classification of the extant echinoderm at a

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  • 8/6/2019 Classification of the Extant Echinoderm at A

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    Classification of the Extant Echinodermata This list also contains links to images.We welcome updates or corrections to this list.

    Crinoidea Isocrinida

    Isocrinidae

    Millericrinida

    Apiocrinidae Bathycrinidae Bourgueticrinidae Phrynocrinidae

    Cyrtocrinida

    Holopusidae Hyocrinidae

    Comatulida

    Comasteridae Colobometridae Eudiocrinidae Himerometridae Mariametridae Zygometridae Aporometridae Notocrinidae Asterometridae Calometridae Charitometridae Ptilometridae Thalassometridae Tropiometridae Antedonidae Apocrinidae Atelecrinidae Pentametrocrinidae

    Asteroidea (after Blake, 1987 and Clark andDowney, 1992)

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    Paxillosida

    Astropectinidae

    25 valid genera

    Ctenodiscididae

    This is a monotypic family including only the genus Ctenodiscus with five species australis , caudatus ,crispatus , orientalis and procurator .

    Goniopectinidae

    3 valid genera.

    Luidiidae

    This family is monotypic containing only the genus Luidia . Members of this family are burrowers intosandy substrates. Members of this family have anywere from 5 to 15 arms. Several species in this taxa areof considerable size. One species Luidia superba from the Galapagos is one of the largest asteroids in theworld.

    Porcellanasteridae

    Eleven genera This is an entirely deep-sea group inhabiting infaunal habitats. Representatives of this groupare found in the deepest of the worlds oceans. Many taxa have cosmopolitan distributions.

    Radiasteridae

    This is a monotypic group including only the genus Radiaster with four species. All are found in relativelydeep-water. Notomyotida

    Benthopectinidae

    12 valid genera. Benthopectinids are an exclusively deep-sea taxon of asteroids found in the Atlantic andPacific Oceans. Valvatida

    Acanthasteridae

    This monotypic multi-armed family represents the genus Acanthaster with two species planci andbrevispinis. Acanthaster is distributed from the East Pacific (Baja California) to the East coast of Africa.

    Acanthaster planci is the notorious Crown of Thorns starfish whose mysterious population outbreaks haveresulted in a perceived threat to the coral reefs of the southwest Pacific.

    Archasteridae

    http://research.calacademy.org/research/izg/echinoderm/images/cots.jpghttp://research.calacademy.org/research/izg/echinoderm/images/cots.jpg
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    This family represents the sole genus Archaster found in the Indo-Pacific. Archaster is an infaunal burrower which superficially resemble members of the family Astropectinidae except that they havesuckered tube feet. Archaster engages in a peculiar mating behavior known as pseudocopulation. Thisoccurs when representatives of opposite genders crawl upon one another and spawn, presumably increasingthe probability of fertilization.

    Asterinidae

    18 valid genera. Asterinids are found throughout the world in tropical and temperate waters.

    Asterodiscididae

    3 genera. All of which were formerly included in both the Goniasteridae and the Oreasteridae. They arefound only in the tropical East and Indo-Pacific.

    Asteropsidae

    5 genera.

    Chaetasteridae

    monotypic (1 genus)

    Ganeriidae

    9 valid genera. Members of this family are mostly poriferivorous and are found exclusively in the AntarcticOcean (and surrounding areas).

    Goniasteridae

    ~ 53 valid genera. This family represents an very diverse and broadly distributed taxa found in all theworlds oceans at all depths.

