&$'()
(*;E7*+"?B6
- ;+!Q0$#15/8+81"?B6
,8+2
%7+8J
%&$'()
(*;E7*+"?B6
- +(18%C,1=#0;%QA
"?B6
- 18%;2
!(*=6C2!$'("#$C -6
:4 ;+
F*# -8618%I8=%&$'( %30&4
80,
90 C!3
100 ;1%=K
O6/8A&A
4ML+(
(251
8)
,/6)&B$,/6
C1++
8
8-32
- 18%;>
%7L;,74U,!0!$,8+2%7+8J"#$%&$'()(*;E7*+
"?B6
- !&1@J
3"#$>86%#$0#1C!32!(;2!(*=6%G2.2
- %&$'(+( !,-#18%;1
70O6-#:O+-
- 2%7+8J%&$'(
12-1
6 ;1%=K )H8:O/>86%#$0#1C!32!(;2!(*=6
C2!$+81)
(*'M0
- 2%7+8J%&$'(
16-2
0 ;1%=K )H8:O/+
(18%;2
!(*=6C2!$!&1@
J3"#$:4+81)
(*'M0
- 2%7+8J%&$'(
12 ;1
%=K"? B6
.2 +( !,-#18%;2
W6O+&6
"#$O6/8A&A
- 2%7+8J%&$'(
32 ;1
%=K )H8:O/,
8=O6
/8A&A,8=S?$
60 ;2#%K;PQ6,
K
48
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
O6/8A&A
(,-#)
A755M,8
(253
7)
,/6)(*;!(B=$:6'N8E
2!#0;5FB#
C1++
8C44
;I(=4
E!&6
1-5
- #&,%818%%#05(A7,!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- 18%;1
70=#0:O+- C!3TA8+'G$!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- 1-#:O/;170!&1@
J3 702
1,7"#$:4 18%;%
(=$,&A"#$:4 TA8+=8A2!/#$ C
!318%;1
7017*$"/8$
- O!&$>81=/8=2
!G1E4
,/6)(*+
(!&1@J
3:4;!Q1
C!3:40-8$
##67U)1
8!&+
1_@J
8 (2
545)
=#0#-#6)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
10-1
60
- T-8
GR
50 ;)
-81&4
22.1
;1%=K
- 2%7+8J%&$'(;E7*+
"?B6+(
!)H8:O/1
8%;>%7L
;,74U,!0!$
- 2%7+8J%&$'( ,&B$C,- 4
0 ;1%=K"?B6
.2 >3+( !=&4
=&B$18%;>
%7L;,74U,C!3=#0
>3,8=:6)
(*'M0
- )H8:O/:4CT%3'&B6
18%C,1=#0;2W61
%3>M1
:4!8=
- )H8:O/;17018%;2!(*=6C2!$UT%$'%/8$"#$UT%U+UP+
;5-6 ;17018%"80O
%F#O&1
4%7;AJ
sat
ellit
e
#6G;4(='
08A'A%%TK
(254
5)
'-A6"#$2!8==#0
)(*;E83;!
(B=$;6FB#;=F*#
C1++
8C44
;I(=4
E!&6
4.5-
45
- +(T
-8 GR
5O 2%3+8J
18 ;1%=K O!&$I8=%&$'(C!3;E
83;!(B=$;2W6;A!8 6
;0F#6
- 18%;>
%7L;,74U,!0!$,8+2%7+8J%&$'()
(*;E7*+"?B6
- +(!&1@
J3:40
-8$;170"?B6
- %&$'(,&B$C,-
27 ;1
%=K"?B6
.2 )H8:O/=
#0CT%3C1%6
- ,/6
)(*.0/%&4%&$'( 4
.5-1
8 ;1%=K E4=
#0)(*+
(!&1@J
3:4;2W6
'#$CI1
!&1@
J3
;'/61
!8$:46
G6"?B6+
8 C!3:4'#$:4C,-+
(1/86:4%-A+1&6
49
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
#;+P
#6
#M.% (
2542
)
,/6#-#6)
(*.0/>81
18%;E
83;!(B=$
;6FB#;=F*#
C1++
8C44
;I(=4
E!&6
3.35
-5.5
0
- T-8
LD
30-L
D50
:6%3=3;A!8
4 ;0
F#6 2%3+8J
3.3
5 C!3 5
.50 ;1%=K
- )H8:O/,/6
#-#6+
(18%;>
%7L;,74U,5/81A-8,/6
#-#6)
(*.+-.0/%&4%&$'(
- ,/6
#-#6)
(* -861
8%I8=%&$'()M1
%30&4
)H8:O/+
(>H86A6UT%U+UP+
7021,7;+F*#
;)(=4
1&4,/6
#-#62
1,7
.,%;+
V (2
547)
'-A6=
#0)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;%FB#%&$
35.5
1-31
9.67
- G
R50
(30) ;)
-81&4
166.
47 ;1
%=K
- 5M0)(*.0/%&4%&$'( 3
19.6
7 ;1%=K )H8:O/+
(#818%:40
-8$ :4
!8=>M0
- 5M0)(*.0/%&4%&$'( 3
5.51
;1%=K ,/61
!/8+(18%;>%7L
;,74U,+
81;+F*#;)(=
41&4
cont
rol
U!4(;!(=
<%8=M)V
(254
7)
'-A6=
#0)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
20-1
00
- T-8
LD
50(3
0) ;)
-81&4
58.1
;1%=K C!3
GR
50(3
0) ;)
-81&4
52.1
3 ;1%=K
- )(*%30&4%&$'( 6
0 ;1%=K E4!&1@
J3:4;2!(*=6
C2!$
.0/C1- :4
0-8$ C
!3:4'(
5+EG#
++-A$
- :6%M-6
M1V
2 ,/6
)(*.0/%&42
%7+8J
%&$'(
20 C!3
40 ;1%=K E4A-8+(18%C,1
=#0+81;+F*#;)(=
41&4,
/6)(*.+-.0/%&4%&$'( 18%,8="#$;6
FB#;=F*#)(*.0/%&42
%7+8J
%&$'( ,&B$C,- 6
0 ;1%=K "?B6.2 C!3>3,8=;E7*+"?B6,
8+2%7+8J%&$'()
(*;E7*+"?B6 :6
,/6)(*.0/%&4%&$'(2%7+8J
60 ;1%=K E4,
/6'("8A C!3%&$'(2%7+8J
80 ;1%=K E4
,/6)(*+
("680;!Q1+
81C!3;>
%7L;,74U,5/81A-8,/6
)(*.+- .0/%&4%&$'(
50
!"!#$%& 4
( '()
5F*#EF5
5F*# G/<?1@
8 '-A6"#$EF5
)(*6H8+8
I8=%&$'(
%&$'()
(*:5/
2%7+8J%&$'()
(*:5/
(;1%=K, G
y) 1/
!"#$%&$'()(*+
(,-#EF5
2/
Cry
ptoc
oryn
e bl
assi
i '8%76(
(254
8)
'-A6=
#0)(*;!(B=$:6
'N8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
20-1
00
- T-8
LD
50(3
0) ;)
-81&4
56.2
4 ;1%=K C!3
GR
50(3
0) ;)
-81&4
52.1
3 ;1%=K
- 18%C,1=#0!0!$;+
F*#2%7+8J
%&$'(;E7*+
"?B6
- E4
18%,8="#$;6
FB#;=F*#
)(*.0/%&42%7+8J%&$'(
40 ;1
%=K"?B6
.2 C!3;17018%
,8=)
&B$O+0)(*2
%7+8J
%&$'(
100 ;1%=K
- )(*2
%7+8J
%&$'(
20 C!3
40 ;1%=K E4!&1@
J3:40
-8$
- :6%M-6
M1V
3 E4
A-8 %&$
'(2%7+8J
80 ;1%=K E4,
/6)(*+("680;!Q1+81C!3
;>%7L
;,74U,5/81A-8,/6
)(*.+-.0/%&4%&$'(
Cry
ptoc
oryn
e
wen
dtii
“gre
en”
2`a6#M4!
(254
8)
;6FB#;=F*#,/6
#-#6)
(*;!(B=$:6'N
8E2!#0;5FB#
C1++
8C44
;I(=4
E!&6
10-5
0
- ,/6
)(*.0/%&4%&$'(2%7+8J
10 ;1%=K +(18%;2
!(*=6C2!$!&1@
J3"#$:4
TF# :4
;%(=ACO!+ :4;2W6%G2O
#1 "#4:4O=
&1;2W6
T!F*6
- ,/6
)(*.0/%&4%&$'(2%7+8J
20 ;1%=K +(18%;>
%7L;,74U,C!3>H86A6O
6-#!0!$
'(:4;2!(*=6
C2!$
!H8,/6;!Q1!$
)*!+,) -
1/ O6
-A=A&0%&$'()
(*C2!$:O/;2W6O
6-A='81!
(SI un
it) ;%(=15F*#O6
-A=A-8
;1%=K (g
ray,
Gy)
2/
'%M2C!3T&0!#1+8>81%8=$8618%<?1@
8)(*'8+8%S%A4%A+.0/
51
!"#$% !"&'(#)*+,(-''. !"/012&'(#)*-)/34567!89:; #"<2%%2'=)%/1267"%,2>-)/345?#>6="')@A:
!"#$% !"&'(#)*+,-.)%/0123"%4156)/789:#52."');<,2/&'(=+> +5 6*?@1!,=6,5; 3 ./A#=+@1/)B/ +,-&'(<*C?1!<01='DEE/ #"6)/78923!@+,-#, :#5 #"!,%1'FGB/+(=*,5/6)/7896H. =%IJ'%'/01 &F5156)/789 K4(E013/@152/+"$;J41#K4"? #"K%@ 68+7')%I1 (Canna generalis) K6'=L,-5; M" (Portulaca grandiflora) K4(=*NE!1> (Chrysanthemum morifolium) LG-;=&O/P4;1/%1'?AE)5K4(6)Q/1F$;/)%?AE)5!31?A+514)5=%IJ'>1<J'9+)B;<AB/ K4(5);!, !"#$% !"&'(#)*$,%3415./A#
LG-;%014);$5R@'(3?@1;%1'#01=/A/%1'?AE)5 %1'C)#=4H$%6)/789 K4(%1'F5156)/789=6H-$=6A-!E01/?/
=.@/ FA;K#; &+8!!1 %8341*3A/ =&O/J"/ ($'8S,, 2546)
1. -493')%B2 (Canna generalis)
68+7')%I16)/78923!@+,- #"E1%%1'T15');<, +,->R/59*'A%1'T15');<,K%!!1K4(?AE)5/A?=C4,5'9=+C:/:45, <U1*)/?AE)5K4(6)Q/1K3@;!31?A+514)5=%IJ'>1<J'9 #"J)B;.H-$K4(FGB/+(=*,5/6)/7896H. ?"%)*!31?A+514)5=%IJ'>1<J'9 2/.@?;&V 6.>. 2543-2545 !,E01/?/ 22 6)/789 #);/,B
2/&V 6.>. 2543 !,E01/?/ 4 6)/789 #"K%@ 6)/789C',!&'(6)/796;I9 6)/789<"!<A'/8.
6)/789=34H$;$'8S, K4(6)/789.!6R6,'/8.
2/&V 6.>. 2544 !,E01/?/ 4 6)/789 #"K%@ 6)/789K#;W+7, 6)/789?)/?A<1 6)/7896A!69=;A/
6)/789<"!');<, 6)/789.!6R6''S, K4(6)/789/X1?''S
2/&V 6.>. 2545 !,E01/?/ 14 6)/789 #"K%@ 6)/789?ANYR 6)/789=6DN6AJ' 6)/7897,'( 6)/7896A*R4>A4&Z 6)/789/W+8! 6)/789<8+7,'1 6)/789K#;?A:'E 6)/789/?4T?, 6)/789&'1:!+59 6)/789/W&?)E%9 6)/7896A!69');<, K4(6)/789K<;=+,5/ :#52/E01/?//,B!, 2 6)/789 +,- !@ #"=%A#E1%%1'T15');<,:#5J'; KJ@=%A#%1'%4156)/789J@$E1%6)/789%415K4"? #"4)%IS(+,-#, #"K%@ 6)/7896A!69');<, K4(6)/789K<;=+,5/
:#56)/7896A!69');<, %4156)/789J@$E1%6)/789%415=34H$;$'8S, LG-; #"E1%%1'T15');<,K%!!1K**=T,5*64)/ (acute irradiation) <@?/6)/789K<;=+,5/ =&O/6)/789+,- #"E1%%1'&4R%=!4D#F$;6)/789%415<"!<A'/8. LG-;=&O/6)/789%415+,-!1E1%%1'T15');<,K%!!1K**=T,5*64)/ (acute irradiation)
(>R/59*'A%1'T15');<,K%!!1K4(?AE)5/A?=C4,5'9=+C:/:45,, 2546)
E8#=#@/+,-!,C8SC@1+1;%1'J41#
1. !,<,<)/<#2< <?5;1! !,4?#415F$;%4,*#$%K&4%23!@ 2. !,F/1#F$;J"/ K4(#$% 23"=4H$%341%3415
3. 4)%IS(2*#@1;+,-<?5;1!
52
6)/7896A*R4>A4&Z 6)/789/?4T?,
6)/789?)/?A<1 6)/789?ANYR
6)/789/W&?)E%9 6)/789/X1?''S
53
%1'2."68+7')%I1&4R%&'(#)*J%KJ@;*'A=?S=%1(%41;U//+0123"#R<?5;1! &$,%K**
4)%IS(#@1;+,- =%A#E1%%1'T15');<,%D=&O/+,-/@1</2EK%@PR"+,-.$*<(<! !"#@1;
68+7 ') %I 1+,- & 4R %& ' (#) **'A = ?S'A !/B0 1 =6A- !C?1!<?5;1!K4(.@ ?5&[$;%)/%1'6);+415F$;#A/*'A=?S'A!J4A-; #"=&O/$5@1;#,
>R/59*'A%1'T15');<,K%!!1 K4(?AE)5/A?=C4,5'9=+C:/:45, !31?A+514)5=%IJ'>1<J'9
C2-9:; 5 68+7')%I1<156)/789J@1; \ +,- #"E1%%1'T15');<,.)%/0123"=%A#%1'%4156)/789 %1'2."&'(:5./9 K4(>R/59*'A%1'T15');<,K%!!1 K4(?AE)5/A?=C4,5'9=+C:/:45, !31?A+514)5=%IJ'>1<J'9
54
2. +-'DE:;>@ F" (Portulaca grandiflora)
K6'=L,-5; M"6)/78923!@+,- #"E1%%1'T15');<,/)B/ 6*?@1&]EE8*)/X1C?A.1');<,&'(58%J9K4( $:L:+& CS(?A+51>1<J'9 !31?A+514)5=%IJ'>1<J'9 #"FGB/+(=*,5/6)/7896H. ?"%)*!31?A+514)5=%IJ'>1<J'9 E01/?/ 10 6)/789 #);/,B
2/&V 6.>. 2542 !,E01/?/ 3 6)/789 #"K%@ 6)/789 !%. 1, 6)/789 !%. 2 K4(6)/789 !%. 3
2/&V 6.>. 2543 !,E01/?/ 2 6)/789 #"K%@ 6)/789.!6R&'(X1')E K4(6)/789K6++A%
2/&V 6.>. 2546 !,E01/?/ 5 6)/789 #"K%@6)/789=%IJ'>1<J'9 60, 6)/789.!6R$)N.)N
6)/789<"!$'8S,, 6)/789X)+',51 K4(6)/789F1?&'(+,& (>R/59*'A%1'T15');<,K%!!1K4(?AE)5/A?=C4,5'9=+C:/:45,, 2546; Wongpiyasatid and Hormchan, 2000)
LG-;<1!1'UK*@;K6'=L,-5; M"J1!4)%IS(F$;#$% #"=&O/ 3 &'(=X+ CH$
- 6)/789#$%L"$/ #"K%@ 6)/789 !%. 3, 6)/789.!6R$)N.)N K4( 6)/789<"!$'8S, - 6)/789#$%%G-;L"$/ #"K%@ 6)/789X)+',51 K4( 6)/789F1?&'(+,&
- 6)/789#$%41 #"K%@ 6)/789 !%. 1, 6)/789 !%. 2, 6)/789.!6R&'(X1')E, 6)/789K6++A%
K4(6)/789=%IJ'>1<J'9 60
E8#=#@/+,-!,C8SC@1+1;%1'J41#
1. !,<,<)/<#2< K4(<?5;1! !,4?#415F$;%4,*#$%K&4%23!@ 2. 4)%IS(F$;#$% !,+)B;#$%L"$/ #$%%G-;L"$/ K4(#$%41 (#$%.)B/=#,5? )
K4(*1;6)/7894)%IS(#$%K&4% &E1%K6'=L,-5; M"+)-? \ &
3. #$%$$%J4$#&V K4(#$%#%
4. #$%*1/+/
5. PGB;/A5!!1=%D*=%<'2/K6'=L,-5; M"&'(=X+#$%41!1%%?@1#$%L"$/
55
6)/789<"!$'8S, (Wongpiyasatid and Hormchan, 2000)
6)/789 !%. 3
6)/789.!6R&'(X1')E
(Wongpiyasatid and Hormchan, 2000)
6)/7896)/789.!6R$)N.)N
C2-9:; 6 K6'=L,-5; M"6)/789J@1; \ +,- #"E1%%1'T15');<,.)%/0123"=%A#%1'%4156)/789 P4;1/%1'?AE)5K4(6)Q/1F$;!31?A+514)5=%IJ'>1<J'9
3. D*G<!2H (Chrysanthemum morifolium)
=*NE!1>6)/78923!@+,-%4156)/789E1%%1'T15');<, #"FGB/+(=*,5/6)/7896H. ?"%)*!31?A+514)5=%IJ'>1<J'9 2/&V 6.>. 2545 E01/?/ 4 6)/789 CH$ 6)/789=%IJ'>1<J'9 60-1,
6)/789=%IJ'>1<J'9 60–2, 6)/789=%IJ'>1<J'9 60–3 K4(6)/789=%IJ'>1<J'9 60-4 LG-;+)B; 4 6)/789 %4156)/789!1E1%%1'T15');<,23"%)*6)/789 Taihei LG-;!,#$%<,!@?;K#; K4(!,6)/789%415+,- !@ #"=%A#E1%%1'T15');<,.)%/0123"=%A#%1'%415:#5J'; E01/?/ 2 6)/789 #"K%@ 6)/79=%IJ'>1<J'9 60-5 #"E1%%1'/01=$1%4,*#$%E1%6)/789%415#$%<,.!6R$!!@?; (Purple Group # 75 B) +,- #"E1%%1'T15');<,K%!!1K**:C'/A% (chronic irradiation) 2/3"$;T15');<,K%!!1 LG-;!,:C*$4J9 –60 %014);K'; 200
CR', =&O/J"/%01=/A#');<,K%!!12/&'A!1S 62.8 =%'59 !1=61(=4,B5;=/HB$=5H-$ K4(6)/789=%IJ'>1<J'9 60 –6 #"E1%%1'/01=$1%4,*#$%E1%6)/789%415#$%<,<"! (Orange Group # 24 D) LG-; #"E1%%1'T15');<,K%!!1K**:C'/A% (chronic irradiation) 2/3"$;T15');<,K%!!1 LG-;!,:C*$4J9–60 %014);
56
K'; 200 CR', =&O/J"/%01=/A#');<,K%!!12/&'A!1S 112 =%'59 !1=61(=4,B5;=/HB$=5H-$ (>R/59*'A%1'T15');<,K%!!1K4(?AE)5/A?=C4,5'9=+C:/:45,, 2546; Lamseejan et al., 2000)
E8#=#@/+,-!,C8SC@1+1;%1'J41#
1. <,#$%=&4,-5/ &E1%6)/789=#A! !,C?1!<?5;1!K4(341%3415!1%FGB/+0123"PR"*'A:XC
!,J)?=4H$%!1%5A-;FGB/
6)/789=%IJ'>1<J'9 60-1 6)/789=%IJ'>1<J'9 60–2
6)/789=%IJ'>1<J'9 60-3 6)/789=%IJ'>1<J'9 60-4
C2-9:; 7 =*NE!1><156)/789J@1; \ +,- #"!1E1%%1'&')*&'8;6)/789:#5%1'2."');<,.)%/0123"=%A#%1'%4156)/789 (+,-!1: http://www.rdi.ku.ac.th/gamma-irradiation/database/Mutation.htm)
57
-''. !"/012&'(#)*
%1'T15');<,'@?!%)*%1'=61(=4,B5;=/HB$=5H-$%)*6''S !"/B01./A#J@1; \ =6H-$.)%/0123"=%A#%1'%4156)/789 +0123"=%A#4)%IS(KJ%J@1; &E1%J"/&%JA =.@/ J"/KC'( 2*#@1; 2*=PH$% 3'H$=%A#KU*
=&O/J"/ +0123"=%A#6)/78923!@ \ $$%!1!1%!15 :#5%1'?AE)56)Q/1&')*&'8;6)/789 F$;%48@!;1/?AE)56''S !"/B01 <U1*)/?AE)5<)J?9/B01<?5;1!K4(6''S !"/B01 <01/)%?AE)5K4(6)Q/1&'(!;/B01EH#
%'!&'(!; 2/&]EE8*)/!,6''S !"/B013415./A#+,-&'(<*C?1!<01='DE2/%1'.)%/0123"=%A#%1'%4156)/789 #"4)%IS(J"/=J,B54; 3'H$J"/KC'( F/1#F$;2* 'R&'@1;2* K4(<,2*=&4,-5/K&4; #"J"/6''S !"/B01EA^?+,-!,C?1!K&4%23!@<?5;1! #"K%@ $/R=*,5< ‘/1/1’ (Anubias nana) $/R=*,5< ‘C$/=E/LA<’ (Anubias congensis) $/R=*,5< ‘*1'9=+$',-’ (Anubias barteri) $/R=*,5< ‘!A/A!1’ (Anubias
minima) :4*A=4,5 (Lobelia cardinalis) K4($:%4/,!1 (Aglonema simplex) =&O/J"/ (%48@!;1/?AE)56''S !"/B01, 2550)
E8#=#@/+,-!,C8SC@1+1;%1'J41#
1. !,4)%IS(J"/KC'( J"/EA^? 2*#@1; <'"1;C?1!K&4%23!@ K4(C?1!341%3415F$;<156)/78923"!1%FGB/
2. <1!1'U&4R%=4,B5; #"2/X1./(+,-E01%)# K4(+01%1'&_A*)JA#RK4')%I1 #";@15
58
'R&K**%1'E)#6''S !"/B012/X1./(F/1#=4D% 6''S !"/B01EA^?+,-<1!1'U=4,B5; #"2/K%"? <'"1;
C?1!<?5;1!$,%K**
:4*A=4,5 (Lobelia cardinalis) $/R=*,5< ‘/1/1’ (Anubias nana)
$:%4/,!1 (Aglonema simplex) %1'E)#J%KJ@;<U1/+,-#"?56''S !"/B01
C2-9:; 8 6''S !"/B01EA^?+,- #"E1%%1'.)%/0123"=%A#%1'%4156)/789:#5%1'T15');<,K%!!1 +,-/01!1K<#;2/;1/?)/&'(!;/"$!=%4"1 C')B;+,- 18 ?)/+,- 24-27 !AU8/15/ 2549 S >R/59K<#;<A/C"1K4(%1'&'(.8!$A!K6C=!H$;+$;71/, (%48@!;1/?AE)56''S !"/B01, 2550)
59
I"$#:-I"$DA:> I$@%2'6="')@A:%)* !"#$% !"&'(#)* +,(-''. !"/012&'(#)*
%1'2."');<,%)* !"#$% !"&'(#)*'?!UG;6''S !"/B01&'(#)*/)B/ 6*?@1!,F"$#,-F"$=<,5
:#5<1!1'U<'8& #"=&O/F"$ \ #);/,B
J2'2@9:; 5 F"$#,-F"$=<,5F$;%1'2."');<,%)* !"#$% !"&'(#)* K4(6''S !"/B01&'(#)*
F"$#, F"$=<,5
1. =&O/?A7,%1'&')*&'8;6)/789+,-=3D/P4 #"'?#='D?:#5+0123" #"4)%IS(23!@ \ 3'H$<1!1'U<'"1;6)/78923!@='D?%?@1%1'&')*&'8;6)/789K**&%JA
1. J"$;!,3/@?5;1/=T61(3'H$3/@?5;1/+,- #"')*$/8N1JAK4"?=+@1/)B/ 2/%1'#01=/A/%1'J@1; \
=%,-5?%)*%1'2."');<, '?!UG;%1'')*PA#.$*P4+,-=%A#FGB/2/%1'2."');<, =/H-$;E1%!,$)/J'15'"15K';!1%31%2." !@UR%?A7,
2. <1!1'U<'"1;4)%IS(+1;6)/78%''!23!@ \
+,- !@=C5!,$5R@ 2/&'(.1%'=#A!J1!7''!.1JA LG-;<1!1'U/01 &2."=&O/K34@;6)/78%''!2/%1'&')*&'8;6)/789J@$ &
2. !@<1!1'U+01 #":#5+)-? & =/H-$;E1%$8&%'S9K4(=C'H-$;!H$+,-=%,-5?F"$;%)*%1'2."');<,!,'1C1K6;!1%
3. !@!,');<,J%C"1;$5R@2/6H.34);E1%T15');<,K4"?