    Mithrodidae

    2 valid genera.This family is closely related to the Ophidiasteridae. It includes two genera found thoughoutthe Indo-Pacific with a small population on the Caribbean side of the Panamic seaway. One genus in thisfamily, Thromidia , is one of the largest asteroids in the world with an arm to arm span of ca. 2.5 feet anddiameter of approx. 5 inches. Genera Include: Thromidia and Mithrodia

    Odontasteridae

    5 Valid genera. Odontasterids apparently feed primarily on encrusting invertebrates, especially poriferans.

    Oreasteridae

    19 Valid genera. Oreasterids are represented primarily in the tropical Indo and East Pacific however twospecies of Oreaster are found in the tropical Atlantic. These large heavily armored asteroids are primarilymicroherbivores and opportunistic scavengers. They are strikingly colored and are often sought after by thetourist industry for their large and ornate tests.

    http://research.calacademy.org/research/izg/echinoderm/images/asterina.jpghttp://research.calacademy.org/research/izg/echinoderm/images/amphiast.jpghttp://research.calacademy.org/research/izg/echinoderm/images/mediastr.jpghttp://research.calacademy.org/research/izg/echinoderm/images/mithrodi.jpghttp://research.calacademy.org/research/izg/echinoderm/images/oreaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/asterina.jpghttp://research.calacademy.org/research/izg/echinoderm/images/amphiast.jpghttp://research.calacademy.org/research/izg/echinoderm/images/mediastr.jpghttp://research.calacademy.org/research/izg/echinoderm/images/mithrodi.jpghttp://research.calacademy.org/research/izg/echinoderm/images/oreaster.jpg
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    Genera include: Oreaster, Protoreaster, Pentaster, Culcita, Choriaster, Rosaster, Pentaceraster, Nidorellia , Anthenea

    Ophidiasteridae

    31 Valid genera. Ophidiasterids occur in both Atlantic and Pacific Oceans. The greatest diversity of thisgroup occurs in the Indo-Pacific. Members of this family are often brilliantly colored (e.g., Fromia,

    Neoferdina ). Some ophidiasterids possess remarkable powers of autotomy and regeneration (e.g., Linckia ).Genera Include: Fromia, Linckia, Neoferdina, Nardoa, Ceratonardoa, Ophidiaster, Tamaria, Phataria,

    Pharia , Leiaster ,

    Poraniidae

    9 Valid genera.

    Sphaerasteridae

    monotypic (1 genus) Velatida

    Caymanostellidae

    These are an entirely deep-sea taxon which may possibly related to Concentricycloids. They are found onlyat abyssal depths in the Atlantic and Pacific Oceans.

    Korethasteridae Myxasteridae

    Myxasterids are reminiscent of pterasterids but are have longer arms. They generally posses more than 5

    arms.Genera include: Myxaster

    Pterasteridae

    Pterasterids are found in cold/ temperate but primarily deep-water habitats although there is one taxonwhich lives in the Indo-Pacific. These asteroids generally possess very short arms relative to their discs andan unusual membrane which is supra-dorsal to the aboral surface of the asteroid itself.Genera include: Pteraster, Hymenodiscus, Diplopteraster, Retaster ,

    Solasteridae

    Solasterid asteroids are often multi-armed. With one exception, most are found in mostly cold/deep-water habitats. Solasterids appear to be predatory/ opportunistic feeders. Genera include: Lophaster, Heterozonias, Solaster, Seriaster Spinulosida

    Echinasteridae

    The species complex of Arctic Henricia has long been a challenge to asteroid population biologists. Thelatter genus, Echinaster is circum- tropical being found in both Pacific and Atlantic oceans.

    http://research.calacademy.org/research/izg/echinoderm/images/nidoreli.jpghttp://research.calacademy.org/research/izg/echinoderm/images/linckia.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pinflata.jpghttp://research.calacademy.org/research/izg/echinoderm/images/solaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/henrcia2.jpghttp://research.calacademy.org/research/izg/echinoderm/images/nidoreli.jpghttp://research.calacademy.org/research/izg/echinoderm/images/linckia.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pinflata.jpghttp://research.calacademy.org/research/izg/echinoderm/images/solaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/henrcia2.jpg
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    Forcipulatida

    Asteriidae

    Asteriids are among the best known and familiar of the asteroids. Representatives of this group includes thewell known Atlantic Asterias and NW Pacific Pisaster . Members of this family all possess four rows of tube feet. Several members of this family are voracious predators of bivalve molluscs.