+0123" !@=&O/$)/J'15K%@!/8I59 3. PR"E(2."');<,J"$;=&O/PR"=.,-5?.1N=T61(#"1/3'H$!,C?1!'R"K4(C?1!.01/1N=&O/$5@1;#, +)B;2/='H-$;F$;');<, %1'2."');<, %1'&')*&'8;6)/7896H.
%1'F5156)/7896H. =&O/J"/
4. 2."+#K+/<1'=C!,2/%1'%01E)#K!4;>)J'R6H.
=.@/ 2."%01E)#=64,B5 `+,-JA# &%)*%4"?5 !"%@$/+01%1'<@;$$% =&O/J"/
4. !,&]EE)5C?*C8!3415&]EE)5+,-+0123" !@&'(<*P4<01='DE =.@/ ./A#6H. ');<,+,-2." &'A!1S');<, '(5(=?41+,-T15');<, '?!UG;?A7,%1'T15');<, =&O/J"/
5. =6A-!C?1!341%3415F$;6)/7896H.23"!1%FGB/
60
!!"# 5
$%&' ()*+%,- ./01234$53.50
$%&'6/
!""!#$ % &'()$*!+,-./"+,-0#1.&)234,3#!'5!6"!#7!'#&5839(:4/"!#0#&)0#;5(&6<;=>60#192?+2' ()$*!,32&@58%@6,!"$*! 64 A6%. (species) B.'8*$6>CD*90E6+,-./" F!6$6,!""$*! 44 A6%.
90E6+,-0#1.&) F!6$6,!""$*! 10 A6%. GH190E6(##I+,-6@F!0#1.&) F!6$6 10 A6%. JK45,C!$%2'!H&'9"LM#?!8M#= 90E6C6*$'5!6234,3"!#?K"L!$% &'9"34'$"&)NHO/5#&583234,3M*/+,-./"+,-0#1.&),!"2348;. PK5#-/'H1 79 GH18*$6,!"90E6"!#?K"L!B.'"!#>A-#&583G",,! 7!'G))973')(H&6 "&)A%@68*$6C#:/M-6(:A2349H3@'5>68Q!(0H/.9A:@/ (9(!19H3@'596:@/9':4/)
B.'NH2349"%.OK@6CH&5 !"2F!"!#7!'#&583>C-"&)+,-./"+,-0#1.&)6&@6 ()$*!8*$6>CD*,3NHC#:/H&"LI1234G8.5//",!9C,:/6"&)(:A/:46 R 234N*!6"!#7!'#&583 B.'8!,!#P F!G6"//"90E6"H;*, +.-.&563@
1. #&583,3NHM*/"!#90H34'6G0H5O/5BS#B,BJ, 9A*6 "!#GM"C&" C#:/O!.C!'+0O/5BS#B,BJ, "!#9(%4,C#:/H. F!6$6BS#B,BJ, 90E6M-6
2. #&583NHM*/H&"LI12!58&IT!6GH18#3#1O/5(:A 9A*6 H&"LI1GS#1G"#6
"!#90H34'6G0H5O/5>) HF!M-6 90E6M-6
3. #&583,3NHM*/"!#90H34'6G0H583./"GH1>) +.-G"* H&"LI1.*!5O/58*$6M*!5 R
"!#90H34'6G0H5O/583./" #U0#*!5O/5./" O6!.O/5./" 90E6M-6
4. #&583,3NHM*/NHM*/"!#9 #%D9M%)BM JK45()$*!#&583,3NH2&@59#*5GH1'&)'&@5"!#9 #%D9M%)BMO/5+,-./"+,-0#1.&)
5. #&583,3NHM*/"!#5/" GH190/#=9JV6M=S$!,/'U*#/.O/59,HV.+,-./"+,-0#1.&)
6. #&583,3NHM*/#1))"!#8:)(&6<;=O/5+,-./"+,-0#1.&) B.'2F!>C-9"%."!#90E6C,&6
7. #&583,3NHM*/$&W &"#A3$%MO/5+,-./"+,-0#1.&) 9A*6 ,3NHM*/#1'19$H!//"./" #1'19$H!"!#)!6O/5./" #1'19$H!9"V)9"34'$ XHX
8. #&583,3NHM*/H&"LI12!58#3#$%2'!CH&5"!#9"V)9"34'$O/5+,-./"+,-0#1.&)
>6"!#7!'#&583>C-G"*+,-./"+,-0#1.&)6&@6 NH"#12)2349"%.OK@6CH&5 !""!#7!'#&583 8!,!#P9"%.+.->6CH!' R H&"LI1#*$,"&6 9A*6 ,3H&"LI1GS#1G"#6#*$,"&)>).*!5 90E6M-6 JK45"!#7!'#&5838!,!#PA&"6F!>C-9"%."!#"H!'(&6<;= K590E60#1B'A6=/'*!5'%45>6"!#0#&)0#;5(&6<;=+,-./"+,-0#1.&) 9(:4/>C-+.-(&6<;=>C,*234,3H&"LI1.3M!,234M-/5"!# B.'97(!1>6"#I3234+,*,3GCH*5O/5'36M!,M-/5"!# C#:/H&"LI1234M-/5"!#+,*8!,!#P2F!>C-9"%.OK@6+.-9,:4/,3"!#0#&)0#;5(&6<;=B.'$%<3/:46
61
.&56&@6"!#7!'#&583 K590E62!59H:/">6"!#0#&)0#;5(&6<;=+,-./"+,-0#1.&)/3"$%<3C6K45234S$#86&)86;6>C-,3"!#?K"L!$% &',!"OK@6
()*+%,-6/
78293:782'%09; !"#8"%+<=+5:+%>?2%;=$"4@A#3:+%'%; '%&=@;5B&->5'%04!C7!<
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
9A*6 >6+JBM(H!8JK, JK45,36@F!90E6/5S=0#1"/)PK5 80 90/#=9JV6M= B.' 1"*/>C-9"%./6;,UH/%8#1 F!6$6,!" 8*5NH>C-9"%."!#90H34'6G0H5 C#:/8#-!5S$!,983'C!'90E6/'*!5,!"M*//5S=0#1"/)O/59JHH= (Kovács and Keresztes, 2002)
CH&5 !"234(:A+.-#&)#&583+0GH-$ 1()H&"LI1"!#"H!'+.-CH!'G)) B.'"!#"H!'2349CV69.*6A&. 9#3'"$*! ,!BS#,%$92A&46 (macromutation) 9A*6 "!#90H34'6G0H5H&"LI1Q!'6/"O/58*$6M*!5 R O/5(:A NHO/5"!#7!'#&583>6A&4$234 1 (M1 generation) B.'+,*8!,!#P#&"L!H&"LI19CH*!63@+$-+.->6#;*6M*/+0 GM*9,:4/7!'#&583G",,!"&)8*$6/:46O/5(:A6/"9C6:/ !"9,HV. (vegetative part)
,&"+.-"!#"H!'G))P!$# JK45/! F!G6"+.-90E6 2 H&"LI1 S:/H&"LI1G#" 90E6"!#"H!'2349"%.OK@62&@5M-6C#:/2&@58*$6O/5(:A (solid mutant) 90E6"!#"H!'2348!,!#P#&"L!H&"LI1"!#"H!'GH1O'!'(&6<;=M*/+0+.- .-$'$%<3"!#O'!'(&6<;=G))+,*/!?&'9(? GH1H&"LI12348/590E6"!#"H!'9(3'5)!58*$6 (partly mutant) C#:/+S9,#! (chimera) "!#"H!'(&6<;=2349"%.OK@697(!18*$6 /! 9"%.)6M-69.3'$"&6 9"%.97(!1)!58*$6O/5"%45 )!58*$6O/5./" )!58*$6O/5>) 90E6M-6 GM*/'*!5+#"VM!,.-$'92S6%SO/5$%<3"!#O'!'(&6<;= 8!,!#P2F!>C-H&"LI1"H!'23490E6+S9,#! 8!,!#P9 #%D+090E6"!#"H!'2&@5M-6+.- B.'"!#M&.'/.>68*$6O/5"%452349"%.+S9,#! 9(:4/2F!>C-M!O-!59 #%D90E6"%45+.- JK45>62348;. "V 1GM""%45234"H!'2&@5"%45 GH16F!+0O'!'(&6<;=90E6M-6234"H!'2&@5M-6M*/+0 (/#;I3 GH1 6$H7$3, 2536)
6/" !"63@'&58!,!#P6F!,!>A->6"!#0Y/5"&6"F! &.G,H5?&M#U(:A B.'()$*!,3"!#6F!#&583,!>A-0#1B'A6="&)+,-./"+,-0#1.&)>6.-!6"!#"F! &.9(H3@'+Z"&)"H-$'+,- (/#6;A, 2545; 8;63'=, 2547)
62
"!#>A-#&583>6"!#0#&)0#;5(&6<;=+,-./"+,-0#1.&)>60#192?+2' 9#%4,,! !""!#234,C!$%2'!H&'9"LM#?!8M#= +.-#&)/6;,&M%>C- &.M&@5C6*$'(H&55!60#,!IUOK@6 >60[ (.?. 2503
B.' .#. /##P 6!S#2##( GH1SI1 +.-2F!"!#$% &'.-!6"!#>A-#&583GH1NHO/5#&583M*/+,-./"+,-0#1.&)CH!'A6%. 9A*6 (;2<#&"L! \-/58!,'*!6 8!$6-/'0#1G0Y5 A)! "H-$'+,- 90E6M-6 B.'+.-,3#!'5!62!5$%A!"!#9"34'$"&)"!#0#&)0#;5(&6<;=(;2<#&"L! >60[ (.?. 2508 9#:4/5 “Radiation-
Induced Somatic Mutations in Plant Breeding” >6"!#0#1A;, The Use of Induced Mutations
in Plant Breeding &.OK@6234"#;5B#, 0#192?/%M!H3 >6#1C$*!5$&6234 25 (]LQ!S,-1 ,%P;6!'6 2507
P&. !"6&@6>60[ (.?. 2518 "V+.-,3"!#M3(%,(=+,-./"+,-0#1.&)/3"S#&@5>60[ (.?. 2518 B.'90E6NH5!6"!#0#&)0#;5(&6<;=\-/58!,'*!6 9#:4/5 “Radiation-Induced Somatic Mutations in
Kalanchoe (Kalanchoe laciniata)” (/#;I3, 2550) !""!##$)#$,5!6$% &'.-!6"!#>A-#&583"&)+,-./"+,-0#1.&)>60#192?+2' 234+.-,3"!#M3(%,(=9N'G(#*#1C$*!50[ (.?. 2510 – 2550 2&@55!6$% &'O/5C6*$'5!6#!A"!#GH19/"A6 >6$!#8!#2!5$%A!"!#GH15!6$% &'234M3(%,(=>6#U0O/5#!'5!6"!#$% &'0#1 F!0[ $%2'!6%(6<= 0^DC!(%9?L 8%45(%,(=M*!5 R ()$*!,3+,-./"+,-0#1.&) #$,2&@5(##I+,-6@F!0#1.&) 2&@58%@6,!"$*! 64 A6%. (species) B.'8*$6>CD*90E6+,-./" F!6$6,!""$*! 44 A6%. 90E6+,-0#1.&) F!6$6,!""$*! 10 A6%. GH190E6(##I+,-6@F!0#1.&) F!6$6 10 A6%. JK45/#;I3 (2550)
+.-#!'5!6+$-$*!,3A6%.O/5+,-./"+,-0#1.&)234C6*$'5!6C#:/);SSHM*!5R 9S'6F!+07!'#&5839(:4/?K"L!$% &'NHO/5#&583M*/"!#90H34'6G0H5H&"LI12!58&IT!6GH1"!#"H!'(&6<;= F!6$6 51 A6%.(:A B.',3#!'5!6(&6<;="H!'GH-$ 12 A6%. +.-G"* (;2<#&"L! 9)D ,!? G(#9J34'5+_- G/Z#%"&6+$B/9HM ,#"MG.5 9""_$' ";CH!)C%6 \-/58!,'*!6 H%HH34 S!#=96A&46 (%2U963' GH1 A$6A,
C6* $ ' 5 !6234 2F ! " ! #?K "L !PK 5 NHO/5 #& 5 83 "& ) +,- . /" +,- 0 # 1.& ), !"234 8; . S: /
,C!$%2'!H&'9"LM#?!8M#= S%.90E6#-/'H1 79 O/5NH5!62!5$%A!"!#234M3(%,(=9N'G(#* 2&@563@96:4/5,! !",3S$!,(#-/,2&@5.-!6);SSH!"# /;0"#I=GH19S#:4/5,:/ #$,PK58%45/F!6$'S$!,81.$">6"!#?K"L!$% &',!"2348;.