    Noteworthy asteriids include: Asterias amurensis , the asteroid introduced to Tasmania from Northern Japan belongs to this family. Several asteriids are of considerable size. Pycnopodia helianthoides found on thePacific Northwest coast of North America can attain a diamter (arm to arm) of almost 3 feet!! Specimens of

    Pisaster brevispinis and Stylasterias forreri can have a diameter (arm to arm) of approx. 2 feet.

    Heliasteridae

    This monotypic family represents the genus Heliaster found from Baja California and along the coast of South America and out to the Galapagos Islands. Heliasterids have 20-35 arms and are intertidal/subtidal

    predators of small molluscs and barnacles. These form a species complex along their range- the taxonomyof which has yet to be fully resolved.

    Labidiasteridae

    All members of this family are multi-armed and are found in moderately deep Atlantic and Pacific oceans.At least two members of this family, Rathbunaster and Labidiaster can catch small crustaceans with their arms and pedicellariae.Genera included in this family: Coronaster, Rathbunaster, Labidiaster, Plazaster

    Neomorphasteridae Pedicellasteridae Zoroasteridae

    Zoroasterids are found only in deep-sea habitats. They possess long spindly arms. Fossil zoroasterids fromAntarctica (Blake and Zinsmeister, 1983) show that some taxa existed in a shallower warmer habitat before'moving' into deep-water habitats.Brisingida Brisingids are a group of exclusively deep-sea asteroids. They are most closely related to the Forcipulatidaand are unusual in possessing 6 or more arms, (usually 12 to 18) and a fused oral disc. Brisingids aresuspension feeders. They hold their long, attenuate arms into the water column using high densities of

    pedicellariae on their arm spines which act as velcro to catch small prey. (Emson and Young, 1994) One brisingid, Midgardia xandaros Downey, 1970 possess the longest arm span of any known asteroid.

    Brisingidae

    Genera include: Astrolirus, Astrostephane, Brisinga, Brisingaster, Stegnobrisinga, Midgardia, Brisingenes, Hymenodiscus/ Brisingella, Parabrisinga, Novodinia

    Freyellidae

    Genera include Colpaster, Freyella, Freyastera, Freyellaster

    Concentricycloidea

    http://research.calacademy.org/research/izg/echinoderm/images/pycnopod.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pisaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/heliastr.jpghttp://research.calacademy.org/research/izg/echinoderm/images/rathbun.jpghttp://research.calacademy.org/research/izg/echinoderm/images/brisinga.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pycnopod.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pisaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/heliastr.jpghttp://research.calacademy.org/research/izg/echinoderm/images/rathbun.jpghttp://research.calacademy.org/research/izg/echinoderm/images/brisinga.jpg
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    Concentricycloids or "sea daisies" are an enigmatic group of echinoderms discovered in 1986 from deep-water off New Zealand on woody debris. This taxa contains the single genus Xyloplax with two species,one in the Pacific and one in the Atlantic. Positive determination of the sister taxon to the concentricycloids

    remains elusive although evidence suggests a relationship with asteroids.

    Ophiuroidea Phyrynophiurida

    Asteronychidae Asteroschematidae Euryalidae Gorgonocephalidae

    Ophiomyxida

    Ophiomyxidae Ophiocanopidae

    Ophiurida

    Ophiocomidae Ophiodermatidae Ophiuridae Ophioleucidae

    This is a deep-sea group, historically placed with the Ophiuridae.Hemieuryalidae Ophionereidae OphiothricidaeOphiothricids are apparently successful in the tropical Pacific. Several taxa are epizoic on octocorals andother invertebrates. Tropical members of this family are also very colorful, often retaining their stunning

    purples and reds after preservation.Amphiuridae Ophiactidae

    Echinoidea Cidaroida

    Cidaridae Psychocidaridae

    Echinothurioida

    Echinothuriidae

    Deep-water and tropical echinoids.Diadematoida

    Aspidodiadematidae Diadematidae Lissodiadematidae Micropygidae

    http://research.calacademy.org/research/izg/echinoderm/images/gorgono.jpghttp://research.calacademy.org/research/izg/echinoderm/images/gorgono.jpg
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    Pedinoida