8*$6O/5(:A2346F!,!7!'#&583 ,3CH&""!#$*!8*$62346F!,!7!'#&5836&@6 M-/58!,!#PO'!'(&6<;=B.'"!#8#-!5M!'/. M!O-!5 M!(%9?L GH18!,!#P(&a6!90E6"%45>C,* C6*/>C,* C#:/(:AM-6>C,*+.- (8%#6;A, 2546) "!# 16F!8*$6>.O/5(:A,!7!'#&5836&@6 OK@6/'U*"&)$*!(:A6&@6O'!'(&6<;=B.'$%<3>.
(8%#6;A, 2536; Auerbach, 1976) JK45()$*!8*$6>CD*+,-./"+,-0#1.&)2346F!,!7!'#&583>60^ ;)&6 190E68*$62349H3@'5>68Q!(0H/.9A:@/ (9(!19H3@'596:@/9':4/) S%.90E6#-/'H1 63 2&@563@96:4/5 !"8*$62349H3@'5>68Q!(0H/.9A:@/6&@6 ,3O6!.9HV" 2F!>C-81.$">6"!#6F!,!7!'#&583 GH18!,!#P7!'+.-S#&@5H1,!"
R /3"2&@5"!#9(!19H3@'596:@/9':4/6&@6/! ,3"!#"H!'(&6<;=S*/6O-!58U5/'U*)-!5GH-$ 9,:4/>A-#&583>6"!#9C634'$6F!>C-9"%."!#"H!'(&6<;="V 12F!>C-9"%."!#"H!'8U5OK@6+0/3" (/#;I3, 2546) 6/" !"63@"!#7!'#&583>68Q!("!#9(!196:@/9':4/6&@6 1>C-"F!96%.M-6"H!'234,3 F!6$69JHH=6-/'2348;. 9A*6 9JHH=9.3'$"V 12F!>C-+.-M-6>C,* 2349"%.OK@690E6(&6<;="H!'2&@5M-6 GM*C!"6F!9/!8*$6O'!'(&6<;=/:462348!,!#PO'!'(&6<;=+.- 9A*6 HF!M-6 "%45 >) C#:/"!#6F!9/!M-6(:A2&@5M-6,!7!'#&5836&@6 0^ ;)&66%',0W%)&M%6-/'"$*!
63
96:4/5 !"96:@/9':4/0#1"/).-$'9JHH=CH!'9JHH=2F!>C-+,*9"%."!#"H!'(&6<;=2;"8*$6 B.'/! ,32&@5H&"LI1O/596:@/9':4/.&@59.%, GH196:@/9':4/234"H!'(&6<;=06"&6 2F!>C-9"%.0^DC!>6"!#S&.9H:/"
(&6<;="H!' (8%#6;A, 2546 ) JK45"!# 1+.-(&6<;="H!'2&@5M-6"VM-/5,3$%<3"!#)&5S&)>C-96:@/9':4/234"H!'(&6<;=(&a6!90E6M-6>C,*OK@6,! JK45 1';*5'!" GH18%@690H:/596:@/234C!"A%@68*$6C#:/M-6+,-./"+,-0#1.&)6&@6
,3O6!.>CD* #$,PK5"!#O6'-!'"V2F!+.-'!"HF!)!"9,:4/923')"&)"!#9(!19H3@'596:@/9':4/
#&583234>A-"&)+,-./"+,-0#1.&) ()$*!,3"!#>A-#&583G",,!,!"2348;.PK5#-/'H1 99 GH1#&5839/V"J= #-/'H1 1 2&@563@96:4/5 !"#&5832&@5 2 A6%. ,3/F!6! >6"!#21H;21H$58U5 8!,!#PN*!6N*!69O-!+0>696:@/9':4/M!'/. M!O-!5+.- JK45B.'0"M%8*$623490E6M!,&",3"!)>)C#:/,3+O (wax) C;-,/'U* (8%#6;A,
2536) GH1+,*()$*!,3#!'5!6"!#>A-#&583/:469H' 8/.SH-/5"&) Q&2#,!? (2548) 234"H*!$$*!#&5832346%',>A->6"!#0#&)0#;5(&6<;=(:A +.-G"*#&5839/V"J=GH1#&583G",,! JK45,3S;I8,)&M%2!5#&5839C,:/6"&6 M*!5"&6234M-6"F!96%.O/5#&58392*!6&@6 B.'#&583G",,!90E6#&583234PU"6F!,!>A-,!""$*!#&5839/V"J= 96:4/5 !"#&583G",,!,3S$!,'!$SH:46M4F!"$*! K5,3/F!6! >6"!#21H;21H$5N*!6$&MP;+.-8U5"$*!#&5839/V"J= (Casarett,
1968) GM*"V'&5+,*,3"!#':6'&6234G6*6/6$*!#&583M*!5A6%."&6 12F!>C-9"%.NHM*!5"&6 GM* 1+0,3NHM*//&M#!"!#"H!'2349"%.OK@6 (/#;I3, 2530")
$%<3"!#7!'#&583GH10#%,!I#&583234>A- ()$*!8*$6>CD*>A-$%<3"!#7!'#&583G))973')(H&6,!"2348;.PK5#-/'H1 79 2&@563@96:4/5,! !""!#7!'#&583G))973')(H&62F!>C-/&M#!"!#"H!'(&6<;=B.'2&4$+0S*/6O-!58U5 >A-#1'19$H!8&@6 O6!.O/58*$6234"H!' (mutated sector) >CD* 5*!'M*/"!#S&.9/!H&"LI16&@6 R //",!+.- (/#;I3, 2539; (3#6;A, 2546; Q&2#,!?, 2548) 8F!C#&)"!#7!'#&583G))9#:@/#&56&@6 ,&" 1>A-"&)(:A2&@5M-6C#:/8*$6O/5(:A234"F!H&59 #%D9M%)BM 6/" !"63@'&5,3"!#7!'#&583G))CH!' R A*$5/!'; (re-current radiation) B.'2F!"!#7!'#&583J@F!CH!' R A&4$/!'; (generation)
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lethality; LD50) C#:/S*!0#%,!I#&583234,3NH2F!>C-M-6(:A,3"!#9 #%D9M%)BMH.H5 50 90/#=9JV6M= (50 % growth reduction; GR50) (8%#6;A, 2536;
(3#6;A, 2546; IAEA, 1977)
64
6/13=:+%>?2%;=$"!"#8"DE378293:782'%09; ?5)9DE+= F
!""!##$)#$,#!'5!6NH"!#$% &'2349"34'$O-/5"&)"!#>A-#&583"&)+,-./"+,-0#1.&) ()$*!#&583,3NHM*/+,-./"+,-0#1.&) .&563@ NHM*/BS#B,BJ,O/5+,-./"+,-0#1.&)
#&5832F!>C-9"%."!#90H34'6G0H5BS#58#-!5O/5BS#B,BJ,+.-5*!' H&"LI123490H34'6G0H59"%. !""!#234BS#B,BJ,PU"2F!>C-GM"C&"96:4/5 !"#&583 GH-$,3"!#9A:4/,M*/"&6 C#://! 2F!>C-A%@68*$6O!.C!'+0 (deletion) JK45"!#O!.C!'+0O/5BS#B,BJ,2F!>C-'36 F!6$6C6K45O!.C!'+0.-$' 8/.SH-/5"&)"!#?K"L!>6 //6%B2"!H&, 234()$*!#&5832F!>C-9"%."!#90H34'6G0H5O/5BS#B,BJ, 9A*6 2F!>C-9"%."!#O!.C#:/C&")#%9$I satellite ("]LI!, 2545) 8F!C#&)"!#9A:4/,M*/"&6 P-!9A:4/,M*/"&69C,:/69.%,"V+,*,3"!#90H34'6G0H59"%.OK@6"&)8%45,3A3$%M6&@6 R GM*"!#9A:4/,/'*!5N%.0"M% 9A*6 ,3A%@68*$6BS#B,BJ,9(%4,OK@6,! (duplication) "!#9A:4/,G))"H&)2%?2!5 (inversion) C#:/9"%."!#9A:4/,8H&)SU*#1C$*!5BS#B,BJ,M*!5SU*"&6 (translocation) 190E6NH>C-8%45,3A3$%M6&@6 R 9"%."!#90H34'6G0H5+.- (8%#%6;A,
2536) 8/.SH-/5"&)#!'5!6>6./5.K5 234()$*!#&583,3NH2F!>C-BS#B,BJ,9"%."!# &)SU*"&6N%.0"M% B.' F!6$69JHH=234()S$!,N%.0"M%O/5BS#B,BJ, 19(%4,OK@6M!,0#%,!I#&5832349(%4,OK@6 ((#2%('=, 2542)
B.'NHO/5#&583234,3M*/BS#B,BJ, 1,!"C#:/6-/'OK@6/'U*"&)0#%,!I#&583234+.-#&) 8/.SH-/5"&)#!'5!6O/56&626! (2525) ()$*!(;2<#&"L!234+.-#&)#&583 12F!>C-9"%.S$!,N%.0"M%O/5BS#B,BJ, B.'90/#=9JV6M=BS#B,BJ,234N%.0"M%,38C8&,(&6<="&)0#%,!I#&583>69A%598-6M#5 6/" !"63@'&5()$*!#&583,3NHM*/ F!6$6O/5BS#B,BJ, B.'"!#?K"L!"&)9)D ,!? ()$*!#&583,3NHM*/ F!6$6O/5BS#B,BJ,B.'()9(3'5 1 >6 4 O/5M-62347!'#&583 B.',3 F!6$6BS#B,BJ,92*!"&) 57 !"0"M%234,3BS#B,BJ,
54 9A*69.3'$"&)>6)&$CH$5 234()$*!9.%,)&$CH$5,3BS#B,BJ,2340H!'#!" 2n=16 GM*9,:4/+.-#&)#&5832F!>C-,3 F!6$6BS#B,BJ,9(%4,OK@6/3" 2 "H;*, S:/ 2n=18 !""!#7!'#&583G",,! 30 9"#'= GH1#&5839/V"J= 40 9"#'= GH1 F!6$6BS#B,BJ, 2n=20 !""!#7!'#&583G",,! 40 9"#'= 8F!C#&)(##I+,-6@F!0#1.&)"V()$*!#&583,3NHM*/BS#B,BJ,9A*69.3'$"&)+,-./"./"+,-0#1.&)/:46 R 9A*6 /9,J/6 (/;+#, 2542)
NHM*/H&"LI12!58&IT!6O/5+,-./"+,-0#1.&)
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
65
NHM*/"!#90H34'6G0H583O/5+,-./"+,-0#1.&)
#&583,3NH2F!>C-9"%."!#90H34'6G0H583O/5+,-./"+,-0#1.&) B.'97(!183O/5>)GH183./"
9"%.0#!"W"!#I=2349#3'"$*! mosaic 8*$6>CD*9"%.OK@696:4/5 !""!#"#1 !'M&$O/5SH/#=B#ZbHH=+,*8,4F!98,/ 6/" !"63@/! ()$*!>)90H34'6 !"839O3'$+090E683/:46 R 9A*6 839CH:/5 C#:/83O!$ 90E6M-6
(/#;I3, 2530") B.'M&$/'*!5O/5+,-./"+,-0#1.&)234+.-#&)NH !"#&5832F!>C-9"%."!#90H34'6G0H5O/583./" GH183>) +.-G"* (;2<#&"L! ( 5 %6M=, 2520) S!#=96A&46 (A&'A;,(H, 2526) 9)D ,!? (A;M%62#, 2532) ";CH!) ((&A#%62#=, 2538) .!CH! ($%2'!";H, 2543) 9?#LT3"-!62/5 (#&A63'=, 2544) (%2U963'
((##I3, 2543; Q&2#,!?, 2548) G(#9J34'5+_- (/#;I3 GH1 6$H7$3, 2534; Wongpiyasatid and
Hormchan, 2000) )&$ 5"H63 (Puripunyavanich and Boonsirichai, 2005) 90E6M-6 B.'"!#90H34'6G0H583./"63@/! 90E6"!#90H34'6G0H583./"2&@5./" C#:/90H34'6G0H597(!1)!58*$6"V+.- (/#;I3 GH1 6$H7$3, 2534) 8*$6>6>)8*$6>CD*,&" 1()H&"LI1>).*!5 9A*6 "H-$'#&M"&2H3 ($6%.!, 2547) +2#'-/'>)GCH,GH1+2#'-/'>)2U* (8;A!.!, 2550) 90E6M-6 JK45H&"LI1"!#.*!5O/5>)8!,!#P9C634'$6F!>C-9"%.+.-98,/ R B.'"!#7!'#&583 (/#;I3, 2550)
NHM*/"!#9 #%D9M%)BMO/5+,-./"+,-0#1.&)
NHO/5#&583234,3M*/+,-./"+,-0#1.&)6&@6 ()$*!#&583,3NH2&@5"!#9#*5GH1'&)'&@5"!#9 #%D9M%)BM GM*,&"()$*!#&583,&" 18*5NHM*/"!#'&)'&@5"!#9 #%D9M%)BM,!""$*!