    Pedinidae

    Salenioida

    Saleniidae

    Phymosomatoida

    Phymosomatidae Stomechinidae Stomopneustidae

    Arbacioida

    Arbaciidae

    Temnopleuroida

    Temnopleuridae Toxopneustidae

    Echinoida

    Echinidae Echinometridae

    Parasaleniidae Strongylocentrotidae

    HolectypoidaClypeasteroida

    Arachnoididae Clypeasteridae

    Laganina

    Fibulariidae Laganidae

    Scutellina

    Astriclypeidae Dendrasteridae Echinarachniidae

    http://research.calacademy.org/research/izg/echinoderm/images/tripneus.jpghttp://research.calacademy.org/research/izg/echinoderm/images/redurchn.jpghttp://research.calacademy.org/research/izg/echinoderm/images/laganum1.jpghttp://research.calacademy.org/research/izg/echinoderm/images/dendrab.jpghttp://research.calacademy.org/research/izg/echinoderm/images/tripneus.jpghttp://research.calacademy.org/research/izg/echinoderm/images/redurchn.jpghttp://research.calacademy.org/research/izg/echinoderm/images/laganum1.jpghttp://research.calacademy.org/research/izg/echinoderm/images/dendrab.jpg
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    Mellitidae

    Rotulina

    Rotulidae

    Cassiduloida

    Cassidulidae Echinolampadidae Pliolampadidae

    Holasteroida

    Calymnidae Holasteridae

    Pourtalesiidae Urechinidae

    Spatangoida

    Toxasteridae Aeropsidae Hemiasteridae Palaeostomatidae Pericosmidae Schizasteridae

    Brissidae Loveniidae Spatangidae

    Neolampadoida

    Neolampadidae

    Holothuroidea Dendrochirotida

    Cucumariidae Paracucumidae Phyllophoridae Placothuriidae Psolidae Sclerodactylidae

    Dactylochirotida

    http://research.calacademy.org/research/izg/echinoderm/images/brissus.jpghttp://research.calacademy.org/research/izg/echinoderm/images/psolus.jpghttp://research.calacademy.org/research/izg/echinoderm/images/brissus.jpghttp://research.calacademy.org/research/izg/echinoderm/images/psolus.jpg
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    Rhopalodinidae Vaneyellidae Ypsilothuriidae

    Aspidochirotida

    Holothuriidae Stichopodidae Synallactidae

    Elasipodida These all represent exclusively deep-sea fauna.

    Deimatidae Elpidiidae Laetmogonidae Pelagothuriidae

    Psychropotidae

    Apodida

    Chiridotidae Myriotrochidae Synaptidae

    Molpadida

    Caudinidae

    Gephyrothuriidae Molpadiidae

    Classification of the Extant Echinodermata This list also contains links to images.We welcome updates or corrections to this list.

    Crinoidea Isocrinida

    Isocrinidae

    Millericrinida

    Apiocrinidae Bathycrinidae

    http://research.calacademy.org/research/izg/echinoderm/images/cuke.jpghttp://research.calacademy.org/research/izg/echinoderm/images/cuke.jpg
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    Bourgueticrinidae Phrynocrinidae

    Cyrtocrinida

    Holopusidae Hyocrinidae

    Comatulida

    Comasteridae Colobometridae Eudiocrinidae Himerometridae Mariametridae Zygometridae

    Aporometridae Notocrinidae Asterometridae Calometridae Charitometridae Ptilometridae Thalassometridae Tropiometridae Antedonidae Apocrinidae Atelecrinidae

    Pentametrocrinidae

    Asteroidea (after Blake, 1987 and Clark andDowney, 1992) Paxillosida

    Astropectinidae

    25 valid genera

    Ctenodiscididae

    This is a monotypic family including only the genus Ctenodiscus with five species australis , caudatus ,crispatus , orientalis and procurator .