- "!#9#*5"!#9 #%D9M%)BM
8F!C#&)NHO/5#&583234,3M*/"!#9#*5"!#9 #%D9M%)BMO/5+,-./"+,-0#1.&)6&@6
9/,/# (2530) ()$*!)&$CH$5 234N*!6"!#7!'#&583 0#%,!I 30 9"#'= ,3"!#9 #%D9M%)BM234.3"$*!"!#+.-#&)#&583#1.&)/:46 JK45>C-NH9A*69.3'$"&)?%#%?&".%c (2542) 2342F!"!#?K"L!>6)&$CH$5 (&6<;=8&MM);M'= ()$*!M-6234+.-#&)#&583G",,*!GH1#&5839/V"J= 10-20 9"#'= ,3"!#9 #%D9M%)BM234.3"$*!M-60"M% ,3#!"GO65'!$>CD* GH1,3#!"d/' F!6$6,!" +CH,3S$!,8,)U#I= GH1>C-M-6>C,*234,!""$*! >6"H-$'+,-9(A#CK5 2346F!B0#BMS/#=,234+.- !""!#9(!19,HV.,!7!'#&583 ()$*!M-6234N*!6"!#7!'#&583 234#1.&)#&583 60 9"#'= ,3O6!.O/5M-6>CD*"$*!M-6234+,*+.-#&)#&583GH1#/.M!'2&@5C,. 6/" !"63@(##I+,-6@F!0#1.&)"V()$*!#&583,3NHM*/"!#9#*5"!#9 #%D9M%)BM9A*69.3'$"&6 9A*6 BH)39H3' 234N*!6"!#7!'#&583 G))9#:@/#&5 33.51 9"#'= ()$*!M-6"H-!,3"!#9 #%D9M%)BM,!"9,:4/923')"&) control
- "!#'&)'&@5"!#9 #%D9M%)BM
NHO/5#&583234,3M*/"!#'&)'&@5"!#9 #%D9M%)BMO/5+,-./"+,-0#1.&) 8!,!#P()+.-2&4$+0"&)"!#?K"L!>6+,-./"+,-0#1.&)2;"A6%.234N*!6"!#7!'#&583 9A*6 "H-$'+,- (<61, 2548)
";CH!)C%6 (8;#$%A, 2526) O%5G.5 (9)D ,!?, 2541) S!#=96A&46 (A&'A;,(H, 2526; /#.3 GH1 A&'A;,(H, 2527) J*/6"H%46+2' (/.%?#, 2535) ./5.K5 ((#2%('=, 2542) 02;,,! (/.%?#, 2533;
9S#:/$&H'=, 2541; $&6$%8!, 2545; $%AD!, 2546) $*!68342%? (/M%6&62=, 2545) //6%B2"!H&, ("]LI!, 2545) /6U9)3'8 (.!$8$##S=, 2545) /9,J/6 (/;+#, 2542) 90E6M-6 B.'H&"LI1234(:AG8.5//">C-9CV6$*!PU"'&@)'&@5C#:/A1H/"!#9 #%D9M%)BM S:/ 19 #%D9M%)BM234A-!"$*!M-60"M%234+,*+.-#&)#&583
66
C#:/9"%.H&"LI1GS#1G"#6 JK4590E6NH96:4/5,! !"#&583,3NH'&)'&@5C#:/A1H/"!#G)*59JHH= JK4590E60#!"W"!#I=2349"%.OK@6)*/'2348;. (/#;I3, 2530) B.'C!"+.-#&)#&5830#%,!I6-/' R 12F!>C-9"%.S$!,H*!A-!>6"!#G)*5M&$O/59JHH= (mitotic delay) JK4590E6G))A&4$S#!$ CH&5 !"6&@6"!#G)*5M&$ 1"H&),!90E60"M% GM*>6"#I3234+.-#&)#&5830#%,!I234,!"OK@6"V 1,3NHM*/"!#'&)'&@5"!#9 #%D9M%)BM,!"OK@6
JK45/! 8*5NH2F!>C-(:AM!'+.->62348;. 2&@563@96:4/5 !"#&583>60#%,!I8U5 2F!>C-9JHH=C';.A15&""!#G)*5M&$/'*!5P!$# GH1+,*,3"!#Zef6S:6"H&),!2F!C6-!234+.-/'*!59.%, >62348;.9JHH= 1M!' 9JHH=/:46234+,*M!'/! G)*5M&$9(%4, F!6$69O-!,!G262349JHH=234M!'+.- >6)!5"#I3()$*!9JHH=234N*!6"!#7!'#&5838!,!#PG)*5M&$M!,0"M%+.-CH!'S#&@5CH&5 !"6&@69JHH=PK5 1M!' (8%#6;A, 2527; /#;I3, 2530")
NHM*/9,HV.+,-./"+,-0#1.&)
#&583,3NHM*/"!#5/"GH190/#=9JV6M=S$!,/'U*#/.O/5+,-./"+,-0#1.&) B.' !"#!'5!6>6+,-./"+,-0#1.&)CH!'A6%. +.-G"* 9?#LT3"-!62/5 (#&A63'=, 2544) 2!6M1$&6 (?%#%#&M6=, 2546) )!6A:46C6U ((%A%M, 2546) GH1)&$CH$5(&6<;=);Ig#%" (Lama et al., 2005) H-$6>C-NH90E6+0>62%?2!59.3'$"&6 S:/ "!#5/"O/59,HV.GH190/#=9JV6M=S$!,/'U*#/.O/5+,-./"+,-0#1.&)H.H59,:4/+.-#&)#&583>60#%,!I2348U5OK@6 #$,PK5"!#()H&"LI1N%.0"M%O/58*$6M*!5 R O/5(:A 6/" !"63@'&5()$*!"!#7!'#&583G))9#:@/#&5 /&M#! 627.5 9"#'= >C-G"*9,HV."H-$'+,-A-!5S*/,2349H3@'5>68Q!(0H/.9A:@/ ,3"!#(&a6!90E6M-6"H-!O6!.>CD* S%.90E6 55.93 90/#=9JV6M= (<&6$=, 2546)
NHM*/#1))"!#8:)(&6<;=O/5+,-./"+,-0#1.&)
#&583,3NHM*/"!#8:)(&6<;=O/5+,-./"+,-0#1.&) B.',3#!'5!6>6+,-./"+,-0#1.&)CH!'A6%.
+.-G"* >6C6-!$&$ ()$*! 234#1.&)#&583 12 9"#'= OK@6+0 ,3NHM*/"!#90E6C,&6O/5C6-!$&$ ();D,3, 2518)
>6G(#9J34'5+_- ()$*!#&5832F!>C-9"%."!#90H34'6G0H5H&"LI19"8#9(?NU-GH19(?9,3' 9A*6 )!5./"+,*,39"8#9(?NU-GH19(?9,3' )!5./",397(!19"8#9(?>.9(?C6K45 (/#;I3 GH1 6$H7$3, 2534;
Wongpiyasatid and Hormchan, 2000) 9A*69.3'$"&)"!#?K"L!O/5(#2%('= (2542) >6./5.K5 ()$*! #&5830#%,!I 2.5 9"#'= OK@6+0 ,3NH2F!>C-S$!,8,)U#I=GH190/#=9JV6M=S$!,5/"O/59#IUH.H5 8/.SH-/5"&)9?#LT$&A# (2536) 234"H*!$$*! "!#A&"6F!>C-9"%."!#"H!'(&6<;=.-$'"!#>A-#&583C!">A-0#%,!I#&5832348U59"%6+0 12F!>C-9"%."!#"H!',!" 6+,*8!,!#PP*!'2/.+08U*#;*6M*/ R +0+.- 96:4/5 !""!#+,*9 #%D(&6<;= (infertility) "!#M!'O/59JHH= 96:@/9':4/ C#:/(:A2&@5M-6 "!#"H!'>6#1.&)BS#B,BJ,983'C!',!" 62F!>C-+,*8!,!#PP*!'2/.+.- 96:4/5 !"9"%.S$!,N%.0"M%OI1G)*56%$9SH3'8G))+,B/J38 90E6NH>C-9JHH=8:)(&6<;=+,*9 #%D(&6<;=C#:/M!'+0 9A*69.3'$"&) (3#6;A (2546)
234"H*!$$*!"!#7!'#&583>C-G"*(:A>60#%,!I8U5 B.'97(!1"!#7!'G))973')(H&6 1"*/>C-9"%.S$!,983'C!'2!58#3#1O/5(:AS*/6O-!5,!" GH1 1,3S$!,N%.0"M%O/5BS#B,BJ,8U5,!" 90E6NH>C-H.S$!,8,)U#I=9(?H5+0
67
NHM*/$&W &"#A3$%MO/5+,-./"+,-0#1.&)
#&583G",,!,3NHM*/$&W &"#A3$%MO/5+,-./"+,-0#1.&) B.',3NHM*/"!#':.C#:/H.#1'19$H!>6A*$5(&a6!"!#M*!5O/5(:A 9A*6 #1'19$H!//"./" #1'19$H!"!#)!6O/5./" #1'19$H!9"V)9"34'$ XHX >6"!#?K"L!"&)G"H.%B/H&8 ()$*!#&583G",,!,3NH2F!>C-G"H.%B/H&8//"./"9#V$OK@6
"!#)!6O/5./"9#V$OK@6 2F!>C-#*6/!';"!#9"V)9"34'$H5 (/#$##I, 2524, 2525; 0#!6/, GH1SI1, 2525) >6";CH!) ()$*!M-6234+.-#&)#&5832F!>C- F!6$6$&6234>A->6"!#//"./"90H34'6G0H5+0
((&A#%62#=, 2538) GH1"!#?K"L!"&))&$CH$5/9,#%"&6839CH:/5 JK450"M% 1+,*8!,!#P//"./"+.->60#192?+2' GM*>6M-6234N*!6"!#7!'#&583()9(3'5 1 M-6 2348!,!#P//"./"+.- ($%A&' GH1 "6"(#, 2549)
NHO/5#&583M*/H&"LI12!58#3#$%2'!CH&5"!#9"V)9"34'$O/5+,-./"+,-0#1.&)
!""!#?K"L!"!#>A-#&5839(:4/0Y/5"&6"F! &.9(H3@'+Z"&)"H-$'+,-Q!'CH&5"!#9"V)9"34'$ ()$*!#&583,3NHM*/"!#H./!';CH&5"!#9"V)9"34'$O/5"H-$'+,- B.'"!#7!'#&583 >6A*$50#%,!I
1.5-2 "%BH9"#'= "*/>C-9"%./!"!#CH;.#*$5O/5./"/'*!5#$.9#V$ 8*5NH>C-/!';"!#0^"G "&6H.H5 (/#6;A, 2545; 8;63'=, 2547) JK45M*!5"&)#!'5!6O/5 A;M%62# (2531) 234?K"L!"&)9)D ,!? ()$*!"!#>A-#&583 0#%,!I 5 GH1 10 9"#'= ,3G6$B6-,2F!>C-9)D ,!?,3/!';"!#0^"G "&66!6OK@6 2&@563@/! 96:4/5,! !"0#%,!I#&583234>A- ,3S$!,GM"M*!5"&6,!" K52F!>C-NH234//",!GM"M*!5"&6 B.'0#%,!I#&5832348U5,!" 1"*/>C-9"%.S$!,983'C!'234#;6G#5G"*(:A+.-,!""$*!0#%,!I#&583234M4F! (/#;I3, 2530)
78293:782'%09; ./0@%%,7825GH+'%09; @;5B&->I8E!"#792*+::+%?;:5H+>I2:/+<@;5B&-J9<>?2%;=$" ./01239"-1234$"< 13=:+%>?2%;=$":; 78293:782'%09; ./0@%%,7825GH+'%09;
"!#7!'#&583>C-G"*+,-./"+,-0#1.&)GH1(##I+,-6@F!0#1.&) 9(:4/A&"6F!>C-9"%."!#"H!'(&6<;=90E6(&6<;=>C,* 234,3S$!,8$'5!, S$!,G0H" ,3H&"LI1234.32!5"!#MH!. ()$*!0#192?+2',3 +,-./"+,-0#1.&)(&6<;=>C,* R !""!#0#&)0#;5(&6<;=B.'"!#>A-#&5832340#18)S$!,8F!9#V GH-$>6(:A 3
A6%. +.-G"* (;2<#&"L! G(#9J34'5+_- GH19)D ,!? F!6$6 22, 10 GH1 6 (&6<;= M!,HF!.&)
6/" !"63@'&5,3(##I+,-6@F!0#1.&)(&6<;=>C,*/3"CH!'A6%.234+.- !""!#7!'#&583 GH1(&6<;=+,-./"
+,-0#1.&)>C,* R 9CH*!63@+.- F!C6*!' *!'G " 6PK59"LM#"#6F!+0O'!'(&6<;=9(%4, F!6$6 9(:4/2F!90E6"!#S-!GH18*5//"GH-$ K590E68#-!5#!'+.->C-G"*9"LM#"#GH10#192?A!M%>C-9(%4,OK@6 GH1H."!#6F!9O-!+,-./"+,-0#1.&) !"M*!50#192? 6/" !"63@>60^ ;)&6'&5,3+,-./"+,-0#1.&)/3"CH!'A6%.
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
68
97H34'GH-$>A-9$H!>6"!#0#&)0#;5(&6<;=9(:4/>C-+.-(&6<;=>C,* 9(3'5 1.2 0[ 92*!6&@6 JK45C!">A-$%<3"!#0#&)0#;5(&6<;=G))0"M%"$*! 1+.-(&6<;=>C,* R /! >A-#1'19$H! 5-10 0[ C#:/6!6"$*!63@ JK45"!#>A-#&583A&"6F!>C-9"%."!#"H!'(&6<;=90E6$%<3"!#C6K45234 1A*$'8*598#%,"!#0#&)0#;5(&6<;=(:AG))9.%,234,3/'U*GH-$M!,0"M% 2&@563@9(:4/9(%4,/&M#!"!#"H!'(&6<;=2349"%.OK@6/'U*GH-$M!,<##,A!M%>C-8U5'%45OK@6 (/#;I3 , 2530)
>60^ ;)&6+.-,3C6*$'5!6234/)#,GH1)#%"!#7!'#&583>C-G"*(:AO/5C6*$'5!6#!A"!#M*!5 R 234,39S#:4/57!'#&583 2F!>C-0#1A!A6 C#:/9"LM#"#2&4$+08!,!#P9O-!PK592SB6BH'363@+.-5*!'OK@6 B.',3NU-9A34'$A!DGH1AF!6!D"!#97(!1.-!6S/'>C-SF!0#K"L! G616F! .UGH GH1S$)S;, /'*!5>"H-A%.
#$,PK5,3"!#>A-92SB6BH'32342&68,&' GH1,3S$!,0H/.Q&',!"'%45OK@6 2F!>C-H.0^DC!>69#:4/5O/5/&6M#!'2349"%.OK@6 !""!#>A-#&58390E6/'*!5,!"
1234$53.50
1. S$#,3"!#6F!(&6<;="H!' !""!#7!'#&5832340#18)S$!,8F!9#V ,!8*598#%,>69A%5(!I%A'=>C-90E6234#U- &",!"OK@6 9(:4/8*598#%,>C-9"%."!#>A-0#1B'A6= !"#&583"&)+,-./"+,-0#1.&),!"OK@6
2. !""!##$)#$,NH5!6$% &' ()$*!"!#?K"L!"!#>A-#&583"&)+,-./"+,-0#1.&)>60#192?+2'6&@6'&5/'U*>6$5 F!"&. JK45'&5,3+,-./"+,-0#1.&)/3"CH!'A6%.234S$##3)9#*5.F!96%6"!#?K"L!M*/+0
3. "!#>A-#&583"&)+,-./"+,-0#1.&)>60#192?+2'6&@6 ()$*!,3"!#>A-0#1B'A6= !"#&5839(3'5 2 A6%.92*!6&@6 B.'97(!1#&583G",,!,3"!#>A-,!"2348;. JK45S$#8*598#%,GH1C!G6$2!5>C-,3"!#>A-0#1BA6= !"#&583/:46 R >C-,!"OK@6 9A*6 #&5836%$M#/6 GH1+///6)3, JK45,3#!'5!6$*!8!,!#PA&"6F!>C-9"%."!#"H!'(&6<;=+.-,!" GM*'&5,3#!S!G(5 GH1>A-"&)M&$/'*!5(:A+.-6-/'
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
JK45M&$9"LM#"#9/590E6);SSH8F!S&D 234 1M-/52F!"!#)&62K"O-/,UH23490E60#1B'A6=9CH*!63@+$-9,:4/6F!(:A+0#&))#%"!#7!'#&583 9(:4/0#1B'A6=O/58*$6#$,GH1M69/5
69
!!"#$%&!'
!"#$ %&'&(. 2545. &#!()&$*#+,-.&/0&#!&1#2+$ Ornithogalum dubium Houtt. 302&#!4#2!)5678&''#!9:'&) &#!.;#<.17=25.$>=?.2>@?. )&*+$'&%',-./&00$1*,
23$)&*+$45+6 "7/8$97/-. 4:;2<$')&(5+%//#=2>'?@$. 2550. ;!!"A'-$=*#B/C:. A34;<*BC2$:
http://www.fisheries.go.th/freshwater/technical/doc_data/paper/panmino.doc, 20
/ D$E2 2550.
6 8 %F46(/&0. 2544. D1E?5!)5678&''#F7@'7G9?&#!.;#<.17=25.$>=?.2>@?E/5805. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
E//H&7 ,//28&/&. 2538. I4JK</5<9BA 22$7;K1./17EK/-2 4>)+=2> 4:;2AE*4B+$, '. 81-86. !
!#25#$&#!H!<(%':/(#&#!A'-0?&A'-H!<0) 8,95(#G/I!)=5F7@ 1. E#L //2 $/./L9$'<$')&(5+A4L%5M'$=2>NK =2>./LN5O 9@$'5 <$'E#L //2 $/)&(5+A3;<H$7& A4L92$E2%PH9)'A3;<./L6*8=*+, /:<6*%Q.
6E/PK)54+- 6 B+/7&)&)5M' F4. 2541. D1E?5!)5678&''#G9?&#!.;#<.17=25.$>=?.2>@?E?5G-$
HF%''#. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. 1E/< $/(5N $/ $ 5225'7/5<9B (6/B+O6/B+<). 2547. !)56781<&#&&)'')$G!)567. %&2%-E/5?<*BC 4.
1E/< $/(5N $/ $ 5225'7/5<9B 9@$'5 <$'./2$#F6%PCK95'7& /L*/)<)&*+$8$97/-A4L6*E1'14+B, /:<6*% Q.
(<(&'7- A.4 ./L%5',-. 2520. &#!JK&L#1)&L"<&!!';)$M%N #5H!<&#!81<D1E?5!)5678&''#F7@'7G9?&#!.H17@2$8H15E?53I!3'3O'81<1)&L"<P#2$?&E?5;>(6&%1;%FM!)&L#. )&*+$'&%',-./&00$1*, (:R$4< /#-23$)&*+$45+.
(&/L'5'*- ':7+L9 :4. 2547. D1E?5!)5678&''#G9?&#!.B!/Q.G/ 3G81<1)&L"<F#56)"R#$E?5 Kalanchoe sp. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
(&/$S/#- %5',:-(5'*/-. 2545. &#!.,$7@2:$*#+,-.&/0&#!&1#2;)$M%N+$0#,1#0?&67E#:302:/M7&#!.;#<.17=25.$>=?.2>@?!9:'&) &#!4#2!)5678&''#. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
H5+H:2%4 9:/&+L85 N&T. 2526. D1E?5!)5678&''#G9?&#!.;#<.17=25H1#22?0I#!N.$()$;)$M%NA:FNOK'. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
H:7&'*/ OF/#L '&"U. 2531. D1E?5!)5678&''#F7@'7G9?. QB'#J+$6P#;H1?0.(>=?. .G03$%&68"./&00$1*, 23$)&*+$45+6 "7/8$97/-.
70
H:7&'*/ OF/#L '&"U. 2532. &#!()&$*#+,-. QB'#J&1#2;)$M%N302+(-.FI$/I&#!.;#<.17=25.$>=?.2>@?!9:'&) &#!4#2!)567. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
N$)9)//E- (5'*/S5 NB)<8-. 2545. &#!E2#2;)$M%N302:/M7&#!.;#<.17=25.$>=?.2>@? 81<D1E?5!)5678&''#G9?2?0E?5 Anubias barteri var. nana F7@.;#<.17=25+$,1?08&-:.
)&*+$'&%',-./&00$1*, 23$)&*+$45+9<J4$'E/&'*/-. =7/62, 95<J-9:)//#-. 2547. D1E?5&#!4#2!)5678&''#8 3I!$/&!9:'&) &#!.;#<.17=25
.$>=?.2>@?E?5 Lobelia cardinalis. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. ,'L :42:7&)5M'-. 2548. D1E?5!)5678&''#G9?&#!.B!/Q.G/ 3GE?5&1-:2A'-6&%1,:#2
1S&D6' #5($/0+$6P#;H1?0.(>=?. )&*+$'&%',-./&00$1*,
23$)&*+$45+6 "7/8$97/-. ,'$,&. 6%/&8%/$+)<8-. 2547. &#!.,$7@2:$*#+,-.&/0&#!&1#2;)$M%NG9?G-$0#:&!<B#2
(Hygrophila difformis) +$6P#;H1?0.(>=?302+(-!)5678&''#. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
,5')- J@$*K<. 2546. D1E?5!)5678&''#G9?&#!5?&E?5.'1T081<&#!.B!/Q.G/ 3GE?5&1-:2A'-+$6P#;H1?0.(>=?. .G03$%&68"./&00$1*, 23$)&*+$45+6 "7/8$97/-.