    Goniopectinidae

    3 valid genera.

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    Luidiidae

    This family is monotypic containing only the genus Luidia . Members of this family are burrowers intosandy substrates. Members of this family have anywere from 5 to 15 arms. Several species in this taxa areof considerable size. One species Luidia superba from the Galapagos is one of the largest asteroids in theworld.

    Porcellanasteridae

    Eleven genera This is an entirely deep-sea group inhabiting infaunal habitats. Representatives of this groupare found in the deepest of the worlds oceans. Many taxa have cosmopolitan distributions.

    Radiasteridae

    This is a monotypic group including only the genus Radiaster with four species. All are found in relativelydeep-water. Notomyotida

    Benthopectinidae

    12 valid genera. Benthopectinids are an exclusively deep-sea taxon of asteroids found in the Atlantic andPacific Oceans. Valvatida

    Acanthasteridae

    This monotypic multi-armed family represents the genus Acanthaster with two species planci andbrevispinis. Acanthaster is distributed from the East Pacific (Baja California) to the East coast of Africa.

    Acanthaster planci is the notorious Crown of Thorns starfish whose mysterious population outbreaks haveresulted in a perceived threat to the coral reefs of the southwest Pacific.

    Archasteridae

    This family represents the sole genus Archaster found in the Indo-Pacific. Archaster is an infaunal burrower which superficially resemble members of the family Astropectinidae except that they havesuckered tube feet. Archaster engages in a peculiar mating behavior known as pseudocopulation. Thisoccurs when representatives of opposite genders crawl upon one another and spawn, presumably increasingthe probability of fertilization.

    Asterinidae

    18 valid genera. Asterinids are found throughout the world in tropical and temperate waters.

    Asterodiscididae

    3 genera. All of which were formerly included in both the Goniasteridae and the Oreasteridae. They arefound only in the tropical East and Indo-Pacific.

    Asteropsidae

    http://research.calacademy.org/research/izg/echinoderm/images/cots.jpghttp://research.calacademy.org/research/izg/echinoderm/images/asterina.jpghttp://research.calacademy.org/research/izg/echinoderm/images/amphiast.jpghttp://research.calacademy.org/research/izg/echinoderm/images/cots.jpghttp://research.calacademy.org/research/izg/echinoderm/images/asterina.jpghttp://research.calacademy.org/research/izg/echinoderm/images/amphiast.jpg
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    5 genera.

    Chaetasteridae

    monotypic (1 genus)

    Ganeriidae

    9 valid genera. Members of this family are mostly poriferivorous and are found exclusively in the AntarcticOcean (and surrounding areas).

    Goniasteridae

    ~ 53 valid genera. This family represents an very diverse and broadly distributed taxa found in all theworlds oceans at all depths.

    Mithrodidae

    2 valid genera.This family is closely related to the Ophidiasteridae. It includes two genera found thoughoutthe Indo-Pacific with a small population on the Caribbean side of the Panamic seaway. One genus in thisfamily, Thromidia , is one of the largest asteroids in the world with an arm to arm span of ca. 2.5 feet anddiameter of approx. 5 inches. Genera Include: Thromidia and Mithrodia

    Odontasteridae

    5 Valid genera. Odontasterids apparently feed primarily on encrusting invertebrates, especially poriferans.

    Oreasteridae

    19 Valid genera. Oreasterids are represented primarily in the tropical Indo and East Pacific however twospecies of Oreaster are found in the tropical Atlantic. These large heavily armored asteroids are primarilymicroherbivores and opportunistic scavengers. They are strikingly colored and are often sought after by thetourist industry for their large and ornate tests.Genera include: Oreaster, Protoreaster, Pentaster, Culcita, Choriaster, Rosaster, Pentaceraster,