'<45 "#- 6*B+'69/B. 2541. &#!.;#<.17=25.$>=?.2>@? 73&.$72.!T&ON81<D1E?5!)5678&''#. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
'5'*'$ 4&C29 :4. 2525. D1E?5!)5678&''#F7@'7G9?;%FM!)&L#1S&D6'. )&*+$'&%',-./&00$1*,
(:R$4< /#-23$)&*+$45+.
O:02B 64&8/57'-6NH$ :4. 2518. &#!JK&L#?/FM/;1E?5!)5678&''9#F7@'7G9?&#!.H17@2$8H151)&L"<&#!.B!/Q.G/ 3GE?5,$-#:):;)$M%NB)&!;!!0/U. )&*+$'&%',-./&00$1*,
23$)&*+$45+6 "7/8$97/-. 6O0(2$8 =%1/('-. 2541. D1E?5!)567G9?&#!.B!/Q.G/ 3G81<1)&L"<[email protected]@2$8H15+$E/5
805. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. . /#- 75?<.VK<. 2548. D1E?5!)5678&''#!9:'&) &#!.;#<.17=25.$>=?.2>@?F7@'7G9?0#,1#.
.G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. ./$'K2 %!M%<"-, A9<,//2 E2 !9, K/)//# 2F4*K<($N A4L K<K$( 3$0H$064&8.
2525. D1E?5!)5678&''#F7@'7G9?1)&L"< #5H!<&#!E?58&10/3?1)6;)$M%N “Spic and
Span 81<;)$M%N Norwich Canary”. /$+<$'I4 $/)&(5+ ./L(@$.W 2525. 9X$O5')&(5+A4L%5M'$A3;<23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
.G00$ ,B/L)&*+$64&8. 2546. E)$2/F>%P?'U$' $/)&(5+, '. 8-16. ! S$E)&H$.U%B)&*+$. 2546.
IS9'>?$)&HR;7:/F2# .?&6#!H!<&? &#!VW&? !'$/6/G.!>@?5 &#!:/B)2. S$E)&H$.U%B)&*+$ E#L6 "7/ @$A%<A9' 23$)&*+$45+6 "7/8$97/-, 'E/.U2.
71
.YZ'K:O4 *58'$':77&+L. 2548. &#!H!) H!%5;)$M%NA'-$=*# Cryptocoryne wendtii “green” 302:/M7&#!.;#<.17=25.$>=?.2>@?!9:'&) &#!4#2!)5678&''#8 .472 ;1)$. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
%/*&%+- 6*&NO$/2B. 2542. D1E?5!)5678&''#G9?1)&L"< #5H!<&#!E?50?50K5 Gloriosa
superba L.. .G03$%&68"./&00$7/B, (:R$4< /#-23$)&*+$45+.
%//#B 8/B9)59N&T. 2543. &#!JK&L#?/FM/;1E?5!)5678&''#G9?&#!.H17@2$8H151)&L"<&#!.B!/Q.G/ 3GE?5;/FS.$72;)$M%N Pearl Wave. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
%5H/&'*/- O:06(&2. 2538. &#!E2#2;)$M%N&%,1# +$6P#;H1?0.(>=?81<&#!F*#+,-&1#2;)$M%N302&#!+(-!)5678&''#. )&*+$'&%',-./&00$1*, 23$)&*+$45+6HB+<[32;.
%5',-)NB */5%+-8&/&. 2546. D1E?5!)5678&''#F7@'7G9?&#!.B!/Q.G/ 3GE?53;M/U )11)5&N+$6P#;H1?0.(>=?. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
%&H&7 9$E/. 2546. D1E?5!)5678&''#G9?&#!.B!/Q.G/ 3G81<1)&L"<F#56)"R#$E?5 #$(>@$,$S. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
%B/':H (K2%: , 9&/':H 4$28/B(5'*/- A4L 9:/&'*/- NB9B.$'. 2544. $/63'BC+)'@$[3>6 &N $/ 4$+%5',:-['6O0(2$8 N>)+/5<9BA 22$/;)2 5O6*E'&E $/6%$L64B?+<6'P?K6+PCK, '. 15-24. ! &#!H!<(%':/F2#&#!:/F2#J#6G!N 81<.FI3$3127$/:.I172!N I!)=5F7@ 8. 20-21 2&X:'$+'
2544. (5N1N+9@$'5 <$'./2$#F6%PCK95'7& /L*/)<)&*+$8$97/-6*E1'14+BA4L9&C<A)N4>K2,
/:<6*% Q.
2546. $/63'BC+)'@$[3>6 &N $/ 4$+%5',:-1N+ $/\$+/5<9B/;)2 5O $/6%$L64B?+<6'P?K6+PCK,
'. 25-45. ! 8F'+-O/& $/\$+/5<9BA 22$A4L)&(5+'&)6E4B+/-6*E1'14+B. 2546. .FI$/I&#!6!-#5;)$M%NA'-0?&A'-H!<0) 0-:2!)5678&''#. 6K 9$/./L KO $/]^ KO/21E/< $/)&(5+6H&<X;$+*KN6*E1'14+B $/%5M'$%5',P=2>NK =2>./LN5ON>)+/5<9BA 22$9F;6 "7/ /. )5'*BC 26-27 2&X:'$+' 2546 # 8F'+-O/& $/\$+/5<9BA 22$A4L)&(5+'&)6E4B+/-6*E1'14+B 23$)&*+$45+6 "7/8$97/-, /:<6*%.
S5*/2$8 %$'%:;2. 2548. &#!4#2!)5678&''#.;>@?.,$7@2:$*#+,-.&/0&#!&1#2;)$M%N+$;/FS.$72+ 09#5F7@E2#2;)$M%N302:/M7HX&(*#. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
2544& $ 8/B9:,//2. 2542. &#!.,$7@2:$*#+,-.&/0&#!&1#2;)$M%N+$. QB'#J0-:2!)5678&''#!9:'&) .FI$/I&#!.;#<.17=25.$>=?.2>@?. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
/</K< )&68"9:)//#. 2528. &#!()&$*#+,-.2? 7!#&1#2;)$M%N+$,1?0F01?5. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
72
/5 H' 1E17, 8&/&%/ 8/BS&010)#&H+-, 9',&H5+ (5'*/-6./2 A4L 69/&28&/& (5'*/-6./2. 2549.
$/H5 '@$[3>6 &NE)$2A././)'['45 "#L*$<95#U$')&*+$JK<I5 6._N 1N+ $/[H>/5<9BA 22$/;)2 5O $/6%$L64B?+<6'P?K6+PCK. :/F2#6#!&*#8;586$ 4 (1): 34-40.
/5H'B+- +$<=,9<. 2544. &#!()&$*#.J!LR7&-#$F?5+,-&1#2;)$M%N302:/M74#2!)5678&''#. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
)'&N$ N)< ><A9'. 2547. &#!JK&L#6SG!?#,#!F7@.,'#<6'81<D1E?5!)5678&''#F7@'7G9?1)&L"<F#56)"R#$:/F2#E?5&1-:2!)G&)F17. )&*+$'&%',-./&00$1*,
23$)&*+$45+6 "7/8$97/-. )/%'&7 A%N&"U-. 2545. &#!.,$7@2:$*#+,-.&/0&#!&1#2;)$M%N+$0#,1#302+(-!)5678&''#
!9:'&) &#!.;#<.17=25.$>=?.2>@?. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. )1/S$9 :48&/&. 2544. D1E?5!)5678&''#G9?&#!.B!/Q.G/ 3G81<D1/G,):29?2E?51/117@;)$M%N
Red Night 81<;)$M%N Apeldoorn +$6P#;H1?0.(>=?. 9522'$./&00$7/B S$E)&H$/5<9B./L+: 7-A4L=K1`1*. E#L)&*+$8$97/-, 23$)&*+$45+6 "7/8$97/-.
)5'6\4&2 8&/&/57'-. 2546. D1E?5!)5678&''#G9?&#!;)Y$#E?5&/@581<+ &%,1# ,/$. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
)5')&9$ %<"-6%B+/. 2545. D1E?5!)5678&''#G9?&#!.;#<.17=25.$>=?.2>@?HF%''#. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
)&HH:7$ /:;<6/PK<. 2537. D1E?53I(/O/$81<!)5678&''# F7@'7G9?&#!&1#2E?5,$-#:): ;)$M%N ‘Double Spathe’ [email protected]=25+$6P#;H1?0.(>=?. )&*+$'&%',-./&00$1*,
23$)&*+$45+6 "7/8$97/-. )&H0$ 6+$)6/86X &< &(. 2546. D1E?5!)5678&''#G9?&#!.;#<.17=25.$>=?.2>@?HF%''#. .G03$
%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. )&H5+ SF/&.G00)$'&H A4L ' %/ O:08&/&H5+. 2549. $/./5O./:<%5',:-O5)34)<9B634PK<1N+/5<9B
A 22$, '. 439 - 444. ! &#!H!<(%'F#5:/(#&#!E?5',#:/F2#1)2.&LG!J#6G!N I!)=5F7@ 44 6#E#;>(. 30 2 /$E2 – 2 :2S$%5',- 2549. 23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
)&*+$ :4 %/22&. 2543. &#!H!) H!%5;)$M%N0#,1#302 :/M7&#!.;#<.17=25.$>=?.2>@?!9:'&) &#!4#2!)5678&''#. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
8/$+:*, J5NE@$. 2547. D1E?5&#!4#2!)5678&''#8 .472 ;1)$G9? Lobelia cardinalis
+$6P#;H1?0.(>=?. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. 8&/&/57'- )H&/(&/$ /. 2546. D1E?5!)5678&''#F7@'7G9?.'1T0;)$M%NF#$G<:)$. .G03$%&68"
./&00$7/B, 23$)&*+$45+6 "7/8$97/-.
73
8&/&85 N&T 9:'*/+$7/. 2542. D1E?5!)567G9?&#!&1#2;)$M%NE?5 ):,1:5;)$M%N6)GG %[email protected]=25+$6P#;H1?0.(>=?. )&*+$'&%',-./&00$1*, 9X$O5'6*E1'14+B%/L(K26 4>$6(>$E:#*3$/4$N /LO5<.
A4L )&H5+ SF/&.G00)$'&H. 2543. D1E?5!)5678&''#G9?&#!&1#2;)$M%NE?5 ):,1:5;)$M%N %"Z!/&. /$+<$' $/)&(5+. 23$)&*+$45+23&N4 8F'+-8$4$+$, 'E/.U2.
8F'+-O/& $/\$+/5<9BA 22$A4L)&(5+'&)6E4B+/-6*E1'14+B. 2546. ;>(;)$M%N&1#2+$H!<.FJAF2.
A34;<*BC2$: http://www.rdi.ku.ac.th/gamma-irradiation/database/Mutation.htm, 20
/ D$E2 2550.
68/"U)5H/ \C@$8$97/-. 2536. /5<9B2&)67H5C' 5O $/./5O./:<%5',:-%PH. :#!6#!:/F2#J#6G!N S!;#
1(1): 35-46.
9$/&'B )$1+O:7/. 2548. D1E?5&#!4#2!)5678&''#8 .472 ;1)$G9?G-$ Cryptocoryne
blassii +$6P#;H1?0.(>=?. .G03$%&68"./&00$7/B, 23$)&*+$45+6 "7/8$97/-. 9@$'5 <$'./2$#F6%PCK95'7&. 2549 . &#!H!) H!%5;)$M%N;>(0-:2!)567. AI;'%5O. %&2%-E/5?<*BC 2.
9@$'5 <$'./2$#F6%PCK95'7& /L*/)<)&*+$8$97/-A4L6*E1'14+B, /:<6*% Q.
2549J. J);FN$/:.I172!N4 ) H!<(#($. <$'6I+A%/;A4L $/./LH$952%5',- 9@$'5 O/&3$/(5N $/N>$'%45<<$'./2$#F 9@$'5 <$'./2$#F6%PCK95'7&, /:<*% Q.
2549E. ;1)55#$.;>@?&#!6!-#56!!IN !#25#$H!<B*#H[ 2547-2548. <$'6I+A%/;./LH$952%5',- 9@$'5 O/&3$/(5N $/N>$'./2$#F 9@$'5 <$'./2$#F6%PCK95'7& /L*/)<)&*+$8$97/-A4L6*E1'14+B, /:<6*% Q.