    Nidorellia , Anthenea

    Ophidiasteridae

    31 Valid genera. Ophidiasterids occur in both Atlantic and Pacific Oceans. The greatest diversity of thisgroup occurs in the Indo-Pacific. Members of this family are often brilliantly colored (e.g., Fromia,

    Neoferdina ). Some ophidiasterids possess remarkable powers of autotomy and regeneration (e.g., Linckia ).Genera Include: Fromia, Linckia, Neoferdina, Nardoa, Ceratonardoa, Ophidiaster, Tamaria, Phataria,

    Pharia , Leiaster ,

    Poraniidae

    9 Valid genera.

    Sphaerasteridae

    http://research.calacademy.org/research/izg/echinoderm/images/mediastr.jpghttp://research.calacademy.org/research/izg/echinoderm/images/mithrodi.jpghttp://research.calacademy.org/research/izg/echinoderm/images/oreaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/nidoreli.jpghttp://research.calacademy.org/research/izg/echinoderm/images/linckia.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pinflata.jpghttp://research.calacademy.org/research/izg/echinoderm/images/mediastr.jpghttp://research.calacademy.org/research/izg/echinoderm/images/mithrodi.jpghttp://research.calacademy.org/research/izg/echinoderm/images/oreaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/nidoreli.jpghttp://research.calacademy.org/research/izg/echinoderm/images/linckia.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pinflata.jpg
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    monotypic (1 genus) Velatida

    Caymanostellidae

    These are an entirely deep-sea taxon which may possibly related to Concentricycloids. They are found onlyat abyssal depths in the Atlantic and Pacific Oceans.

    Korethasteridae Myxasteridae

    Myxasterids are reminiscent of pterasterids but are have longer arms. They generally posses more than 5arms.Genera include: Myxaster

    Pterasteridae

    Pterasterids are found in cold/ temperate but primarily deep-water habitats although there is one taxonwhich lives in the Indo-Pacific. These asteroids generally possess very short arms relative to their discs andan unusual membrane which is supra-dorsal to the aboral surface of the asteroid itself.Genera include: Pteraster, Hymenodiscus, Diplopteraster, Retaster ,

    Solasteridae

    Solasterid asteroids are often multi-armed. With one exception, most are found in mostly cold/deep-water habitats. Solasterids appear to be predatory/ opportunistic feeders. Genera include: Lophaster,

    Heterozonias, Solaster, Seriaster Spinulosida

    EchinasteridaeThe species complex of Arctic Henricia has long been a challenge to asteroid population biologists. Thelatter genus, Echinaster is circum- tropical being found in both Pacific and Atlantic oceans.Forcipulatida

    Asteriidae

    Asteriids are among the best known and familiar of the asteroids. Representatives of this group includes thewell known Atlantic Asterias and NW Pacific Pisaster . Members of this family all possess four rows of tube feet. Several members of this family are voracious predators of bivalve molluscs.

    Noteworthy asteriids include: Asterias amurensis , the asteroid introduced to Tasmania from Northern Japan belongs to this family. Several asteriids are of considerable size. Pycnopodia helianthoides found on thePacific Northwest coast of North America can attain a diamter (arm to arm) of almost 3 feet!! Specimens of

    Pisaster brevispinis and Stylasterias forreri can have a diameter (arm to arm) of approx. 2 feet.

    Heliasteridae

    This monotypic family represents the genus Heliaster found from Baja California and along the coast of South America and out to the Galapagos Islands. Heliasterids have 20-35 arms and are intertidal/subtidal

    http://research.calacademy.org/research/izg/echinoderm/images/solaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/henrcia2.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pycnopod.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pisaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/heliastr.jpghttp://research.calacademy.org/research/izg/echinoderm/images/solaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/henrcia2.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pycnopod.jpghttp://research.calacademy.org/research/izg/echinoderm/images/pisaster.jpghttp://research.calacademy.org/research/izg/echinoderm/images/heliastr.jpg
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    predators of small molluscs and barnacles. These form a species complex along their range- the taxonomyof which has yet to be fully resolved.

    Labidiasteridae

    All members of this family are multi-armed and are found in moderately deep Atlantic and Pacific oceans.At least two members of this family, Rathbunaster and Labidiaster can catch small crustaceans with their arms and pedicellariae.Genera included in this family: Coronaster, Rathbunaster, Labidiaster, Plazaster

    Neomorphasteridae Pedicellasteridae Zoroasteridae

    Zoroasterids are found only in deep-sea habitats. They possess long spindly arms. Fossil zoroasterids fromAntarctica (Blake and Zinsmeister, 1983) show that some taxa existed in a shallower warmer habitat before'moving' into deep-water habitats.Brisingida Brisingids are a group of exclusively deep-sea asteroids. They are most closely related to the Forcipulatidaand are unusual in possessing 6 or more arms, (usually 12 to 18) and a fused oral disc. Brisingids aresuspension feeders. They hold their long, attenuate arms into the water column using high densities of

    pedicellariae on their arm spines which act as velcro to catch small prey. (Emson and Young, 1994) One brisingid, Midgardia xandaros Downey, 1970 possess the longest arm span of any known asteroid.

    Brisingidae

    Genera include: Astrolirus, Astrostephane, Brisinga, Brisingaster, Stegnobrisinga, Midgardia, Brisingenes, Hymenodiscus/ Brisingella, Parabrisinga, Novodinia

    Freyellidae

    Genera include Colpaster, Freyella, Freyastera, Freyellaster

    Concentricycloidea Concentricycloids or "sea daisies" are an enigmatic group of echinoderms discovered in 1986from deep-water off New Zealand on woody debris. This taxa contains the single genus

    Xyloplax with two species, one in the Pacific and one in the Atlantic. Positive determination of the sister taxon to the concentricycloids remains elusive although evidence suggests a relationship with

    asteroids.

    Ophiuroidea Phyrynophiurida

    Asteronychidae Asteroschematidae Euryalidae Gorgonocephalidae

    Ophiomyxida

    Ophiomyxidae

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    Ophiocanopidae

    Ophiurida

    Ophiocomidae

    Ophiodermatidae Ophiuridae Ophioleucidae

    This is a deep-sea group, historically placed with the Ophiuridae.Hemieuryalidae Ophionereidae OphiothricidaeOphiothricids are apparently successful in the tropical Pacific. Several taxa are epizoic on octocorals andother invertebrates. Tropical members of this family are also very colorful, often retaining their stunning

    purples and reds after preservation.Amphiuridae Ophiactidae

    Echinoidea Cidaroida

    Cidaridae Psychocidaridae

    Echinothurioida

    Echinothuriidae

    Deep-water and tropical echinoids.Diadematoida

    Aspidodiadematidae Diadematidae Lissodiadematidae Micropygidae

    Pedinoida

    Pedinidae

    Salenioida

    Saleniidae

    Phymosomatoida

    Phymosomatidae Stomechinidae Stomopneustidae

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    Calymnidae Holasteridae Pourtalesiidae Urechinidae

    Spatangoida Toxasteridae Aeropsidae Hemiasteridae Palaeostomatidae Pericosmidae Schizasteridae Brissidae Loveniidae Spatangidae

    Neolampadoida

    Neolampadidae

    Holothuroidea Dendrochirotida

    Cucumariidae Paracucumidae Phyllophoridae Placothuriidae Psolidae Sclerodactylidae

    Dactylochirotida

    Rhopalodinidae Vaneyellidae Ypsilothuriidae

    Aspidochirotida

    Holothuriidae Stichopodidae Synallactidae

    Elasipodida These all represent exclusively deep-sea fauna.

    Deimatidae

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    Elpidiidae Laetmogonidae Pelagothuriidae Psychropotidae

    Apodida Chiridotidae Myriotrochidae Synaptidae

    Molpadida

    Caudinidae Gephyrothuriidae Molpadiidae