9&/':H 4$28/B(5'*/-. 2527. :/(#;)$M%J#6G!N!)567. E#L)&*+$8$97/- 23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
2536. &#!&1#2;)$M%NE?5;>(. aG''BC%5OO4&HH&C<, /:<6*% Q.
2540. &#!&1#2;)$M%NE?5;>(. %&2%-E/5?<*BC 2. 9@$'5 %&2%-23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
2546. )&,B $/A4LJ5?'7K'[' $/./5O./:<%5',:-=2>NK =2>./LN5O1N+ $/[H>/5<9BA 22$, '. 10-24. ! 8F'+-O/& $/\$+/5<9BA 22$A4L)&(5+'&)6E4B+/-6*E1'14+B. 2546. .FI$/I&#!6!-#5;)$M%NA'-0?&A'-H!<0) 0-:2!)5678&''#. 6K 9$/./L KO $/]^ KO/21E/< $/)&(5+6H&<X;$+*KN6*E1'14+B $/%5M'$%5',:-=2>NK =2>./LN5ON>)+/5<9BA 22$9F;6 "7/ /. )5'*BC 26-27 2&X:'$+' 2546 # 8F'+-O/& $/\$+/5<9BA 22$A4L)&(5+'&)6E4B+/-6*E1'14+B 23$)&*+$45+6 "7/8$97/-, /:<6*%.
9:(&7/$ 6%H/E<. 2548. D1E?5!)5678&''#F7@'7G9?1)&L"<F#56)"R#$E?5;!!"A'-$=*#+ ;#2 Cryptocoryne balansae Gagnep. )&*+$'&%',-./&00$1*,
23$)&*+$45+6 "7/8$97/-.
74
9:H$N$ 8/BO:06/PK<. 2550. &#!.,$7@2:$*#+,-.&/0&#!&1#2;)$M%N+$AF!2-?2+ 8,1'09#5302+(-!)5678&''#. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-
9:'B+- (5'*/8/B. 2547. D1E?58I1.O72'81<&#!4#2!)5678&''#G9?6!7!:/F2#81<I%"P#;E?50?&&1-:2A'-6&%1,:#2. )&*+$'&%',-./&00$1*, 23$)&*+$45+6*E1'14+B%/L(K26 4>$,'O:/B.
9:2'$ &(=%bF/+-. 2528. &#!&1#2;)$M%NE?5 73&.$72302&#!.;#<.17=25.$>=?.2>@?!9:'&) &#!4#2!)5678&''#. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
9:/)&H )//#= /1/('-. 2526. D1E?5!)56781<6#!.I'7G9?&%,1# ,/$;)$M%N1#!N3&[email protected]=25+$6P#;H1?0.(>=?. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
KN&8/ /LA9H5+. 2533. $/8c "$ $/./5O./:<%5',:-.*:22$ (Curcuma sparganifolia) 1N+ $/[H>/5<9BA 22$. :. .&LG!J#6G!N (:/F2.) 26 (1): 6-11.
2535. I4JK</5<9BA 22;$7;KNK `;K' 4&C'=*+. :. .&LG!J#6G!N (:/F2.) 24 (2):
125-135.
K7&'5'*- (5'S&/2+-. 2545. D1E?5!)5678&''#F7@'7G9?&#!.B!/Q;)Y$#E?5.$>=?.2>@?69:$,):E?5:9#$67@F/JF7@.;#<.17=25+$6P#;H1?0.(>=?. )&*+$'&%',-./&00$1*,
23$)&*+$45+JK'A ;'.
KS&H$7& H&NO:/B, %<8-+:*, ')4O:06/PK< A4L %&*5 "- %:*,)/H5+. 2543. &#!.;#<.17=250#,1#.;>@?&#!H!) H!%5;)$M%N. /$+<$' $/)&(5+. 9X$O5')&(5+A4L]^ KO/2 $/6 "7/4@$.$< 23$)&*+$45+6*E14+B/$H2<E44>$''$. A34;<*BC2$: http://dcms.thailis.or.th, 18 2&X:'$+'
2550.
K/NB 93)5H/&'*/- A4L H5+H:2%4 9:/&+L85 N&T. 2527. .FI$/I&#!H!) H!%5;)$M%NI#!N.$()@$302&#!?# !)5678&''#E"<.17=25+$,1?0F01?5. /$+<$' $/)&(5+ ./L(@$.W 2527.
9X$O5')&(5+A4L%5M'$A3;<23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
K/':H K5<9:J8&/&. 2545. D1E?5!)5678&''#81<&#!B%9'.?M#$?1G9?&#!&*#B)0.;17=2A\ 81<&#!+(-$=*#2#2>0?#2%0?&&1-:2A'-,:#2;)$M%N ?' #17 F7@4#2!)567. )&*+$'&%',-./&00$1*, 23$)&*+$45+6*E1'14+B%/L(K26 4>$,'O:/B.
K/)//# 2F4*K<($N. 2524. D1E?5!)5678&''9#G9?8&10/3?1)6. .G03$%&68"./&00$1*,
23$)&*+$45+6 "7/8$97/-. 2525. D1E?5!)5678&''#F7@'7G9?1)&L"< #5H!<&#!E?58&10/3?1)6.
)&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-. K/)//# )<"-)$'&H. 2548. H)'H2%5',:-[32;($ $/\$+/5<9B. .I,&#!.&LG! 29(4): 154-158.
75
K/:#B )<8-.Y+L9X&7+-. 2530 . .?&6#!I*#6?$:/(#:/M7H!) H!%5;)$M%N302&#!&1#2;)$M%N. S$E)&H$/5<9B./L+: 7-A4L=K1`1*. E#L)&*+$8$97/- 23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
2530J. .?&6#!I*#6?$:/(#!)567(7::/F2#. S$E)&H$/5<9B./L+: 7-A4L=K1`1*. E#L)&*+$8$97/- 23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
2539. .?&6#!I*#6?$:/(#&#!+(-!)56781<A?3O3FH+$&#!.&LG!.!>@?5&#!+(-!)567 +$&#!H!) H!%5;)$M%N;>(. S$E)&H$/5<9B./L+: 7-A4L=K1`1*. E#L)&*+$8$97/- 23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
2540. .?&6#!I*#6?$:/(#!)567&) (7:/G81<6/@58:01-?' .!>@?5!)56781<6/@5'7(7:/G.
S$E)&H$/5<9B./L+: 7-A4L=K1`1*. E#L)&*+$8$97/- 23$)&*+$45+6 "7/8$97/-, /:<6*% Q.
2546. E)$2/F>6OP?K<7>'6 BC+) 5O/5<9BA4L $/[H>/5<9B[' $/./5O./:<%5',:-=2>NK =2>./LN5O,
'. 1-9. ! 8F'+-O/& $/\$+/5<9BA 22$A4L)&(5+'&)6E4B+/-6*E1'14+B. 2546. .FI$/I&#!6!-#5;)$M%NA'-0?&A'-H!<0) 0-:2!)5678&''#. 6K 9$/./L KO $/]^ KO/21E/< $/)&(5+6H&<X;$+*KN6*E1'14+B $/%5M'$%5',:-=2>NK =2>./LN5ON>)+/5<9BA 22$9F;6 "7/ /. )5'*BC 26-27 2&X:'$+' 2546 # 8F'+-O/& $/\$+/5<9BA 22$A4L)&(5+'&)6E4B+/-6*E1'14+B 23$)&*+$45+6 "7/8$97/-, /:<6*%.
2550. &#!+(-!)567+$&#!;)Y$#;)$M%NA'-0?&A'-H!<0) . S$E)&H$/5<9B./L+: 7-A4L=K1`1*. E#L)&*+$8$97/- 23$)&*+$45+6 "7/8$97/-. A34;<*BC2$:http://www.doa.go.th/web-itc/web2005/news_doa/x_ray_flower5.pdf, 5 9&<3$E2 2550
A4L ')4\)B /:;<,'6 B+/7&. 2534. D1E?5!)5678&''#G9?&#!.H17@2$8H151)&L"<81<670?&8;!.O7@25A]-. /$+<$' $/)&(5+. S$E)&H$/5<9B./L+: 7-A4L=K1`1*. E#L)&*+$8$97/-, 23$)&*+$45+6 "7/8$97/-.
2536. /5<9B 5O $/ 4$+%5',:-['=2>NK =2>./LN5O. :. :/F2#J#6G!N.&LG! 26 (4-6):
113-127 '.
K50H:4B H&'9:J. 2546. JK&L#6SG!?#,#!F7@.,'#<6'81<D1E?5!)5678&''#F7@'7G9?1)&L"<F#56)"R#$:/F2#E?5&1-:2 ):676-'. )&*+$'&%',-./&00$1*,
23$)&*+$45+6 "7/8$97/- K$S$S/#- )5M')&6HB+/. 2539. D1E?5!)5678&''#G9?&#!.B!/Q.G/ 3G 81<&#!D1/G,):29?2
E?51/117@+$6P#;H1?0.(>=?. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-. K$/+$ K$(6(/&0. 2548. D1E?5&#!4#2!)5678&''#G9?&#!.H17@2$8H15E?51)&L"<F#5
6)"R#$:/F2#81<1#2;/';N07.?T$.?+$&1-:2. ; (Musa acuminata ‘BEP’).
)&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
76
K:=/ 6/PK<#/<E-. 2542. &#!.;#<.17=25.$>=?.2>@?81<&#!()&$*#+,-.&/0&#!&1#2+$G-$?.'O?$302+(-!)5678&''#. )&*+$'&%',-./&00$1*, 23$)&*+$45+6 "7/8$97/-.
6K2K/ */5%+-6(/&0. 2530. D1E?5!)567F7@'7G9?&#!.B!/Q.G/ 3GE?5 ):6%M#6/3$ 1.
.G03$%&68"./&00$7/B, 9X$O5'6*E1'14+B%/L(K26 4>$6(>$E:#*3$/4$N /LO5<. Auerbach C. 1976. Mutation research : problems, results and perspectives / Charlotte
Auerbach. Chapman & Hall, London.
Broertjes, C. and A.M. van Harten. 1978. Application of mutation breeding methods in
the improvement of vegetatively propagated crops : an interpretive literature
review. Elsevier Scienfic Publishing, Amsterdam.
Casarett, A. P. 1968. Radiation Biology. Prentice-Hall Inc. Englewood Cliffs, New
Jersey.
Gottschalk, W. and G. Wolff. 1983. Induced Mutations in Plant Breeding. Springer-
Verlag, Berlin.
IAEA. 1977. Manual on Mutation Breeding. Technical Reports Series No. 119. 2nd
ed.
Joint FAO/IAEA Division of Atomic Energy in Food and Agriculture, Vienna.
Kovács E. and Á. Keresztes. 2002. Effect of Gamma and UV B/C radiation on Plant Cells.
Micron 33: 199-110.
Lama S., Aeksomtramaet L. and K. Kanchanapoom. 2005. Induced Mutation in Nelumbo
nucifera Gaertn. by Gamma Irradiation. 31st Congress on Science and
Technology of Thailand at Suranaree University of Technology, 18 – 20 October
2005. Available Source: http://www.scisoc.or.th/stt/31/sec_b/paper/stt31_B0015.pdf,
9 July 2008. (in Thai Abstract)
Lamseejan, S., Jompuk, P., Wongpiyasatid, A., Deeseepan, S. and P. Kwanthammachart.
2000. Gamma-reys Induced Morphological Changes in Chrysanthemum
(Chrysanthemum morifolium). Kasetsart J. (Nat. Sci.) 34 (3): 417-422.
Puripunyavanich V. and K. Boonsirichai. 2005. Effect of Gamma Irradiation in
‘Jongkolnee’ Water lily. 31st Congress on Science and Technology of Thailand at
Suranaree University of Technology, 18 – 20 October 2005. Available Source:
http://www.scisoc.or.th/stt/31/sec_f/paper/stt31_F0054.pdf, 9 July 2008. (in Thai
Abstract)
Wongpiyasatid, A. and P. Hormchan. 2000. New Mutants of Perenial Portulaca grandiflora
Through Gamma Radiation. Kasetsart J. (Nat. Sci.) 34 (3): 408-416.